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<document id="DA89A4DDE72451D98488544632F1E7A0" ID="10.11646/zootaxa.4205.3.6" ID-DOI="10.11646/zootaxa.4205.3.6" ID-GBIF-Dataset="d8d24aba-900e-4dcc-97cd-adb094ebe3ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="194329" ID-ZooBank="66FB30C3-66BC-4D79-9361-03F5C9F22498" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481096845972" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="03D7CE3F344CF322A8CF6480FC65F979" docLanguage="en" docName="zootaxa.4205.3.6.pdf" docOrigin="Zootaxa 4205 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmaenus (Geoscydmaenus) isaloensis Franz" docType="treatment" docVersion="8" lastPageNumber="271" masterDocId="FFEEB647344FF324A8586722FFEEFFA9" masterDocTitle="Review of Scydmaenus (Geoscydmaenus) of Madagascar (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="274" masterPageNumber="265" pageNumber="268" updateTime="1698396176682" updateUser="plazi">
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<mods:title id="28123B36BB03ADBC6D74EC7C72B3057A">Review of Scydmaenus (Geoscydmaenus) of Madagascar (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="947B73C38A56E2510572C912A3ABCEF7">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="F1754B9C6D3B5C813A56802DC79D5B78">2016</mods:date>
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<treatment id="03D7CE3F344CF322A8CF6480FC65F979" ID-DOI="http://doi.org/10.5281/zenodo.6053365" ID-GBIF-Taxon="124956957" ID-Zenodo-Dep="6053365" LSID="urn:lsid:plazi:treatment:03D7CE3F344CF322A8CF6480FC65F979" httpUri="http://treatment.plazi.org/id/03D7CE3F344CF322A8CF6480FC65F979" lastPageId="6" lastPageNumber="271" pageId="3" pageNumber="268">
<subSubSection id="C3642CA2344CF327A8CF6480FDA2FC77" pageId="3" pageNumber="268" type="nomenclature">
<paragraph id="8BC17F29344CF327A8CF6480FD34FC15" blockId="3.[151,730,930,990]" box="[151,730,930,956]" pageId="3" pageNumber="268">
<heading id="D089C845344CF327A8CF6480FD34FC15" bold="true" box="[151,730,930,956]" fontSize="11" level="1" pageId="3" pageNumber="268" reason="1">
<taxonomicName id="4C7E04AA344CF327A8CF6480FD34FC15" ID-CoL="4W3VF" authority="Franz" authorityName="Franz" box="[151,730,930,956]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="isaloensis" subGenus="Geoscydmaenus">
<emphasis id="B90AA33B344CF327A8CF6480FD34FC15" bold="true" box="[151,730,930,956]" pageId="3" pageNumber="268">
<emphasis id="B90AA33B344CF327A8CF6480FEC0FC15" bold="true" box="[151,302,930,956]" italics="true" pageId="3" pageNumber="268">Scydmaenus</emphasis>
(
<emphasis id="B90AA33B344CF327A9676480FDECFC15" bold="true" box="[319,514,930,956]" italics="true" pageId="3" pageNumber="268">Geoscydmaenus</emphasis>
)
<emphasis id="B90AA33B344CF327AA4C6480FD67FC15" bold="true" box="[532,649,930,956]" italics="true" pageId="3" pageNumber="268">isaloensis</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC17F29344CF327A8CF64E7FDA2FC77" blockId="3.[151,730,930,990]" box="[151,588,965,990]" pageId="3" pageNumber="268">
(
<figureCitation id="134563AC344CF327A8C764E7FECEFC77" box="[159,288,965,990]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="3" pageNumber="268">Figs 1, 34</figureCitation>
,
<figureCitation id="134563AC344CF327A97364E4FEA6FC77" box="[299,328,966,990]" captionStart="FIGURES 9 11" captionStartId="5.[151,264,1613,1636]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@5.[151,1436,193,1592]" captionTargetPageId="5" captionText="FIGURES 9 11. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus madagassicus Franz (9 10) and Sc. isaloensis Franz (11). Pterothorax and abdomen in ventral view (9); pterothorax, transparent mount in ventral view (10); profemoral ventral tooth of male in lateral view (11). Abbreviations: aest 3, metanepisternum; ar, anterior ridge; lf, lateral fovea; lmfa, lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus." httpUri="https://zenodo.org/record/267455/files/figure.png" pageId="3" pageNumber="268">11</figureCitation>
,
<figureCitation id="134563AC344CF327A90C64E4FE72FC77" box="[340,412,966,990]" captionStart="FIGURES 12 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="3" pageNumber="268">12, 14</figureCitation>
,
<figureCitation id="134563AC344CF327A9F064E7FE1EFC77" box="[424,496,965,990]" captionStart="FIGURES 16 21" captionStartId="7.[151,264,1906,1929]" captionTargetBox="[164,1415,224,1854]" captionTargetId="figure@7.[151,1436,193,1885]" captionTargetPageId="7" captionText="FIGURES 16 21. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (16 18) and Sc. madagassicus Franz (19 21). Aedeagus in ventral (16, 19) and lateral (17 18, 20) view (with partly exserted flagellum in Fig. 18); spermatheca (21)." httpUri="https://zenodo.org/record/194333/files/figure.png" pageId="3" pageNumber="268">1618</figureCitation>
