treatments-xml/data/03/86/A4/0386A43AEF46A2048B85F901FA13F872.xml

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<mods:title id="5D888B802CCB3C473468302D97312AAB">Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus</mods:title>
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Key to male imagos of 
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<emphasis id="B95BC93EEF46A2078AA1F901FDB5F96A" bold="true" box="[435,573,1733,1759]" italics="true" pageId="49" pageNumber="350">Campsurus</emphasis>
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1 Pedestals touching each other basally or separated by only a short distance, less than 0.7× the width of a pedestal (e.g. 
<figureCitation id="131409A9EF46A2078E5AF8C8FA10F894" box="[1352,1432,1804,1825]" captionStart="FIGURES 35 – 50" captionStartId="10.[151,264,1859,1882]" captionTargetBox="[218,1378,208,1822]" captionTargetId="figure@10.[212,1381,207,1822]" captionTargetPageId="10" captionText="FIGURES 35 – 50. Campsurus essequibo, imago. Holotype male genitalia: 35, general v. v.; 36, detail of penes, v. v. Female adult: 37, abdominal sternum VIII; 38, detail of sockets. Male imago: 39 – 42, genitalia, variations, v. v.; 43, genitalia after treatment with KOH; 44 – 47, genitalia, variations, l. v.; 48, male mesofurcasternum. SEM photographs: 49, genitalia, v. v. (with detail of microlepides); 50, detail of penes, v. v." httpUri="https://zenodo.org/record/838446/files/figure.png" pageId="49" pageNumber="350">Figs. 35</figureCitation>
, 
<figureCitation id="131409A9EF46A2078BF3F8EDFF73F88B" box="[225,251,1833,1854]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="49" pageNumber="350">75</figureCitation>
, 
<figureCitation id="131409A9EF46A2078A17F8EDFE96F88B" box="[261,286,1833,1854]" captionStart="FIGURES 85 – 93" captionStartId="24.[151,264,1909,1932]" captionTargetBox="[168,1418,342,1885]" captionTargetId="figure@24.[163,1418,337,1885]" captionTargetPageId="24" captionText="FIGURES 85 – 93. Campsurus povilla sp. nov., male imago: 85, genitalia, v. v.; 86, detail of pedestals, v. v.; 87, detail of penes, v. v. Nymph: 88, left mandibular tusk, detail of inner tubercles; 89, left mandible, occlusal view (the basal part and molae were displaced during slide preparation; 90, same, detail of apex; 91, right mandible, occlusal view. Female abdominal sternum VIII with detail of socket: 92, C. povilla sp. nov.; 93, C. zunigae sp. nov." httpUri="https://zenodo.org/record/838456/files/figure.png" pageId="49" pageNumber="350">85</figureCitation>
)..............................................................................................2
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- Pedestals distinctly separated at the middle, by a distance equal to or greater than the width of a pedestal (e.g. 
<figureCitation id="131409A9EF46A2078E12F882FACAF8EE" box="[1280,1346,1862,1883]" captionStart="FIGURES 1 – 12" captionStartId="4.[151,264,1834,1857]" captionTargetBox="[229,1357,377,1813]" captionTargetId="figure@4.[229,1357,373,1813]" captionTargetPageId="4" captionText="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." httpUri="https://zenodo.org/record/838440/files/figure.png" pageId="49" pageNumber="350">Figs. 1</figureCitation>
, 
<figureCitation id="131409A9EF46A2078E5CF882FAEEF8EE" box="[1358,1382,1862,1883]" captionStart="FIGURES 25 – 34" captionStartId="7.[151,264,1939,1962]" captionTargetBox="[212,1363,202,1896]" captionTargetId="figure@7.[212,1363,193,1896]" captionTargetPageId="7" captionText="FIGURES 25 – 34. Campsurus salobra sp. nov., adult: 25, male genitalia, v. v.; 26, same, l. v.; 27, female sternum VIII; 28, detail of female sockets. SEM photographs: 29, male genitalia, v. v. (with detail of microtrichiae); 30, apical portion of penes, v. v.; 31, female sterna VIII – X; 32, female sockets (with detail of microtrichia); 33, egg, convex side; 34, detail of capped pole." httpUri="https://zenodo.org/record/838444/files/figure.png" pageId="49" pageNumber="350">25</figureCitation>
, 
<figureCitation id="131409A9EF46A2078E60F882FA10F8EE" box="[1394,1432,1862,1883]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="49" pageNumber="350">100</figureCitation>
, 
<figureCitation id="131409A9EF46A2078BF3F8A0FE8EF8CC" box="[225,262,1892,1913]" captionStart="FIGURES 117 – 122" captionStartId="32.[151,264,1660,1683]" captionTargetBox="[196,1392,193,1620]" captionTargetId="figure@32.[196,1392,193,1639]" captionTargetPageId="32" captionText="FIGURES 117 – 122. Campsurus inusitatus sp. nov., male imago, SEM photographs: 117, genitalia, v. v. (g? = gonopore, mf = median furrow of styliger, ml = main lobe of penes, sl = secondary lobe); left mandible, v. v.; 118, detail of microlepides on sternum IX; 119, detail of penes and pedestals (note that penean membrane was broken during preparation); 120, detail of penean lobes, v. v. Line drawings: 121, male genitalia, v. v.; 122, male terminalia, l. v. (t. IX – t. X = tergum IX and X, s. IX = sternum IX, c = cercus base, p = paraproct, ml = main lobe of penes, sl = secondary lobe)." httpUri="https://zenodo.org/record/838466/files/figure.png" pageId="49" pageNumber="350">117</figureCitation>
, 
<figureCitation id="131409A9EF46A2078A02F8A0FEBEF8CC" box="[272,310,1892,1913]" captionStart="FIGURES 138 – 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 – 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="49" pageNumber="350">138</figureCitation>
, 
<figureCitation id="131409A9EF46A2078A52F8A0FEEDF8CC" box="[320,357,1892,1913]" captionStart="FIGURES 151 – 162" captionStartId="42.[151,264,1947,1970]" captionTargetBox="[239,1346,202,1918]" captionTargetId="figure@42.[239,1346,195,1918]" captionTargetPageId="42" captionText="FIGURES 151 – 162. Campsurus litaninensis, adult: 151, male genitalia, v. v.; 152 – 153, details of pedestal, v. v; 154, genitalia, l. v.; 155 – 156, details of penes, l. v.; 157, detail of female socket, v. v.; 158, detail of pedestal, v. v. SEM: 159, male genitalia, v. v.; 160, detail of microtrichia on male sternum IX; 161, detail of pedestal, v. v.; 162, detail of penes, v. v." httpUri="https://zenodo.org/record/838476/files/figure.png" pageId="49" pageNumber="350">151</figureCitation>
).......................................................................................12
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<paragraph id="8B90152CEF46A2078B85F845FA14F823" blockId="49.[151,1436,1804,2030]" box="[151,1436,1921,1942]" pageId="49" pageNumber="350">
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2(1) Pedestal with inner distal corner always more developed than outer corner; outer corner sometimes absent (e.g. 
