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<document id="70F7964A672967DC0A240255E449D66A" ID-CLB-Dataset="7176" ID-DOI="10.5252/zoosystema2021v43a10" ID-GBIF-Dataset="120db7ca-e885-4d5e-bc3d-83af0671f8d1" ID-ISSN="1638-9387" ID-Zenodo-Dep="4720674" ID-ZooBank="urn:lsid:zoobank.org:pub:2763DA07-4D8F-4CA2-BB63-E5BC26470296" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445312570" checkinUser="felipe" docAuthor="Drolshagen, Bastian &amp; Bäckstam, Christian M." docDate="2021" docId="0386CD35FF89FFB576B5FDEA47AB90BE" docLanguage="en" docName="Zoosystema.43.10.163-196.pdf.imf" docOrigin="Zoosystema 43 (10)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Linothele septentrionalis Drolshagen &amp; Bäckstam 2021, n. sp." docType="treatment" docUuid="514DF6CE-3DC9-4416-B71B-1C60C698EBEB" docUuidSource="ZooBank" docVersion="8" lastPageNumber="182" masterDocId="FFBFB54DFF9CFFA0757CFF9F44099755" masterDocTitle="A taxonomic review of the mygalomorph spider genus Linothele Karsch, 1879 (Araneae, Dipluridae)" masterLastPageNumber="196" masterPageNumber="163" pageNumber="182" updateTime="1698937603326" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9A9AD8A0FE7B41745520396A51BC3442">A taxonomic review of the mygalomorph spider genus Linothele Karsch, 1879 (Araneae, Dipluridae)</mods:title>
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<mods:roleTerm id="12D683679E39421F40FFDAE47B321232">Author</mods:roleTerm>
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<mods:namePart id="27613069AF2CB027C9A3FA45AD93857F">Drolshagen, Bastian</mods:namePart>
<mods:affiliation id="D19699E031AE1B6C921A730131235EF4">Kolberger Strasse 12 F, 76139 Karlsruhe (Germany) bdrolshagen @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="83CAFDE4384181A531337C4F3C6103C2" type="email">bdrolshagen@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CF72655196F41E2EFB2BD50163B9E9A6">Bäckstam, Christian M.</mods:namePart>
<mods:affiliation id="94B62ACC295DDB23425C624A5490358F">Swedish Museum of Natural History, Department of Zoology, Box 50007, 104 05 Stockholm (Sweden) christian. backstam @ nrm. se</mods:affiliation>
<mods:nameIdentifier id="8811D4C744B18B2B1F23EA11BA4E7A16" type="email">backstam@nrm.se</mods:nameIdentifier>
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<mods:title id="81ED05C83DF9F46D0B0FC86314E557EB">Zoosystema</mods:title>
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<mods:date id="E6BCE7A99CCF884C56DDB5ECC0A5DC7B">2021</mods:date>
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<mods:number id="8B2C0C9FBB908DA4AC29CC8F44526D5D">43</mods:number>
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<treatment id="0386CD35FF89FFB576B5FDEA47AB90BE" ID-DOI="http://doi.org/10.5281/zenodo.4721106" ID-GBIF-Taxon="181600919" ID-Zenodo-Dep="4721106" LSID="urn:lsid:zoobank.org:act:514DF6CE-3DC9-4416-B71B-1C60C698EBEB" httpUri="http://treatment.plazi.org/id/0386CD35FF89FFB576B5FDEA47AB90BE" lastPageNumber="182" pageId="21" pageNumber="182">
<subSubSection id="C3352FA8FF89FFB576B5FDEA411A95C5" box="[969,1299,629,656]" pageId="21" pageNumber="182" type="nomenclature">
<paragraph id="8B907C23FF89FFB576B5FDEA411A95C5" blockId="21.[969,1299,629,688]" box="[969,1299,629,656]" pageId="21" pageNumber="182">
<heading id="D0D8CB4FFF89FFB576B5FDEA411A95C5" box="[969,1299,629,656]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="182" reason="2">
<taxonomicName id="4C2F07A0FF89FFB576B5FDEA40D895C5" ID-CoL="98YY3" authority="Drolshagen &amp; Bäckstam, 2021" authorityName="Drolshagen &amp; Bäckstam" authorityYear="2021" box="[969,1233,629,656]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="182" phylum="Arthropoda" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="B95BA031FF89FFB576B5FDEA40D895C5" bold="true" box="[969,1233,629,656]" italics="true" pageId="21" pageNumber="182">Linothele septentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2681D4AFF89FFB571A4FDE9411A95C5" box="[1240,1299,630,656]" pageId="21" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57164FD09419095B1" pageId="21" pageNumber="182" type="description">