,
<figureCitation id="134563AC344CF327A9A464E4FDADFC77" box="[508,579,966,990]" captionStart="FIGURES 22 25" captionStartId="8.[151,264,804,827]" captionTargetBox="[163,1428,208,766]" captionTargetId="figure@8.[151,1436,193,784]" captionTargetPageId="8" captionText="FIGURES 22 25. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (22 23) and Sc. madagassicus Franz (24 25). Aedeagus in ventral (22, 24) and lateral (23, 25) views." httpUri="https://zenodo.org/record/194334/files/figure.png" pageId="3" pageNumber="268">2223</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3642CA2344CF327A8CF632FFC6BFB8B" box="[151,901,1037,1059]" pageId="3" pageNumber="268" type="reference_group">
<paragraph id="8BC17F29344CF327A8CF632FFC6BFB8B" blockId="3.[151,901,1037,1059]" box="[151,901,1037,1059]" pageId="3" pageNumber="268">
<treatmentCitationGroup id="AB6E5807344CF327A8CF632FFC6BFB8B" box="[151,901,1037,1059]" pageId="3" pageNumber="268">
<taxonomicName id="4C7E04AA344CF327A8CF632FFD00FB8B" ID-CoL="4W3VF" authority="Franz, 1986: 372" authorityName="Franz" authorityPageNumber="372" authorityYear="1986" box="[151,750,1037,1059]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="isaloensis" subGenus="Geoscydmaenus">
<emphasis id="B90AA33B344CF327A8CF632FFEFAFB8B" box="[151,276,1037,1058]" italics="true" pageId="3" pageNumber="268">Scydmaenus</emphasis>
(
<emphasis id="B90AA33B344CF327A97B632FFE29FB8B" box="[291,455,1037,1058]" italics="true" pageId="3" pageNumber="268">Geoscydmaenus</emphasis>
)
<emphasis id="B90AA33B344CF327A98E632FFDD4FB8B" box="[470,570,1037,1058]" italics="true" pageId="3" pageNumber="268">isaloensis</emphasis>
<treatmentCitation id="0ADF5938344CF327AA18632CFD00FB8B" author="Franz" box="[576,750,1037,1059]" page="372" pageId="3" pageNumber="268" year="1986">
<bibRefCitation id="EFEF02D8344CF327AA18632CFD52FB8A" author="Franz" box="[576,700,1037,1059]" pageId="3" pageNumber="268" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986</bibRefCitation>
: 372
</treatmentCitation>
</taxonomicName>
, figs 339341.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3642CA2344CF327A8CF636DFAEDFACD" pageId="3" pageNumber="268" type="materials_examined">
<paragraph id="8BC17F29344CF327A8CF636DFB86FB7D" blockId="3.[151,1437,1103,2029]" pageId="3" pageNumber="268">
<materialsCitation id="3B167574344CF327A8CF636DFE34FB7D" ID-GBIF-Occurrence="1413803492" collectionCode="NHMW" collectorName="Massif de l'Isalo &amp; H. Franz &amp; Mg" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="3" pageNumber="268" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B90AA33B344CF327A8CF636DFD5AFBC1" bold="true" box="[151,692,1103,1128]" pageId="3" pageNumber="268">
<typeStatus id="54C5C18B344CF327A8CF636DFF3CFBC1" box="[151,210,1103,1128]" pageId="3" pageNumber="268">Type</typeStatus>
material.
<typeStatus id="54C5C18B344CF327A902636DFE29FBC1" box="[346,455,1103,1128]" pageId="3" pageNumber="268" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3693FB9344CF327A985636DFD40FBC1" box="[477,686,1103,1128]" name="Madagascar" pageId="3" pageNumber="268">MADAGASCAR</collectingCountry>
)
</emphasis>
:
<specimenCount id="9D78B4A0344CF327AA936372FD0CFBCE" box="[715,738,1104,1127]" pageId="3" pageNumber="268" type="male"></specimenCount>
, two labels (
<figureCitation id="134563AC344CF327ABDD636DFC21FBC1" box="[901,975,1103,1128]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="3" pageNumber="268">Fig. 1</figureCitation>
): &quot;S-Madagaskar /
<collectorName id="268B1AFF344CF327AC98636DFA68FBC1" box="[1216,1414,1103,1128]" pageId="3" pageNumber="268">Massif de l'Isalo</collectorName>
/ lg.
<collectorName id="268B1AFF344CF327A8E86356FEE0FB24" box="[176,270,1140,1165]" pageId="3" pageNumber="268">H.Franz</collectorName>
1969&quot; [white, printed] with &quot;
<collectorName id="268B1AFF344CF327AADD6356FD41FB25" box="[645,687,1140,1164]" pageId="3" pageNumber="268">Mg</collectorName>
20&quot; handwritten in blue on the reverse side;
<taxonomicName id="4C7E04AA344CF327ACAF6356FA6AFB24" box="[1271,1412,1140,1165]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="274" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="4C7E04AA344CF327A8C663B5FEB7FB19" box="[158,345,1175,1200]" class="Insecta" family="Staphylinidae" genus="Geoscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="267" phylum="Arthropoda" rank="genus">Geoscydmaenus</taxonomicName>
) /
<taxonomicName id="4C7E04AA344CF327A92463B5FE07FB19" box="[380,489,1175,1200]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="isaloensis">isaloensis</taxonomicName>
m. / det.