<figureCitation id="131409A9EF46A2078E1DF845FADDF823" box="[1295,1365,1921,1942]" captionStart="FIGURES 85 – 93" captionStartId="24.[151,264,1909,1932]" captionTargetBox="[168,1418,342,1885]" captionTargetId="figure@24.[163,1418,337,1885]" captionTargetPageId="24" captionText="FIGURES 85 – 93. Campsurus povilla sp. nov., male imago: 85, genitalia, v. v.; 86, detail of pedestals, v. v.; 87, detail of penes, v. v. Nymph: 88, left mandibular tusk, detail of inner tubercles; 89, left mandible, occlusal view (the basal part and molae were displaced during slide preparation; 90, same, detail of apex; 91, right mandible, occlusal view. Female abdominal sternum VIII with detail of socket: 92, C. povilla sp. nov.; 93, C. zunigae sp. nov." httpUri="https://zenodo.org/record/838456/files/figure.png" pageId="49" pageNumber="350">Fig. 85</figureCitation>
).....3
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<paragraph id="8B90152CEF46A2078B85F85AFA1DF806" blockId="49.[151,1436,1804,2030]" box="[151,1429,1950,1971]" pageId="49" pageNumber="350">
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- Pedestal with outer distal corner more developed than inner corner, generally forming a long parastylus (e.g. 
<figureCitation id="131409A9EF46A2078FE7F85AFAEFF806" box="[1269,1383,1950,1971]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="49" pageNumber="350">Figs. 53–59</figureCitation>
)..6
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<paragraph id="8B90152CEF46A2078B85F878FA14F864" blockId="49.[151,1436,1804,2030]" box="[151,1436,1979,2001]" pageId="49" pageNumber="350">
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3(2) FW &lt;
<quantity id="4CD7B8C9EF46A2078A0CF878FED0F864" box="[286,344,1980,2001]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="49" pageNumber="350" unit="mm" value="7.0">7 mm</quantity>
; penes blade-like, not curved (
<figureCitation id="131409A9EF46A207896FF878FD4BF864" box="[637,707,1980,2001]" captionStart="FIGURES 85 – 93" captionStartId="24.[151,264,1909,1932]" captionTargetBox="[168,1418,342,1885]" captionTargetId="figure@24.[163,1418,337,1885]" captionTargetPageId="24" captionText="FIGURES 85 – 93. Campsurus povilla sp. nov., male imago: 85, genitalia, v. v.; 86, detail of pedestals, v. v.; 87, detail of penes, v. v. Nymph: 88, left mandibular tusk, detail of inner tubercles; 89, left mandible, occlusal view (the basal part and molae were displaced during slide preparation; 90, same, detail of apex; 91, right mandible, occlusal view. Female abdominal sternum VIII with detail of socket: 92, C. povilla sp. nov.; 93, C. zunigae sp. nov." httpUri="https://zenodo.org/record/838456/files/figure.png" pageId="49" pageNumber="350">Fig. 85</figureCitation>
)............................................ .. 
<taxonomicName id="4C2F6EAFEF46A2078FF9F878FAC1F864" ID-CoL="6V9V" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1259,1353,1980,2001]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="350" phylum="Arthropoda" rank="species" species="povilla" status="sp. nov.">
<emphasis id="B95BC93EEF46A2078FF9F878FAC1F864" box="[1259,1353,1980,2001]" italics="true" pageId="49" pageNumber="350">C. povilla</emphasis>
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<emphasis id="B95BC93EEF46A2078E5DF87FFA14F864" bold="true" box="[1359,1436,1979,2001]" pageId="49" pageNumber="350">sp. nov.</emphasis>
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- FW&gt; 
<quantity id="4CD7B8C9EF46A2078A0BF81DFEDAF85B" box="[281,338,2009,2030]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="49" pageNumber="350" unit="mm" value="9.0">9 mm</quantity>
; penes variable, never blade-like, generally cylindrical or conical and always curved ventrally or medially.... 4
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4(3) Penes with only one finger-like lobe, ventrally curved.......................................................................................... 
<taxonomicName id="4C2F6EAFEF45A2048974FF71FD48FF7F" ID-CoL="QF4Q" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[614,704,181,202]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="violaceus">
<emphasis id="B95BC93EEF45A2048974FF71FD48FF7F" box="[614,704,181,202]" italics="true" pageId="50" pageNumber="351">violaceus</emphasis>
</taxonomicName>
group (see 
<bibRefCitation id="EFBE68DDEF45A2048823FF71FB83FF7F" box="[817,1035,181,202]" pageId="50" pageNumber="351" refString="Molineri, C., Salles, F. F. &amp; Emmerich, D. (2015 a) Revision of Campsurus violaceus species group (Ephemeroptera: Polymitarcyidae) with new synonymies and nomina dubia in Campsurus Eaton, 1868. Zootaxa, 3920 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 3920.1.3" type="journal article">
Molineri 
<emphasis id="B95BC93EEF45A2048899FF71FC34FF7F" box="[907,956,181,202]" italics="true" pageId="50" pageNumber="351">et al.</emphasis>
(2015a)
</bibRefCitation>
for identification of the six valid species)
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FF16FA13FF52" blockId="50.[151,1436,151,1991]" box="[151,1435,210,231]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FF16FA13FF52" box="[151,1435,210,231]" pageId="50" pageNumber="351">- Penes variable, but not finger-like, formed by two lobes (the ventral one may be reduced but always is visible)...........5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FF2BFA14FEE9" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FF2BFA14FE97" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FF2BFA14FE97" pageId="50" pageNumber="351">
5(4) Penes flat, not curved ventrally, and located in the same plane as forceps......................................................................... 