<paragraph id="8B907C23FF89FFB57164FD0940CD95E5" blockId="21.[969,1299,629,688]" box="[1048,1220,662,688]" pageId="21" pageNumber="182">
(
<figureCitation id="131460A6FF89FFB57163FD09407895E5" box="[1055,1137,662,688]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Figs 13</figureCitation>
;
<figureCitation id="131460A6FF89FFB57101FD0940B595E5" box="[1149,1212,662,688]" captionStart="FIG" captionStartId="27.[134,144,1434,1452]" captionTargetBox="[221,1378,215,1382]" captionTargetId="graphics-856@27.[718,907,630,966]" captionTargetPageId="27" captionText="FIG. 17. — Patterns on opisthosoma in Linothele Karsch, 1879: A-E, L. curvitarsis Karsch, 1879; F-H, L. macrothelifera Strand, 1908; I, J, L. septentrionalis n. sp.; K-M, L. spinosa n. sp.; A, L. curvitarsis indet. holotype (ZMB Arach-458), dorsal view; B, C, L. curvitarsis male (NHRS-KASI 000000032); B, dorsal view; C, ventral view; D, Diplura soricina Simon, 1889 female (MNHN (339)), dorsal view; E, Diplura soricina male (MNHN (339)), dorsal view; F, L. macrothelifera imm. male (MCZ-76020), dorsal view; G, H, L. macrothelifera undet. (MCZ-76021); G, dorsal view; H, ventral view; I, J, L. septentrionalis n. sp. female holotype (MCZ- 76012); I, dorsal view; J, ventral view; K-M, L. spinosa n. sp. male holotype (NHRS-KASI 000000036); K, dorsal view; L, ventral view; M, L. spinosa n. sp. female paratype (NHRS-KASI 000000037), dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4720732" httpUri="https://zenodo.org/record/4720732/files/figure.png" pageId="21" pageNumber="182">17I, J</figureCitation>
)
</paragraph>
<paragraph id="8B907C23FF89FFB5763FFD4D419095B1" blockId="21.[835,1433,722,740]" box="[835,1433,722,740]" pageId="21" pageNumber="182">
<uri id="FFBE7021FF89FFB5763FFD4D419095B1" box="[835,1433,722,740]" pageId="21" pageNumber="182">
urn:lsid:zoobank.org:act:
<uuid id="FF8946F6FF89FFB57170FD4D419095B1" box="[1036,1433,722,740]" pageId="21" pageNumber="182">514DF6CE-3DC9-4416-B71B-1C60C698EBEB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651FC9047BD9415" pageId="21" pageNumber="182" type="materials_examined">
<paragraph id="8B907C23FF89FFB57651FC9047BD9415" blockId="21.[813,1455,782,833]" pageId="21" pageNumber="182">
TYPE MATERIAL. —
<materialsCitation id="3B47767EFF89FFB5768EFC9147BD9415" ID-GBIF-Occurrence="3091159372" accessionNumber="MCZ 76012" collectionCode="MCZ" collectorName="Nathan Banks" country="Bahamas" location="Bahamas" pageId="21" pageNumber="182" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B95BA031FF89FFB5768EFC9140C39473" bold="true" box="[1010,1226,782,806]" pageId="21" pageNumber="182">
<typeStatus id="5494C281FF89FFB5768EFC91405A9473" box="[1010,1107,782,806]" pageId="21" pageNumber="182" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F3383CB3FF89FFB57123FC9140B09473" box="[1119,1209,782,806]" name="Bahamas" pageId="21" pageNumber="182">Bahamas</collectingCountry>
</emphasis>
<specimenCount id="9D29B7AAFF89FFB571AEFC8E40EA9470" box="[1234,1251,785,805]" pageId="21" pageNumber="182" type="female"></specimenCount>
;
<collectorName id="26DA19F5FF89FFB5718EFC9141769473" box="[1266,1407,782,806]" pageId="21" pageNumber="182">Nathan Banks</collectorName>
leg.; MCZ 76012.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651FCFD407694FB" pageId="21" pageNumber="182" type="etymology">
<paragraph id="8B907C23FF89FFB57651FCFD407694FB" blockId="21.[813,1456,865,942]" pageId="21" pageNumber="182">
ETYMOLOGY. — The specific epithet is a Latin adjective, meaning northern and refers to the most northern locality in
<taxonomicName id="4C2F07A0FF89FFB57055FCE3418994C1" authorityName="Karsch" authorityYear="1879" box="[1321,1408,892,916]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF89FFB57055FCE3418994C1" box="[1321,1408,892,916]" italics="true" pageId="21" pageNumber="182">Linothele</emphasis>