<determinerName id="7E1D93ED344CF327AA0C63B5FD5CFB19" box="[596,690,1175,1200]" pageId="3" pageNumber="268">H.Franz</determinerName>
&quot; [white, handwritten and printed]; &quot;
<specimenCount id="9D78B4A0344CF327AC0C63BAFB85FB19" box="[1108,1131,1176,1200]" pageId="3" pageNumber="268" type="male"></specimenCount>
&quot; [white, printed]; &quot;
<typeStatus id="54C5C18B344CF327AD1263BAFA7DFB19" box="[1354,1427,1176,1200]" pageId="3" pageNumber="268">Typus</typeStatus>
&quot; [red, handwritten] (
<collectionCode id="ED6FE7EC344CF327A928639EFE3DFB7D" box="[368,467,1212,1236]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="268">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B167574344CF327A9BC639EFB8AFB7D" ID-GBIF-Occurrence="1413803490" box="[484,1124,1212,1237]" collectionCode="NHMW" collectorName="Massif de l'Isalo &amp; H. Franz &amp; Mg" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="3" pageNumber="268" specimenCount="2" typeStatus="paratype">
<emphasis id="B90AA33B344CF327A9BC639EFD88FB7C" bold="true" box="[484,614,1212,1237]" pageId="3" pageNumber="268">
<typeStatus id="54C5C18B344CF327A9BC639EFD8FFB7C" box="[484,609,1212,1237]" pageId="3" pageNumber="268" type="paratype">Paratypes</typeStatus>
:
</emphasis>
12 exx., same data as for
<typeStatus id="54C5C18B344CF327ABD1639EFC05FB7C" box="[905,1003,1212,1237]" pageId="3" pageNumber="268" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED6FE7EC344CF327ABA2639EFBB2FB7D" box="[1018,1116,1212,1236]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="268">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC17F29344CF327A89F63FDFAEDFACD" blockId="3.[151,1437,1103,2029]" pageId="3" pageNumber="268">
<materialsCitation id="3B167574344CF327A89F63FDFCD0FAB4" ID-GBIF-Occurrence="1413803489" collectingDate="2013-01-17" collectingDateMax="2013-01-18" collectingDateMin="2013-01-17" collectorName="Isalo N. P. &amp; L. S. Rahanitriniaina &amp; E. M. Rabotoson" country="Madagascar" elevation="730" location="Ananalava forest" pageId="3" pageNumber="268" specimenCount="1">
<emphasis id="B90AA33B344CF327A89F63FDFDF1FB51" bold="true" box="[199,543,1247,1272]" pageId="3" pageNumber="268">Additional material studied:</emphasis>
16 exx.,
<collectingCountry id="F3693FB9344CF327AAD763FDFCF6FB51" box="[655,792,1247,1272]" name="Madagascar" pageId="3" pageNumber="268">Madagascar</collectingCountry>
,
<collectorName id="268B1AFF344CF327AB7E63FDFC76FB5E" box="[806,920,1247,1272]" pageId="3" pageNumber="268">Isalo N.P.</collectorName>
,
<location id="8EA129F2344CF327ABFE63FDFB85FB51" LSID="urn:lsid:plazi:treatment:03D7CE3F344CF322A8CF6480FC65F979:8EA129F2344CF327ABFE63FDFB85FB51" box="[934,1131,1247,1272]" country="Madagascar" name="Ananalava forest" pageId="3" pageNumber="268">Ananalava forest</location>
,
<elevation id="0053981A344CF327AC2063C2FB16FB5E" box="[1144,1272,1248,1272]" metricMagnitude="2" metricUnit="m" metricValue="7.3" metricValueMax="7.409999999999999" metricValueMin="7.19" pageId="3" pageNumber="268" unit="m" value="730.0" valueMax="741.0" valueMin="719.0">
<quantity id="4C86D2CC344CF327AC2063C2FB16FB5E" box="[1144,1272,1248,1272]" metricMagnitude="2" metricUnit="m" metricValue="7.3" metricValueMax="7.409999999999999" metricValueMin="7.19" pageId="3" pageNumber="268" unit="m" value="730.0" valueMax="741.0" valueMin="719.0">719-741 m</quantity>
</elevation>
,
<collectingDate id="EF84A001344CF327AD5E63FDFA76FB51" box="[1286,1432,1247,1272]" pageId="3" pageNumber="268" value="2013-01-17" valueMax="2013-01-18" valueMin="2013-01-17">
<date id="FFC059E9344CF327AD5E63FDFA76FB51" box="[1286,1432,1247,1272]" pageId="3" pageNumber="268" value="2013-01-17" valueMax="2013-01-18" valueMin="2013-01-17">17-18.i.2013</date>
</collectingDate>
, sifted leaf litter,
<collectorName id="268B1AFF344CF327A9166226FDDFFAB4" box="[334,561,1284,1309]" pageId="3" pageNumber="268">L.S. Rahanitriniaina</collectorName>
&amp;
<collectorName id="268B1AFF344CF327AA0D6227FCE5FAB4" box="[597,779,1284,1309]" pageId="3" pageNumber="268">E.M. Rabotoson</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B167574344CF327AB126226FC07FAE9" ID-GBIF-Occurrence="1413803494" collectingDate="2013-01-21" collectorName="Isalo N. P. &amp; L. S. Rahanitriniaina &amp; E. M. Rabotoson" country="Madagascar" elevation="852" location="Piscine" pageId="3" pageNumber="268" specimenCount="1">