<emphasis id="B95BC93EEF45A204895BFEC9FD17FE97" box="[585,671,269,290]" italics="true" pageId="50" pageNumber="351">albifilum</emphasis>
group (see Molineri &amp; Salles (2013) for identification of the five valid species)
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FEEEFA14FEE9" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FEEEFA14FEE9" pageId="50" pageNumber="351">
- Penes curved ventrally, each lobe twisted outwards........................................................................................ 
<taxonomicName id="4C2F6EAFEF45A2048923FE83FDE2FEE9" ID-CoL="699XL" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[561,618,327,348]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B95BC93EEF45A2048923FE83FDE2FEE9" box="[561,618,327,348]" italics="true" pageId="50" pageNumber="351">major</emphasis>
</taxonomicName>
group (see 
<bibRefCitation id="EFBE68DDEF45A20489C9FE83FC70FEE9" author="Molineri" box="[731,1016,327,348]" pageId="50" pageNumber="351" refString="Molineri, C. &amp; Emmerich, D. (2010) New species and new stage descriptions of Campsurus major species-group (Polymitarcyidae, Campsurinae), with first report of silk-case construction in a mayfly nymphs. Aquatic Insects, 32, 265 - 280." type="journal article" year="2010">Molineri &amp; Emmerich (2010)</bibRefCitation>
for identification of the three valid species)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FEA1FA14FE22" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FEA1FA14FECF" blockId="50.[151,1436,151,1991]" box="[151,1436,356,378]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FEA1FA14FECF" box="[151,1436,356,378]" pageId="50" pageNumber="351">
6(2) Pedestal very long, subrectangular; parastylus relatively short and rounded (
<figureCitation id="131409A9EF45A20488ABFEA1FBA5FECF" box="[953,1069,357,378]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">Figs. 75–76</figureCitation>
)............... 
<taxonomicName id="4C2F6EAFEF45A2048FEAFEA1FAC0FECF" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1272,1352,357,378]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="janae" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FEAFEA1FAC0FECF" box="[1272,1352,357,378]" italics="true" pageId="50" pageNumber="351">C. janae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFEA0FA14FECF" box="[1359,1436,356,378]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFEA0FA14FECF" bold="true" box="[1359,1436,356,378]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FE46FA14FE22" blockId="50.[151,1436,151,1991]" box="[151,1436,386,407]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FE46FA14FE22" box="[151,1436,386,407]" pageId="50" pageNumber="351">
- Pedestal large, but not as above; parastylus longer than above (e.g. 
<figureCitation id="131409A9EF45A2048877FE46FC5FFE22" box="[869,983,386,407]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 51–52</figureCitation>
, 
<figureCitation id="131409A9EF45A20488F0FE46FC76FE22" box="[994,1022,386,407]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">72</figureCitation>
).................................7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FE5BFA13FE67" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FE5BFA14FE00" blockId="50.[151,1436,151,1991]" box="[151,1436,415,437]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FE5BFA14FE00" box="[151,1436,415,437]" pageId="50" pageNumber="351">
7(6) Penes large, with ear-like base and outer margin granulated (
<figureCitation id="131409A9EF45A2048837FE5BFC10FE01" box="[805,920,415,436]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">Figs. 71, 73</figureCitation>
).......................... 
<taxonomicName id="4C2F6EAFEF45A2048FCEFE5BFAC0FE01" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1244,1352,415,436]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="pereirae" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FCEFE5BFAC0FE01" box="[1244,1352,415,436]" italics="true" pageId="50" pageNumber="351">C. pereirae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFE5BFA14FE00" box="[1359,1436,415,437]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFE5BFA14FE00" bold="true" box="[1359,1436,415,437]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FE79FA13FE67" blockId="50.[151,1436,151,1991]" box="[151,1435,445,466]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FE79FA13FE67" box="[151,1435,445,466]" pageId="50" pageNumber="351">
- Penes variable, but never with granulated outer margin (e.g., 
<figureCitation id="131409A9EF45A2048839FE79FC16FE67" box="[811,926,445,466]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 53–59</figureCitation>
, 
<figureCitation id="131409A9EF45A20488BAFE79FC62FE67" box="[936,1002,445,466]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">68–70</figureCitation>
).................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FE1EFA14FD9F" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FE1EFA13FDB9" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FE1EFA13FDB9" pageId="50" pageNumber="351">
8(7) Forceps very short, surpassing the apex of parastylus at a distance less than half of forceps length (
<figureCitation id="131409A9EF45A2048F80FE1EFA81FE5A" box="[1170,1289,474,495]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 51–53</figureCitation>
)............................................................................................... 
<taxonomicName id="4C2F6EAFEF45A2048FD9FE33FA13FDB9" ID-CoL="QF3S" authority="Traver" authorityName="Traver ." authorityYear="1947" box="[1227,1435,503,524]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="cuyuniensis">
<emphasis id="B95BC93EEF45A2048FD9FE33FADEFDB9" box="[1227,1366,503,524]" italics="true" pageId="50" pageNumber="351">C. cuyuniensis</emphasis>
Traver
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FDD1FA14FD9F" blockId="50.[151,1436,151,1991]" box="[151,1436,533,554]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FDD1FA14FD9F" box="[151,1436,533,554]" pageId="50" pageNumber="351">
- Forceps longer, surpassing the apex of parastylus by a distance at least 0.6× the forceps length (
<figureCitation id="131409A9EF45A2048F66FDD1FB60FD9F" box="[1140,1256,533,554]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 54, 56</figureCitation>
)............. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FDF6FA13FD37" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FDF6FA14FDFD" blockId="50.[151,1436,151,1991]" box="[151,1436,562,584]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FDF6FA14FDFD" box="[151,1436,562,584]" pageId="50" pageNumber="351">
9(8) Penes with spatulate apex (
<figureCitation id="131409A9EF45A2048AF7FDF6FDA4FDF2" box="[485,556,562,583]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Fig. 55</figureCitation>
); parastylus apically rounded (
<figureCitation id="131409A9EF45A2048841FDF6FC41FDF2" box="[851,969,562,583]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 54–55</figureCitation>
)..................... 