</taxonomicName>
zoogeography; the gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651FC50404394B2" box="[813,1098,975,999]" pageId="21" pageNumber="182" type="materials_examined">
<paragraph id="8B907C23FF89FFB57651FC50404394B2" blockId="21.[813,1098,975,999]" box="[813,1098,975,999]" pageId="21" pageNumber="182">
<typeStatus id="5494C281FF89FFB57651FC50475494B0" box="[813,861,975,998]" pageId="21" pageNumber="182">TYPE</typeStatus>
LOCALITY. —
<collectingCountry id="F3383CB3FF89FFB57697FC50404C94B2" box="[1003,1093,975,999]" name="Bahamas" pageId="21" pageNumber="182">Bahamas</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651FB974149934A" box="[813,1344,1031,1056]" pageId="21" pageNumber="182" type="distribution">
<paragraph id="8B907C23FF89FFB57651FB974149934A" blockId="21.[813,1344,1031,1056]" box="[813,1344,1031,1056]" pageId="21" pageNumber="182">
DISTRIBUTION. — Only known from the
<typeStatus id="5494C281FF89FFB571B8FB9740E79375" box="[1220,1262,1032,1056]" pageId="21" pageNumber="182">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651FBDE405893FD" pageId="21" pageNumber="182" type="diagnosis">
<paragraph id="8B907C23FF89FFB57651FBDE405893FD" blockId="21.[813,1456,1088,1192]" pageId="21" pageNumber="182">
DIAGNOSIS. — The female of
<taxonomicName id="4C2F07A0FF89FFB5712CFBDF4138930D" authority="Drolshagen &amp; Bäckstam, 2021" authorityName="Drolshagen &amp; Bäckstam" authorityYear="2021" box="[1104,1329,1088,1112]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="182" phylum="Arthropoda" rank="species" species="septentrionalis" status="sp. nov.">
<emphasis id="B95BA031FF89FFB5712CFBDF4138930D" box="[1104,1329,1088,1112]" italics="true" pageId="21" pageNumber="182">Linothele septentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2681D4AFF89FFB57044FBDE4164930D" box="[1336,1389,1089,1112]" pageId="21" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
differs from those of all other
<taxonomicName id="4C2F07A0FF89FFB57178FBC440539326" authorityName="Karsch" authorityYear="1879" box="[1028,1114,1115,1139]" class="Arachnida" family="Dipluridae" genus="Linothele" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B95BA031FF89FFB57178FBC440539326" box="[1028,1114,1115,1139]" italics="true" pageId="21" pageNumber="182">Linothele</emphasis>
</taxonomicName>
by the combined presence of maculae (
<figureCitation id="131460A6FF89FFB57628FBEA47A393D8" box="[852,938,1141,1165]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Fig. 13B</figureCitation>
), presence of a pattern on the sternum and absence of maxillary cuspules (
<figureCitation id="131460A6FF89FFB57690FB0F404C93FD" box="[1004,1093,1168,1192]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Fig. 13A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651FB4B408490DE" pageId="21" pageNumber="182" type="description">
<paragraph id="8B907C23FF89FFB57651FB4B47B493B9" blockId="21.[813,957,1236,1325]" box="[813,957,1236,1262]" pageId="21" pageNumber="182">DESCRIPTION</paragraph>
<paragraph id="8B907C23FF89FFB57651FB6C476B9258" blockId="21.[813,957,1236,1325]" box="[813,866,1267,1293]" pageId="21" pageNumber="182">
<emphasis id="B95BA031FF89FFB57651FB6C476B9258" box="[813,866,1267,1293]" italics="true" pageId="21" pageNumber="182">Male</emphasis>
</paragraph>
<paragraph id="8B907C23FF89FFB57651FA8C47AB9278" blockId="21.[813,957,1236,1325]" box="[813,930,1299,1325]" pageId="21" pageNumber="182">Unknown.</paragraph>
<paragraph id="8B907C23FF89FFB57651FACC47EB9238" blockId="21.[811,1457,1363,1932]" box="[813,994,1363,1389]" pageId="21" pageNumber="182">
<emphasis id="B95BA031FF89FFB57651FACC47EB9238" box="[813,994,1363,1389]" italics="true" pageId="21" pageNumber="182">
Female (
<typeStatus id="5494C281FF89FFB576F8FACC47D29238" box="[900,987,1363,1389]" pageId="21" pageNumber="182" type="holotype">holotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="8B907C23FF89FFB57651FAEC408490DE" blockId="21.[811,1457,1363,1932]" pageId="21" pageNumber="182">