2 exx.,
<collectingCountry id="F3693FB9344CF327ABC46226FBCBFAB4" box="[924,1061,1284,1309]" name="Madagascar" pageId="3" pageNumber="268">Madagascar</collectingCountry>
,
<collectorName id="268B1AFF344CF327AC686226FB71FAB5" box="[1072,1183,1284,1309]" pageId="3" pageNumber="268">Isalo N.P.</collectorName>
,
<location id="8EA129F2344CF327ACF26226FB15FAB4" LSID="urn:lsid:plazi:treatment:03D7CE3F344CF322A8CF6480FC65F979:8EA129F2344CF327ACF26226FB15FAB4" box="[1194,1275,1284,1309]" country="Madagascar" name="Piscine" pageId="3" pageNumber="268">Piscine</location>
naturelle,
<elevation id="0053981A344CF327AD2A6226FF41FA96" metricMagnitude="2" metricUnit="m" metricValue="8.52" pageId="3" pageNumber="268" unit="m" value="852.0">
<quantity id="4C86D2CC344CF327AD2A6226FF41FA96" metricMagnitude="2" metricUnit="m" metricValue="8.52" pageId="3" pageNumber="268" unit="m" value="852.0">852 m</quantity>
</elevation>
,
<collectingDate id="EF84A001344CF327A8E46205FEC7FAE9" box="[188,297,1319,1344]" pageId="3" pageNumber="268" value="2013-01-21">
<date id="FFC059E9344CF327A8E46205FEC7FAE9" box="[188,297,1319,1344]" pageId="3" pageNumber="268" value="2013-01-21">21.i.2013</date>
</collectingDate>
, sifted leaf litter,
<collectorName id="268B1AFF344CF327A9A9620AFD38FAE9" box="[497,726,1319,1344]" pageId="3" pageNumber="268">L.S. Rahanitriniaina</collectorName>
&amp;
<collectorName id="268B1AFF344CF327AAA4620AFC5AFAE9" box="[764,948,1319,1344]" pageId="3" pageNumber="268">E.M. Rabotoson</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B167574344CF327ABAE620AFAEDFACD" ID-GBIF-Occurrence="1413803491" collectingDate="2013-01-20" collectionCode="MMBC" collectorName="Isalo N. P. &amp; L. S. Rahanitriniaina &amp; E. M. Rabotoson" country="Madagascar" elevation="894" location="Oasis forest" pageId="3" pageNumber="268" specimenCount="1">
2 exx.,
<collectingCountry id="F3693FB9344CF327AC136205FB3AFAE9" box="[1099,1236,1319,1344]" name="Madagascar" pageId="3" pageNumber="268">Madagascar</collectingCountry>
,
<collectorName id="268B1AFF344CF327ACB96205FABFFA96" box="[1249,1361,1319,1344]" pageId="3" pageNumber="268">Isalo N.P.</collectorName>
,
<location id="8EA129F2344CF327AD066205FF34FACC" LSID="urn:lsid:plazi:treatment:03D7CE3F344CF322A8CF6480FC65F979:8EA129F2344CF327AD066205FF34FACC" country="Madagascar" name="Oasis forest" pageId="3" pageNumber="268">Oasis forest</location>
,
<elevation id="0053981A344CF327A8BC626EFEC3FACD" box="[228,301,1356,1380]" metricMagnitude="2" metricUnit="m" metricValue="8.94" pageId="3" pageNumber="268" unit="m" value="894.0">
<quantity id="4C86D2CC344CF327A8BC626EFEC3FACD" box="[228,301,1356,1380]" metricMagnitude="2" metricUnit="m" metricValue="8.94" pageId="3" pageNumber="268" unit="m" value="894.0">894 m</quantity>
</elevation>
,
<collectingDate id="EF84A001344CF327A961626EFE4BFACC" box="[313,421,1356,1381]" pageId="3" pageNumber="268" value="2013-01-20">
<date id="FFC059E9344CF327A961626EFE4BFACC" box="[313,421,1356,1381]" pageId="3" pageNumber="268" value="2013-01-20">20.i.2013</date>
</collectingDate>
, sifted leaf litter,
<collectorName id="268B1AFF344CF327AA31626EFCA2FACC" box="[617,844,1356,1381]" pageId="3" pageNumber="268">L.S. Rahanitriniaina</collectorName>
&amp;
<collectorName id="268B1AFF344CF327AB37626FFBC8FACC" box="[879,1062,1356,1381]" pageId="3" pageNumber="268">E.M. Rabotoson</collectorName>
leg. (
<collectionCode id="ED6FE7EC344CF327AC3D626EFB2EFACD" box="[1125,1216,1356,1380]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="268">MMBC</collectionCode>
, cPJ).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3642CA2344CF327A89F624DFC5BFA5C" pageId="3" pageNumber="268" type="diagnosis">
<paragraph id="8BC17F29344CF327A89F624DFC5BFA5C" blockId="3.[151,1437,1103,2029]" pageId="3" pageNumber="268">
<emphasis id="B90AA33B344CF327A89F624DFE4FFA21" bold="true" box="[199,417,1391,1416]" pageId="3" pageNumber="268">Revised diagnosis.</emphasis>
Head broadest near posterior margin of vertex, strongly narrowing anteriorly; profemur of male with clavate part strongly convex dorsally and ventrally, with long ventral projection located near middle, its apex hook-shaped; protibia of male abruptly broadened distally and slightly but distinctly curved; aedeagus with deeply notched apex, lateral apical lobes distinctly separated at middle.