<taxonomicName id="4C2F6EAFEF45A2048FD4FDF6FAC1FDF2" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1222,1353,562,583]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="nessimiani" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FD4FDF6FAC1FDF2" box="[1222,1353,562,583]" italics="true" pageId="50" pageNumber="351">C. nessimiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFDF6FA14FDFD" box="[1359,1436,562,584]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFDF6FA14FDFD" bold="true" box="[1359,1436,562,584]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FD8BFA13FD37" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FD8BFA13FD37" pageId="50" pageNumber="351">
- Main lobe of penes apically pointed (
<figureCitation id="131409A9EF45A2048958FD8BFD37FDD1" box="[586,703,591,612]" captionStart="FIGURES 35 – 50" captionStartId="10.[151,264,1859,1882]" captionTargetBox="[218,1378,208,1822]" captionTargetId="figure@10.[212,1381,207,1822]" captionTargetPageId="10" captionText="FIGURES 35 – 50. Campsurus essequibo, imago. Holotype male genitalia: 35, general v. v.; 36, detail of penes, v. v. Female adult: 37, abdominal sternum VIII; 38, detail of sockets. Male imago: 39 – 42, genitalia, variations, v. v.; 43, genitalia after treatment with KOH; 44 – 47, genitalia, variations, l. v.; 48, male mesofurcasternum. SEM photographs: 49, genitalia, v. v. (with detail of microlepides); 50, detail of penes, v. v." httpUri="https://zenodo.org/record/838446/files/figure.png" pageId="50" pageNumber="351">Figs. 36, 39</figureCitation>
, 
<figureCitation id="131409A9EF45A20489D8FD8BFD6DFDD1" box="[714,741,591,612]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">70</figureCitation>
); parastylus apically pointed (in lateral view, may be rounded in ventral view) (
<figureCitation id="131409A9EF45A2048A36FDA9FEFDFD37" box="[292,373,621,642]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 57</figureCitation>
, 
<figureCitation id="131409A9EF45A2048A6DFDA9FE10FD37" box="[383,408,621,642]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">69</figureCitation>
)...................................................................................10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FD4EFA13FD09" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FD4EFA13FD2A" blockId="50.[151,1436,151,1991]" box="[151,1435,650,671]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FD4EFA13FD2A" box="[151,1435,650,671]" pageId="50" pageNumber="351">
10(9) Main lobe of penes relatively long and slender (
<figureCitation id="131409A9EF45A20489B9FD4EFCEEFD2A" box="[683,870,650,671]" captionStart="FIGURES 35 – 50" captionStartId="10.[151,264,1859,1882]" captionTargetBox="[218,1378,208,1822]" captionTargetId="figure@10.[212,1381,207,1822]" captionTargetPageId="10" captionText="FIGURES 35 – 50. Campsurus essequibo, imago. Holotype male genitalia: 35, general v. v.; 36, detail of penes, v. v. Female adult: 37, abdominal sternum VIII; 38, detail of sockets. Male imago: 39 – 42, genitalia, variations, v. v.; 43, genitalia after treatment with KOH; 44 – 47, genitalia, variations, l. v.; 48, male mesofurcasternum. SEM photographs: 49, genitalia, v. v. (with detail of microlepides); 50, detail of penes, v. v." httpUri="https://zenodo.org/record/838446/files/figure.png" pageId="50" pageNumber="351">Figs. 35–36, 39–47</figureCitation>
).............................. 
<taxonomicName id="4C2F6EAFEF45A2048FCEFD4EFA13FD2A" ID-CoL="QF3Y" authority="Traver" authorityName="Traver" authorityYear="1947" box="[1244,1435,650,671]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="essequibo">
<emphasis id="B95BC93EEF45A2048FCEFD4EFADEFD2A" box="[1244,1366,650,671]" italics="true" pageId="50" pageNumber="351">C. essequibo</emphasis>
Traver
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FD63FA13FD09" blockId="50.[151,1436,151,1991]" box="[151,1435,679,700]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FD63FA13FD09" box="[151,1435,679,700]" pageId="50" pageNumber="351">
- Main lobe of penes stouter, main lobe with wider base (
<figureCitation id="131409A9EF45A20489E7FD63FCE1FD09" box="[757,873,679,700]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 56–59</figureCitation>
, 
<figureCitation id="131409A9EF45A2048861FD63FC39FD09" box="[883,945,679,700]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">68–70</figureCitation>
)...................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FD01FA1EFC87" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FD01FA14FD4D" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FD01FA14FD4D" pageId="50" pageNumber="351">
11(10) Penes relatively slender at base; parastylus becoming abruptly thinner on apical third in lateral view (
<figureCitation id="131409A9EF45A2048FB8FD01FAA9FD6F" box="[1194,1313,709,730]" captionStart="FIGURES 51 – 59" captionStartId="12.[151,264,1910,1933]" captionTargetBox="[172,1421,199,1890]" captionTargetId="figure@12.[172,1421,193,1890]" captionTargetPageId="12" captionText="FIGURES 51 – 59. male imagos. Campsurus cuyuniensis: 51, paratype forceps (CUIC 2434); 52, same but from another paratype; 53, male genitalia from French Guiana specimen (FAMU), v. v. Campsurus nessimiani sp. nov.: 54, genitalia, v. v.; 55, same, l. v. Campsurus dasilvai sp. nov.: 56, genitalia, v. v.; 57, same, l. v.; 58, genitalia, v. v. (SEM), with detail of microlepides on sternum IX; 59, details of penes, v. v. (SEM)." httpUri="https://zenodo.org/record/838448/files/figure.png" pageId="50" pageNumber="351">Figs. 56–59</figureCitation>
)............................................................................................... 