CL = 3.9. CT = 11. MC = 0. Colouration in alcohol: Prosoma, chelicerae, legs and pedipalps pale, yellow; sternum (length: 2.0) dark, with bright longitudinal pattern (
<figureCitation id="131460A6FF89FFB57197FA2C41419298" box="[1259,1352,1459,1485]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Fig. 13A</figureCitation>
); opisthosoma with distinct mid-dorsal pattern, consisting of quadrate spots anteriorly, which become more rectangular posteriorly, or can be interconnected, forming longitudinal lines, laterally with several spots; ventrally with longitudinal lines and spots (
<figureCitation id="131460A6FF89FFB5760CF9CD47D79138" box="[880,990,1618,1645]" captionStart="FIG" captionStartId="27.[134,144,1434,1452]" captionTargetBox="[221,1378,215,1382]" captionTargetId="graphics-856@27.[718,907,630,966]" captionTargetPageId="27" captionText="FIG. 17. — Patterns on opisthosoma in Linothele Karsch, 1879: A-E, L. curvitarsis Karsch, 1879; F-H, L. macrothelifera Strand, 1908; I, J, L. septentrionalis n. sp.; K-M, L. spinosa n. sp.; A, L. curvitarsis indet. holotype (ZMB Arach-458), dorsal view; B, C, L. curvitarsis male (NHRS-KASI 000000032); B, dorsal view; C, ventral view; D, Diplura soricina Simon, 1889 female (MNHN (339)), dorsal view; E, Diplura soricina male (MNHN (339)), dorsal view; F, L. macrothelifera imm. male (MCZ-76020), dorsal view; G, H, L. macrothelifera undet. (MCZ-76021); G, dorsal view; H, ventral view; I, J, L. septentrionalis n. sp. female holotype (MCZ- 76012); I, dorsal view; J, ventral view; K-M, L. spinosa n. sp. male holotype (NHRS-KASI 000000036); K, dorsal view; L, ventral view; M, L. spinosa n. sp. female paratype (NHRS-KASI 000000037), dorsal view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4720732" httpUri="https://zenodo.org/record/4720732/files/figure.png" pageId="21" pageNumber="182">Fig. 17I, J</figureCitation>
); maculae present, see
<figureCitation id="131460A6FF89FFB571B2F9CD41419139" box="[1230,1352,1618,1644]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Figure 13B</figureCitation>
. Clypeus: narrow. Sternum, labium and maxillae: see
<figureCitation id="131460A6FF89FFB57187F9ED417F91D9" box="[1275,1398,1650,1676]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Figure 13A</figureCitation>
. Palp measurements: 7.6 (2.5,1.4,1.7,2.0). Leg measurements: I: 13.0 (3.3, 1.5, 3.0, 2.9, 2.3); II: 10.8 (2.9, 1.5, 2.0, 2.6, 1.8); III: 10.6 (2.8, 1.3, 2.1, 2.4, 2.0); IV: 14.9 (2.4, 3.8, 3.2, 1.5, 4.0). Leg formula: 4123. Scopula divided. Preening-combs absent. Leg tarsi integral. Spinnerets: PMS: 1.3, separated by less than length of segment (0.7); PLS: 5.9 (1.6, 1.9, 2.4), apical segments rigid. Spermatheca: consisting of two stalks with basal lobes, see
<figureCitation id="131460A6FF89FFB57177F8EE408490DE" box="[1035,1165,1905,1931]" captionStart="FIG" captionStartId="22.[132,143,797,815]" captionTargetBox="[203,1392,221,753]" captionTargetPageId="22" captionText="FIG. 13. — Linothele septentrionalis n. sp: A-C, L. septentrionalis n. sp. female holotype (MCZ-76012); A, maxilla, labium and sternum, ventral view; B, left palp, retrolateral view; C, spermatheca, dorsal view. Scale bars: A, B, 1 mm; C, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720718" httpUri="https://zenodo.org/record/4720718/files/figure.png" pageId="21" pageNumber="182">Figure 13C.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3352FA8FF89FFB57651F82D47AB90BE" pageId="21" pageNumber="182" type="description">
<paragraph id="8B907C23FF89FFB57651F82D47FB909F" blockId="21.[813,1010,1970,2027]" box="[813,1010,1970,1996]" pageId="21" pageNumber="182">NATURAL HISTORY</paragraph>
<paragraph id="8B907C23FF89FFB57651F84E47AB90BE" blockId="21.[813,1010,1970,2027]" box="[813,930,2001,2027]" pageId="21" pageNumber="182">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>