</paragraph>
</subSubSection>
<subSubSection id="C3642CA2344CF322A89F62DDFB0FF905" lastPageId="6" lastPageNumber="271" pageId="3" pageNumber="268" type="description">
<paragraph id="8BC17F29344CF327A89F62DDFBFDF995" blockId="3.[151,1437,1103,2029]" pageId="3" pageNumber="268">
<emphasis id="B90AA33B344CF327A89F62DDFE9AF9B1" bold="true" box="[199,372,1535,1560]" pageId="3" pageNumber="268">Redescription.</emphasis>
Body of male (
<figureCitation id="134563AC344CF327AA6C62DDFD94F9B1" box="[564,634,1535,1560]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="3" pageNumber="268">Fig. 3</figureCitation>
) strongly elongate and slender, strongly flattened, yellowish to light brown and covered with light brown setae; BL
<quantity id="4C86D2CC344CF327AAF06106FCA5F995" box="[680,843,1572,1597]" metricMagnitude="-4" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.299999999999999" metricValueMin="7.6" pageId="3" pageNumber="268" unit="mm" value="0.7949999999999999" valueMax="0.83" valueMin="0.76">0.760.83 mm</quantity>
(mean
<quantity id="4C86D2CC344CF327ABC66106FBE8F995" box="[926,1030,1572,1596]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="268" unit="mm" value="0.8">0.80 mm</quantity>
).
</paragraph>
<paragraph id="8BC17F29344CF327A89F6165FD7AF829" blockId="3.[151,1437,1103,2029]" pageId="3" pageNumber="268">
Head (
<figureCitation id="134563AC344CF327A9496165FE98F9C9" box="[273,374,1607,1632]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="3" pageNumber="268">Figs 34</figureCitation>
) in dorsal view subtriangular with weakly rounded sides, broadest shortly in front of occipital constriction and strongly narrowing anteriorly, HL
<quantity id="4C86D2CC344CF327AA89614EFC9AF92D" box="[721,884,1644,1669]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="3" pageNumber="268" unit="mm" value="0.14" valueMax="0.15" valueMin="0.13">0.130.15 mm</quantity>
(mean
<quantity id="4C86D2CC344CF327AB9F614EFBC0F92D" box="[967,1070,1644,1668]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="268" unit="mm" value="0.14">0.14 mm</quantity>
), HW
<quantity id="4C86D2CC344CF327AC2E614EFAF6F92D" box="[1142,1304,1644,1669]" metricMagnitude="-4" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="268" unit="mm" value="0.155" valueMax="0.16" valueMin="0.15">0.150.16 mm</quantity>
(mean
<quantity id="4C86D2CC344CF327AD33614EFF29F901" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="268" unit="mm" value="0.15">0.15 mm</quantity>
); vertex and frons confluent, evenly but weakly convex; supraantennal tubercles feebly marked. Dorsal surface of head covered with shallow but distinct and relatively large punctures distributed mostly on sides of vertex and frons, with much smaller punctures on median area (
<figureCitation id="134563AC344CF327AABB61F5FCC9F959" box="[739,807,1751,1776]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="3" pageNumber="268">Fig. 4</figureCitation>
), but the size of finely punctate region is variable and in some specimens large punctures occupy nearly entire frons and vertex; setae short and sparse, nearly recumbent. Antennae (
<figureCitation id="134563AC344CF327A949603DFE8DF891" box="[273,355,1823,1848]" captionStart="FIGURES 12 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="3" pageNumber="268">Fig. 12</figureCitation>
) short, AnL
<quantity id="4C86D2CC344CF327A9AA603DFD7BF891" box="[498,661,1823,1848]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="3" pageNumber="268" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.330.35 mm</quantity>
(mean
<quantity id="4C86D2CC344CF327AAB16002FCBFF891" box="[745,849,1824,1848]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="268" unit="mm" value="0.34">0.34 mm</quantity>
); scape and pedicel strongly elongate, antennomeres IIIV each distinctly elongate, VI about as long as broad, VII and VIII distinctly transverse, IX and X about as long as broad, XI about 1.61.7× as long as broad.