<taxonomicName id="4C2F6EAFEF45A2048FF3FD26FAC1FD42" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1249,1353,738,759]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="dasilvai" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FF3FD26FAC1FD42" box="[1249,1353,738,759]" italics="true" pageId="50" pageNumber="351">C. dasilvai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFD26FA14FD4D" box="[1359,1436,738,760]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFD26FA14FD4D" bold="true" box="[1359,1436,738,760]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FD3BFA1EFC87" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FD3BFA1EFC87" pageId="50" pageNumber="351">
- Penes relatively wider at base; parastylus becoming gradually thinner toward the apex in lateral view (
<figureCitation id="131409A9EF45A2048FA7FD3BFAA4FCA1" box="[1205,1324,767,788]" captionStart="FIGURES 68 – 76" captionStartId="18.[151,264,1825,1848]" captionTargetBox="[265,1305,209,1801]" captionTargetId="figure@18.[260,1326,193,1805]" captionTargetPageId="18" captionText="FIGURES 68 – 76. Male adults. Campsurus zunigae sp. nov.: 68, genitalia of imago, v. v.; 69, genitalia of subimago, l. v.; 70, detail of penes of imago, l. v. Campsurus pereirae sp. nov., imago: 71, genitalia, v. v.; 72, detail of pedestal, v. v.; 73, detail of penes, v. v.; 74, genitalia, l. v. (forceps omitted). Campsurus janae sp. nov., imago: 75, genitalia, v. v.: 76, detail of penes, l. v." httpUri="https://zenodo.org/record/838452/files/figure.png" pageId="50" pageNumber="351">Figs. 68–70</figureCitation>
).............................................................................................. 
<taxonomicName id="4C2F6EAFEF45A2048FCEFCD9FACBFC87" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1244,1347,797,818]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="zunigae" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FCEFCD9FACBFC87" box="[1244,1347,797,818]" italics="true" pageId="50" pageNumber="351">C. zunigae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5BFCD8FA1EFC87" box="[1353,1430,796,818]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5BFCD8FA1EFC87" bold="true" box="[1353,1430,796,818]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FCFEFA14FC3F" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FCFEFA13FCD9" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FCFEFA13FCD9" pageId="50" pageNumber="351">
12(1) Sternum IX strongly projected distally (surpassing pedestals and covering the base of penes) (
<figureCitation id="131409A9EF45A2048F76FCFEFB79FCFA" box="[1124,1265,826,847]" captionStart="FIGURES 151 – 162" captionStartId="42.[151,264,1947,1970]" captionTargetBox="[239,1346,202,1918]" captionTargetId="figure@42.[239,1346,195,1918]" captionTargetPageId="42" captionText="FIGURES 151 – 162. Campsurus litaninensis, adult: 151, male genitalia, v. v.; 152 – 153, details of pedestal, v. v; 154, genitalia, l. v.; 155 – 156, details of penes, l. v.; 157, detail of female socket, v. v.; 158, detail of pedestal, v. v. SEM: 159, male genitalia, v. v.; 160, detail of microtrichia on male sternum IX; 161, detail of pedestal, v. v.; 162, detail of penes, v. v." httpUri="https://zenodo.org/record/838476/files/figure.png" pageId="50" pageNumber="351">Figs. 151, 159</figureCitation>
).................................................................................................. 
<taxonomicName id="4C2F6EAFEF45A2048FD8FC93FA13FCD9" ID-CoL="QF46" authority="Spieth" authorityName="Spieth ." authorityYear="1943" box="[1226,1435,855,876]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="litaninensis">
<emphasis id="B95BC93EEF45A2048FD8FC93FAD0FCD9" box="[1226,1368,855,876]" italics="true" pageId="50" pageNumber="351">C. litaninensis</emphasis>
Spieth
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FCB1FA14FC3F" blockId="50.[151,1436,151,1991]" box="[151,1436,885,906]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FCB1FA14FC3F" box="[151,1436,885,906]" pageId="50" pageNumber="351">
- Sternum IX not projected as above, never surpassing the pedestals (
<figureCitation id="131409A9EF45A204887AFCB1FC4BFC3F" box="[872,963,885,906]" captionStart="FIGURES 117 – 122" captionStartId="32.[151,264,1660,1683]" captionTargetBox="[196,1392,193,1620]" captionTargetId="figure@32.[196,1392,193,1639]" captionTargetPageId="32" captionText="FIGURES 117 – 122. Campsurus inusitatus sp. nov., male imago, SEM photographs: 117, genitalia, v. v. (g? = gonopore, mf = median furrow of styliger, ml = main lobe of penes, sl = secondary lobe); left mandible, v. v.; 118, detail of microlepides on sternum IX; 119, detail of penes and pedestals (note that penean membrane was broken during preparation); 120, detail of penean lobes, v. v. Line drawings: 121, male genitalia, v. v.; 122, male terminalia, l. v. (t. IX – t. X = tergum IX and X, s. IX = sternum IX, c = cercus base, p = paraproct, ml = main lobe of penes, sl = secondary lobe)." httpUri="https://zenodo.org/record/838466/files/figure.png" pageId="50" pageNumber="351">Figs. 117</figureCitation>
, 
<figureCitation id="131409A9EF45A20488DCFCB1FC7CFC3F" box="[974,1012,885,906]" captionStart="FIGURES 123 – 128" captionStartId="34.[156,269,1818,1841]" captionTargetBox="[192,1394,445,1797]" captionTargetId="figure@34.[192,1394,445,1797]" captionTargetPageId="34" captionText="FIGURES 123 – 128. Campsurus indivisus, adult, SEM: 123, male terminalia, v. v.; 124, detail of pedestal, v. v. (p = pedestal, ic = inner corner); 125, detail of penes and pedestals, v. v. (bp = base of penes, g = gonopore, ml = main lobe of penes, sl = secondary lobe); 126, detail of apical portion of penes, v. v; 127, gonopores (g) at base of penes, v. v.; 128, microlepides on male sternum IX." httpUri="https://zenodo.org/record/838468/files/figure.png" pageId="50" pageNumber="351">123</figureCitation>
, 
<figureCitation id="131409A9EF45A20488ECFCB1FB0DFC3F" box="[1022,1157,885,906]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">100, 103, 107</figureCitation>
).....................13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FC56FA13FC71" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FC56FA13FC12" blockId="50.[151,1436,151,1991]" box="[151,1435,914,935]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FC56FA13FC12" box="[151,1435,914,935]" pageId="50" pageNumber="351">