</paragraph>
<paragraph id="8BC17F29344CF327A89F60AFFD8DF844" blockId="3.[151,1437,1103,2029]" pageId="3" pageNumber="268">
Pronotum in dorsal view (
<figureCitation id="134563AC344CF327A9B060AEFDA2F80D" box="[488,588,1932,1957]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="3" pageNumber="268">Figs 34</figureCitation>
) oval, broadest between middle and anterior third; PL
<quantity id="4C86D2CC344CF327ACF560AEFABEF80D" box="[1197,1360,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="3" pageNumber="268" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.210.23 mm</quantity>
(mean
<quantity id="4C86D2CC344CF327A8CF6092FEEEF861" box="[151,256,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="268" unit="mm" value="0.22">0.22 mm</quantity>
), PW
<quantity id="4C86D2CC344CF327A91F6092FE02F861" box="[327,492,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="3" pageNumber="268" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.200.23 mm</quantity>
(mean
<quantity id="4C86D2CC344CF327AA1B6092FD43F861" box="[579,685,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="268" unit="mm" value="0.21">0.21 mm</quantity>
). Punctures on pronotal disc shallow and diffused, inconspicuous; setae short, sparse and nearly recumbent.
</paragraph>
<paragraph id="8BC17F29344BF320A89F67B5FCBAFF5E" blockId="4.[151,1437,151,248]" pageId="4" pageNumber="269">
Elytra (
<figureCitation id="134563AC344BF320A94267B5FEB5FF19" box="[282,347,151,176]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="4" pageNumber="269">Fig. 3</figureCitation>
) oval and strongly elongate, broadest near anterior third, EL
<quantity id="4C86D2CC344BF320AC5A67B5FB4BFF06" box="[1026,1189,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="4" pageNumber="269" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.430.45 mm</quantity>
(mean
<quantity id="4C86D2CC344BF320ACAF67BAFAB0FF19" box="[1271,1374,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="269" unit="mm" value="0.44">0.44 mm</quantity>
), EW
<quantity id="4C86D2CC344BF320A8CF679EFED5FF7D" box="[151,315,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="4" pageNumber="269" unit="mm" value="0.265" valueMax="0.28" valueMin="0.25">0.250.28 mm</quantity>
(mean
<quantity id="4C86D2CC344BF320A9C9679EFE15FF7D" box="[401,507,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="269" unit="mm" value="0.27">0.27 mm</quantity>
), EI 1.551.80. Elytra covered with distinct and dense punctures, which are small and with diffused margins; setae similar to those on pronotum.
</paragraph>
<caption id="DF012FA1344BF320A8CF607BFB0CF862" httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="4" pageNumber="269" targetBox="[168,1436,295,1860]" targetPageId="4">
<paragraph id="8BC17F29344BF320A8CF607BFB0CF862" blockId="4.[151,1436,1881,1996]" pageId="4" pageNumber="269">
<emphasis id="B90AA33B344BF320A8CF607BFED2F8C6" bold="true" box="[151,316,1881,1904]" pageId="4" pageNumber="269">FIGURES 18.</emphasis>
<taxonomicName id="4C7E04AA344BF320A91C6078FD90F8C6" authority="Geoscydmaenus" authorityName="Geoscydmaenus" box="[324,638,1882,1903]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="B90AA33B344BF320A91C6078FE2FF8C6" box="[324,449,1882,1903]" italics="true" pageId="4" pageNumber="269">Scydmaenus</emphasis>
(
<emphasis id="B90AA33B344BF320A98A6078FD98F8C6" box="[466,630,1882,1903]" italics="true" pageId="4" pageNumber="269">Geoscydmaenus</emphasis>
)
</taxonomicName>
of Madagascar.
<taxonomicName id="4C7E04AA344BF320AB746078FBB6F8C6" authority="Franz" authorityName="Franz" box="[812,1112,1882,1903]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="isaloensis">
<emphasis id="B90AA33B344BF320AB746078FBF8F8C6" box="[812,1046,1882,1903]" italics="true" pageId="4" pageNumber="269">Scydmaenus isaloensis</emphasis>
Franz
</taxonomicName>
(1, 34) and
<taxonomicName id="4C7E04AA344BF320ACBF6078FF3FF827" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="madagassicus">
<emphasis id="B90AA33B344BF320ACBF6078FA72F8C6" box="[1255,1436,1882,1903]" italics="true" pageId="4" pageNumber="269">Sc. madagassicus</emphasis>
Franz
</taxonomicName>
(2, 58). Set of original labels of holotype (12); dorsal habitus of male (3); head and prothorax in dorsal (45) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex.
</paragraph>
</caption>
<caption id="DF012FA1344AF321A8CF616FFBDBF977" httpUri="https://zenodo.org/record/267455/files/figure.png" pageId="5" pageNumber="270" targetBox="[151,1436,193,1584]" targetPageId="5">
<paragraph id="8BC17F29344AF321A8CF616FFBDBF977" blockId="5.[151,1436,1613,1758]" pageId="5" pageNumber="270">
<emphasis id="B90AA33B344AF321A8CF616FFEA6F9CA" bold="true" box="[151,328,1613,1636]" pageId="5" pageNumber="270">FIGURES 911.</emphasis>
<taxonomicName id="4C7E04AA344AF321A908616CFD62F9CA" authority="Geoscydmaenus" authorityName="Geoscydmaenus" box="[336,652,1614,1635]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="B90AA33B344AF321A908616CFE23F9CA" box="[336,461,1614,1635]" italics="true" pageId="5" pageNumber="270">Scydmaenus</emphasis>
(
<emphasis id="B90AA33B344AF321A986616CFD6CF9CA" box="[478,642,1614,1635]" italics="true" pageId="5" pageNumber="270">Geoscydmaenus</emphasis>
)
</taxonomicName>
of Madagascar.