13(12) Paraprocts large, clearly protruding as in 
<figureCitation id="131409A9EF45A204897EFC56FD4FFC12" box="[620,711,914,935]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 108</figureCitation>
, 
<figureCitation id="131409A9EF45A20489C3FC56FD7FFC12" box="[721,759,914,935]" captionStart="FIGURES 117 – 122" captionStartId="32.[151,264,1660,1683]" captionTargetBox="[196,1392,193,1620]" captionTargetId="figure@32.[196,1392,193,1639]" captionTargetPageId="32" captionText="FIGURES 117 – 122. Campsurus inusitatus sp. nov., male imago, SEM photographs: 117, genitalia, v. v. (g? = gonopore, mf = median furrow of styliger, ml = main lobe of penes, sl = secondary lobe); left mandible, v. v.; 118, detail of microlepides on sternum IX; 119, detail of penes and pedestals (note that penean membrane was broken during preparation); 120, detail of penean lobes, v. v. Line drawings: 121, male genitalia, v. v.; 122, male terminalia, l. v. (t. IX – t. X = tergum IX and X, s. IX = sternum IX, c = cercus base, p = paraproct, ml = main lobe of penes, sl = secondary lobe)." httpUri="https://zenodo.org/record/838466/files/figure.png" pageId="50" pageNumber="351">122</figureCitation>
, 
<figureCitation id="131409A9EF45A2048813FC56FA13FC12" box="[769,1435,914,935]" captionStart="FIGURES 123 – 128" captionStartId="34.[156,269,1818,1841]" captionTargetBox="[192,1394,445,1797]" captionTargetId="figure@34.[192,1394,445,1797]" captionTargetPageId="34" captionText="FIGURES 123 – 128. Campsurus indivisus, adult, SEM: 123, male terminalia, v. v.; 124, detail of pedestal, v. v. (p = pedestal, ic = inner corner); 125, detail of penes and pedestals, v. v. (bp = base of penes, g = gonopore, ml = main lobe of penes, sl = secondary lobe); 126, detail of apical portion of penes, v. v; 127, gonopores (g) at base of penes, v. v.; 128, microlepides on male sternum IX." httpUri="https://zenodo.org/record/838468/files/figure.png" pageId="50" pageNumber="351">123..................................................14</figureCitation>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FC6BFA13FC71" blockId="50.[151,1436,151,1991]" box="[151,1435,943,964]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FC6BFA13FC71" box="[151,1435,943,964]" pageId="50" pageNumber="351">- Paraprocts of usual size and not protruding................................................................19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FC09FA14FB8F" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FC09FA13FC4A" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FC09FA13FC4A" pageId="50" pageNumber="351">
14(13) Sternum IX with broad posteromedian emargination (
<figureCitation id="131409A9EF45A20489E3FC09FCC7FC57" box="[753,847,973,994]" captionStart="FIGURES 117 – 122" captionStartId="32.[151,264,1660,1683]" captionTargetBox="[196,1392,193,1620]" captionTargetId="figure@32.[196,1392,193,1639]" captionTargetPageId="32" captionText="FIGURES 117 – 122. Campsurus inusitatus sp. nov., male imago, SEM photographs: 117, genitalia, v. v. (g? = gonopore, mf = median furrow of styliger, ml = main lobe of penes, sl = secondary lobe); left mandible, v. v.; 118, detail of microlepides on sternum IX; 119, detail of penes and pedestals (note that penean membrane was broken during preparation); 120, detail of penean lobes, v. v. Line drawings: 121, male genitalia, v. v.; 122, male terminalia, l. v. (t. IX – t. X = tergum IX and X, s. IX = sternum IX, c = cercus base, p = paraproct, ml = main lobe of penes, sl = secondary lobe)." httpUri="https://zenodo.org/record/838466/files/figure.png" pageId="50" pageNumber="351">Figs. 117</figureCitation>
, 
<figureCitation id="131409A9EF45A2048849FC09FC09FC57" box="[859,897,973,994]" captionStart="FIGURES 123 – 128" captionStartId="34.[156,269,1818,1841]" captionTargetBox="[192,1394,445,1797]" captionTargetId="figure@34.[192,1394,445,1797]" captionTargetPageId="34" captionText="FIGURES 123 – 128. Campsurus indivisus, adult, SEM: 123, male terminalia, v. v.; 124, detail of pedestal, v. v. (p = pedestal, ic = inner corner); 125, detail of penes and pedestals, v. v. (bp = base of penes, g = gonopore, ml = main lobe of penes, sl = secondary lobe); 126, detail of apical portion of penes, v. v; 127, gonopores (g) at base of penes, v. v.; 128, microlepides on male sternum IX." httpUri="https://zenodo.org/record/838468/files/figure.png" pageId="50" pageNumber="351">123</figureCitation>
), penes with long and relatively thin lobes (
<figureCitation id="131409A9EF45A2048E2BFC09FA10FC57" box="[1337,1432,973,994]" captionStart="FIGURES 117 – 122" captionStartId="32.[151,264,1660,1683]" captionTargetBox="[196,1392,193,1620]" captionTargetId="figure@32.[196,1392,193,1639]" captionTargetPageId="32" captionText="FIGURES 117 – 122. Campsurus inusitatus sp. nov., male imago, SEM photographs: 117, genitalia, v. v. (g? = gonopore, mf = median furrow of styliger, ml = main lobe of penes, sl = secondary lobe); left mandible, v. v.; 118, detail of microlepides on sternum IX; 119, detail of penes and pedestals (note that penean membrane was broken during preparation); 120, detail of penean lobes, v. v. Line drawings: 121, male genitalia, v. v.; 122, male terminalia, l. v. (t. IX – t. X = tergum IX and X, s. IX = sternum IX, c = cercus base, p = paraproct, ml = main lobe of penes, sl = secondary lobe)." httpUri="https://zenodo.org/record/838466/files/figure.png" pageId="50" pageNumber="351">Figs. 120</figureCitation>
, 
<figureCitation id="131409A9EF45A2048BF3FC2EFE80FC4A" box="[225,264,1002,1023]" captionStart="FIGURES 123 – 128" captionStartId="34.[156,269,1818,1841]" captionTargetBox="[192,1394,445,1797]" captionTargetId="figure@34.[192,1394,445,1797]" captionTargetPageId="34" captionText="FIGURES 123 – 128. Campsurus indivisus, adult, SEM: 123, male terminalia, v. v.; 124, detail of pedestal, v. v. (p = pedestal, ic = inner corner); 125, detail of penes and pedestals, v. v. (bp = base of penes, g = gonopore, ml = main lobe of penes, sl = secondary lobe); 126, detail of apical portion of penes, v. v; 127, gonopores (g) at base of penes, v. v.; 128, microlepides on male sternum IX." httpUri="https://zenodo.org/record/838468/files/figure.png" pageId="50" pageNumber="351">126</figureCitation>
)................................................................................................15
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FBC3FA14FB8F" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FBC3FA14FB8F" pageId="50" pageNumber="351">
- Sternum IX with straight hind margin (
<figureCitation id="131409A9EF45A2048941FBC3FC85FBA9" box="[595,781,1031,1052]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 100, 103, 107</figureCitation>
) or with thin median notch (as in 
<taxonomicName id="4C2F6EAFEF45A2048F5FFBC3FB6EFBA9" ID-CoL="QF4H" authorityName="Spieth" authorityYear="1943" box="[1101,1254,1031,1052]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="pedicellarius">
<emphasis id="B95BC93EEF45A2048F5FFBC3FB6EFBA9" box="[1101,1254,1031,1052]" italics="true" pageId="50" pageNumber="351">C. pedicellarius</emphasis>
</taxonomicName>
); penes with short, subtriangullar main lobe and small, rounded secondary lobe (
<figureCitation id="131409A9EF45A2048839FBE1FC61FB8F" box="[811,1001,1061,1082]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 101, 104, 109</figureCitation>
)................................. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FB86FA14FBC0" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FB86FA13FBE2" blockId="50.[151,1436,151,1991]" box="[151,1435,1090,1111]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FB86FA13FBE2" box="[151,1435,1090,1111]" pageId="50" pageNumber="351">
15(14) Parastylus well-developed (
<figureCitation id="131409A9EF45A2048AF8FB86FDB6FBE2" box="[490,574,1090,1111]" captionStart="FIGURES 123 – 128" captionStartId="34.[156,269,1818,1841]" captionTargetBox="[192,1394,445,1797]" captionTargetId="figure@34.[192,1394,445,1797]" captionTargetPageId="34" captionText="FIGURES 123 – 128. Campsurus indivisus, adult, SEM: 123, male terminalia, v. v.; 124, detail of pedestal, v. v. (p = pedestal, ic = inner corner); 125, detail of penes and pedestals, v. v. (bp = base of penes, g = gonopore, ml = main lobe of penes, sl = secondary lobe); 126, detail of apical portion of penes, v. v; 127, gonopores (g) at base of penes, v. v.; 128, microlepides on male sternum IX." httpUri="https://zenodo.org/record/838468/files/figure.png" pageId="50" pageNumber="351">Fig. 124</figureCitation>
)....................................................... 
<taxonomicName id="4C2F6EAFEF45A2048FF5FB86FA13FBE2" ID-CoL="QF43" authority="Ulmer" authorityName="Ulmer ." authorityYear="1942" box="[1255,1435,1090,1111]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="indivisus">
<emphasis id="B95BC93EEF45A2048FF5FB86FAD0FBE2" box="[1255,1368,1090,1111]" italics="true" pageId="50" pageNumber="351">C. indivisus</emphasis>
Ulmer
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FB9BFA14FBC0" blockId="50.[151,1436,151,1991]" box="[151,1436,1119,1141]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FB9BFA14FBC0" box="[151,1436,1119,1141]" pageId="50" pageNumber="351">
- Parastylus absent (
<figureCitation id="131409A9EF45A2048A86FB9BFE6CFBC1" box="[404,484,1119,1140]" captionStart="FIGURES 117 – 122" captionStartId="32.[151,264,1660,1683]" captionTargetBox="[196,1392,193,1620]" captionTargetId="figure@32.[196,1392,193,1639]" captionTargetPageId="32" captionText="FIGURES 117 – 122. Campsurus inusitatus sp. nov., male imago, SEM photographs: 117, genitalia, v. v. (g? = gonopore, mf = median furrow of styliger, ml = main lobe of penes, sl = secondary lobe); left mandible, v. v.; 118, detail of microlepides on sternum IX; 119, detail of penes and pedestals (note that penean membrane was broken during preparation); 120, detail of penean lobes, v. v. Line drawings: 121, male genitalia, v. v.; 122, male terminalia, l. v. (t. IX – t. X = tergum IX and X, s. IX = sternum IX, c = cercus base, p = paraproct, ml = main lobe of penes, sl = secondary lobe)." httpUri="https://zenodo.org/record/838466/files/figure.png" pageId="50" pageNumber="351">Fig. 119</figureCitation>
)............................................................. 
<taxonomicName id="4C2F6EAFEF45A2048FDBFB9BFAC0FBC1" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1225,1352,1119,1140]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="inusitatus" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FDBFB9BFAC0FBC1" box="[1225,1352,1119,1140]" italics="true" pageId="50" pageNumber="351">C. inusitatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFB9BFA14FBC0" box="[1359,1436,1119,1141]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFB9BFA14FBC0" bold="true" box="[1359,1436,1119,1141]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FBB9FA13FB1A" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FBB9FA1DFB27" blockId="50.[151,1436,151,1991]" box="[151,1429,1149,1170]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FBB9FA1DFB27" box="[151,1429,1149,1170]" pageId="50" pageNumber="351">
16(14) Sternum IX medially notched (see 
<bibRefCitation id="EFBE68DDEF45A204893DFBB9FD2DFB27" author="Spieth" box="[559,677,1149,1170]" pageId="50" pageNumber="351" refString="Spieth, H. T. (1943) Taxonomic Studies on the Ephemeroptera III. Some interesting Ephemerids from Surinam and other Neotropical localities. American Museum Novitates, 1244, 1 - 13." type="journal article" year="1943">Spieth 1943</bibRefCitation>
: 
<figureCitation id="131409A9EF45A20489BDFBB9FD7DFB27" box="[687,757,1149,1170]" captionStart="FIGURES 13 – 24" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[235,1359,202,1827]" captionTargetId="figure@5.[211,1367,193,1852]" captionTargetPageId="5" captionText="FIGURES 13 – 24. Campsurus latipennis, nymph. 13, head, d. v. (long setae omitted); 14, left mandible, d. v. (row of filtering setae indicated with dashed line); 15, right mandible, v. v. (filtering setae as in previous figure); 16, apex of right mandibular tusk; 17, foreleg, d. v. (filtering setae not shown); 18, foreleg, v. v. (only filtering setae are indicated, with dashed lines); 19, foretarsal claw; 20, middle leg, d. v.; 21, middle tarsal claw; 22, hind leg, d. v.; 23, hind tarsal claw; 24, paraprocts with spines indicated (arrow)." httpUri="https://zenodo.org/record/838442/files/figure.png" pageId="50" pageNumber="351">Fig. 15</figureCitation>
).................................... 