<taxonomicName id="4C7E04AA344AF321AB61616CFB7EF9CA" authority="Franz" authorityName="Franz" box="[825,1168,1614,1635]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="madagassicus">
<emphasis id="B90AA33B344AF321AB61616CFBA3F9CA" box="[825,1101,1614,1635]" italics="true" pageId="5" pageNumber="270">Scydmaenus madagassicus</emphasis>
Franz
</taxonomicName>
(910) and
<taxonomicName id="4C7E04AA344AF321AD49616CFF3FF92B" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="isaloensis">
<emphasis id="B90AA33B344AF321AD49616CFA72F9CA" box="[1297,1436,1614,1635]" italics="true" pageId="5" pageNumber="270">Sc. isaloensis</emphasis>
Franz
</taxonomicName>
(11). Pterothorax and abdomen in ventral view (9); pterothorax, transparent mount in ventral view (10); profemoral ventral tooth of male in lateral view (11). Abbreviations: aest3, metanepisternum; ar, anterior ridge; lf, lateral fovea; lmfa, lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus.
</paragraph>
</caption>
<paragraph id="8BC17F29344AF321A89F602EFD0FF826" blockId="5.[151,1436,1803,2007]" pageId="5" pageNumber="270">
Legs (
<figureCitation id="134563AC344AF321A9556029FEB9F88A" box="[269,343,1803,1828]" captionStart="FIGURES 1 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 4) and Sc. madagassicus Franz (2, 5 8). Set of original labels of holotype (1 2); dorsal habitus of male (3); head and prothorax in dorsal (4 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="5" pageNumber="270">Figs 3</figureCitation>
,
<figureCitation id="134563AC344AF321A93C6029FE6FF88A" box="[356,385,1803,1827]" captionStart="FIGURES 9 11" captionStartId="5.[151,264,1613,1636]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@5.[151,1436,193,1592]" captionTargetPageId="5" captionText="FIGURES 9 11. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus madagassicus Franz (9 10) and Sc. isaloensis Franz (11). Pterothorax and abdomen in ventral view (9); pterothorax, transparent mount in ventral view (10); profemoral ventral tooth of male in lateral view (11). Abbreviations: aest 3, metanepisternum; ar, anterior ridge; lf, lateral fovea; lmfa, lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus." httpUri="https://zenodo.org/record/267455/files/figure.png" pageId="5" pageNumber="270">11</figureCitation>
,
<figureCitation id="134563AC344AF321A9D66029FE45F88A" box="[398,427,1803,1827]" captionStart="FIGURES 12 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="5" pageNumber="270">14</figureCitation>
) relatively short and robust; profemur (
<figureCitation id="134563AC344AF321AB356029FC51F88A" box="[877,959,1803,1828]" captionStart="FIGURES 12 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="5" pageNumber="270">Fig. 14</figureCitation>
) with large clavate part similarly strongly convex dorsally and ventrally, with ventral submedian projection bearing small apical denticle directed toward trochanter and long seta at base of projection (
<figureCitation id="134563AC344AF321AAFE6071FD16F8C2" box="[678,760,1875,1900]" captionStart="FIGURES 9 11" captionStartId="5.[151,264,1613,1636]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@5.[151,1436,193,1592]" captionTargetPageId="5" captionText="FIGURES 9 11. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus madagassicus Franz (9 10) and Sc. isaloensis Franz (11). Pterothorax and abdomen in ventral view (9); pterothorax, transparent mount in ventral view (10); profemoral ventral tooth of male in lateral view (11). Abbreviations: aest 3, metanepisternum; ar, anterior ridge; lf, lateral fovea; lmfa, lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus." httpUri="https://zenodo.org/record/267455/files/figure.png" pageId="5" pageNumber="270">Fig. 11</figureCitation>
); protibia (
<figureCitation id="134563AC344AF321AB2F6071FC27F8C2" box="[887,969,1875,1900]" captionStart="FIGURES 12 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="5" pageNumber="270">Fig. 14</figureCitation>
) strongly and abruptly thickened slightly distally to middle, and slightly but distinctly curved.
</paragraph>
<paragraph id="8BC17F29344AF321A89F60B9FB30F87E" blockId="5.[151,1436,1803,2007]" pageId="5" pageNumber="270">
Aedeagus (
<figureCitation id="134563AC344AF321A91560B9FE39F81D" box="[333,471,1947,1972]" captionStart="FIGURES 16 21" captionStartId="7.[151,264,1906,1929]" captionTargetBox="[164,1415,224,1854]" captionTargetId="figure@7.[151,1436,193,1885]" captionTargetPageId="7" captionText="FIGURES 16 21. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (16 18) and Sc. madagassicus Franz (19 21). Aedeagus in ventral (16, 19) and lateral (17 18, 20) view (with partly exserted flagellum in Fig. 18); spermatheca (21)." httpUri="https://zenodo.org/record/194333/files/figure.png" pageId="5" pageNumber="270">Figs 1618</figureCitation>
,
<figureCitation id="134563AC344AF321A9B460B9FDDCF81A" box="[492,562,1947,1971]" captionStart="FIGURES 22 25" captionStartId="8.[151,264,804,827]" captionTargetBox="[163,1428,208,766]" captionTargetId="figure@8.[151,1436,193,784]" captionTargetPageId="8" captionText="FIGURES 22 25. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (22 23) and Sc. madagassicus Franz (24 25). Aedeagus in ventral (22, 24) and lateral (23, 25) views." httpUri="https://zenodo.org/record/194334/files/figure.png" pageId="5" pageNumber="270">2223</figureCitation>
) elongate, AeL
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; median lobe in ventral view with distinctly demarcated apical portion bearing a pair of lateral apical lobes separated at middle by deep notch.
</paragraph>
<caption id="DF012FA13449F322A8CF62E8FB7EF9A9" httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="6" pageNumber="271" targetBox="[151,1427,210,1461]" targetPageId="6">
<paragraph id="8BC17F293449F322A8CF62E8FB7EF9A9" blockId="6.[151,1436,1482,1536]" pageId="6" pageNumber="271">
<emphasis id="B90AA33B3449F322A8CF62E8FEBDFA49" bold="true" box="[151,339,1482,1505]" pageId="6" pageNumber="271">FIGURES 1215.</emphasis>
<taxonomicName id="4C7E04AA3449F322A90262E9FD7DFA48" authority="Geoscydmaenus" authorityName="Geoscydmaenus" box="[346,659,1483,1505]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="B90AA33B3449F322A90262E9FE39FA49" box="[346,471,1483,1504]" italics="true" pageId="6" pageNumber="271">Scydmaenus</emphasis>
(
<emphasis id="B90AA33B3449F322A9BD62E9FD67FA49" box="[485,649,1483,1504]" italics="true" pageId="6" pageNumber="271">Geoscydmaenus</emphasis>
)
</taxonomicName>
of Madagascar.
<taxonomicName id="4C7E04AA3449F322AB6262E9FB8CFA48" authority="Franz" authorityName="Franz" box="[826,1122,1483,1505]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="isaloensis">
<emphasis id="B90AA33B3449F322AB6262E9FBCFFA49" box="[826,1057,1483,1504]" italics="true" pageId="6" pageNumber="271">Scydmaenus isaloensis</emphasis>
Franz
</taxonomicName>
(12, 14) and
<taxonomicName id="4C7E04AA3449F322ACB162EEFF3FFA56" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="madagassicus">
<emphasis id="B90AA33B3449F322ACB162EEFAE6FA48" box="[1257,1288,1484,1505]" italics="true" pageId="6" pageNumber="271">Sc.</emphasis>
<emphasis id="B90AA33B3449F322AD5662E9FA72FA49" box="[1294,1436,1483,1504]" italics="true" pageId="6" pageNumber="271">madagassicus</emphasis>
Franz
</taxonomicName>
(13, 15). Right antenna of male in dorsal view (1213); right fore leg of male in anterior view.
</paragraph>
</caption>
<paragraph id="8BC17F293449F322A89F6105FB0FF905" blockId="6.[151,1437,1575,1744]" pageId="6" pageNumber="271">
<emphasis id="B90AA33B3449F322A89F6105FECBF9E9" bold="true" box="[199,293,1575,1600]" pageId="6" pageNumber="271">Female.</emphasis>
Similar to male, except for distinctly more slender profemora lacking ventral projections and nearly straight and only slightly, gradually thickened protibiae. BL
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(mean
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); HL
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(mean
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), HW
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(mean
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), AnL
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; PL
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(mean
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), PW
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(mean
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); EL
<quantity id="4C86D2CC3449F322AA6A61B1FD76F902" box="[562,664,1683,1707]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="271" unit="mm" value="0.43">0.43 mm</quantity>
, EW
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(mean
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), EI 1.621.70.
</paragraph>
</subSubSection>
<subSubSection id="C3642CA23449F322A89F6195FC65F979" box="[199,907,1719,1744]" pageId="6" pageNumber="271" type="distribution">
<paragraph id="8BC17F293449F322A89F6195FC65F979" blockId="6.[151,1437,1575,1744]" box="[199,907,1719,1744]" pageId="6" pageNumber="271">
<emphasis id="B90AA33B3449F322A89F6195FE8EF979" bold="true" box="[199,352,1719,1744]" pageId="6" pageNumber="271">Distribution.</emphasis>
Central-southern
<collectingCountry id="F3693FB93449F322AA726195FD5DF979" box="[554,691,1719,1744]" name="Madagascar" pageId="6" pageNumber="271">Madagascar</collectingCountry>
, Ihorombe Region.
</paragraph>
</subSubSection>
</treatment>
</document>