<taxonomicName id="4C2F6EAFEF45A2048FABFBB9FA1DFB27" ID-CoL="QF4H" authority="Spieth" authorityName="Spieth" authorityYear="1943" box="[1209,1429,1149,1170]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="pedicellarius">
<emphasis id="B95BC93EEF45A2048FABFBB9FADAFB27" box="[1209,1362,1149,1170]" italics="true" pageId="50" pageNumber="351">C. pedicellarius</emphasis>
Spieth
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FB5EFA13FB1A" blockId="50.[151,1436,151,1991]" box="[151,1435,1178,1199]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FB5EFA13FB1A" box="[151,1435,1178,1199]" pageId="50" pageNumber="351">
- Sternum IX without such notch (
<figureCitation id="131409A9EF45A2048908FB5EFD51FB1A" box="[538,729,1178,1199]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 100, 103, 107</figureCitation>
)......................................................... 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FB73FA13FB5F" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FB73FA14FB78" blockId="50.[151,1436,151,1991]" box="[151,1436,1207,1229]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FB73FA14FB78" box="[151,1436,1207,1229]" pageId="50" pageNumber="351">
17(16) Penes dorsally with microspines on outer margin (
<figureCitation id="131409A9EF45A20489D0FB73FC96FB79" box="[706,798,1207,1228]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 108</figureCitation>
, 
<figureCitation id="131409A9EF45A204883AFB73FCC3FB79" box="[808,843,1207,1228]" captionStart="FIGURES 111 – 116" captionStartId="29.[151,264,1822,1845]" captionTargetBox="[212,1374,445,1802]" captionTargetId="figure@29.[212,1374,445,1802]" captionTargetPageId="29" captionText="FIGURES 111 – 116. Campsurus demeni sp. nov.: 111, penes and pedestal, with detail of microlepides on sternum IX, v. v. (SEM). Campsurus lucidus (Bolivia): 112, male mesosternum (p = mesofurcasternal plate, im = inner margin of plate, L = leg); 113, female abdominal sterna VI – IX (g = gonopore and eggs, sk = sockets, roman numbers = number of abdominal segment); 114, same, detail after KOH treatment. Campsurus sinamari sp. nov. (KOH treated): 115, sternum VIII; 116, detail of sockets. Arrows indicate socket´s openings." httpUri="https://zenodo.org/record/838464/files/figure.png" pageId="50" pageNumber="351">111</figureCitation>
)................................ 
<taxonomicName id="4C2F6EAFEF45A2048FFAFB73FAC1FB79" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1256,1353,1207,1228]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="demeni" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FFAFB73FAC1FB79" box="[1256,1353,1207,1228]" italics="true" pageId="50" pageNumber="351">C. demeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFB73FA14FB78" box="[1359,1436,1207,1229]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFB73FA14FB78" bold="true" box="[1359,1436,1207,1229]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FB11FA13FB5F" blockId="50.[151,1436,151,1991]" box="[151,1435,1237,1258]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FB11FA13FB5F" box="[151,1435,1237,1258]" pageId="50" pageNumber="351">
- Penes smooth, without microspines as above (
<figureCitation id="131409A9EF45A2048989FB11FCA2FB5F" box="[667,810,1237,1258]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 101, 104</figureCitation>
)..................................................18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FB36FA14FA90" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FB36FA13FAB2" blockId="50.[151,1436,151,1991]" box="[151,1435,1266,1287]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FB36FA13FAB2" box="[151,1435,1266,1287]" pageId="50" pageNumber="351">
18(17) Parastylus long and slender, at least 2× length of pedestal (
<figureCitation id="131409A9EF45A2048801FB36FCECFAB2" box="[787,868,1266,1287]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Fig. 100</figureCitation>
)..................... 
<taxonomicName id="4C2F6EAFEF45A2048F7FFB36FA13FAB2" ID-CoL="QF48" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy ." authorityYear="1924" box="[1133,1435,1266,1287]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="lucidus">
<emphasis id="B95BC93EEF45A2048F7FFB36FB46FAB2" box="[1133,1230,1266,1287]" italics="true" pageId="50" pageNumber="351">C. lucidus</emphasis>
Needham &amp; Murphy
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FACBFA14FA90" blockId="50.[151,1436,151,1991]" box="[151,1436,1295,1317]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FACBFA14FA90" box="[151,1436,1295,1317]" pageId="50" pageNumber="351">
- Parastylus short, less than 1.5× the length of pedestal (
<figureCitation id="131409A9EF45A20489F8FACBFCF0FA91" box="[746,888,1295,1316]" captionStart="FIGURES 100 – 110" captionStartId="27.[151,264,1800,1823]" captionTargetBox="[208,1388,291,1770]" captionTargetId="figure@27.[193,1394,265,1779]" captionTargetPageId="27" captionText="FIGURES 100 – 110. Male imagos. Campsurus lucidus: 100, genitalia, v. v.; 101, detail of penes, v. v.; 102, detail of pedestal, v. v. Campsurus sinamari sp. nov.: 103, genitalia, v. v.; 104, detail of penes, v. v.; 105 – 106, details of pedestal, v. v. Campsurus demeni sp. nov.: 107, genitalia, v. v.; 108, male terminalia, l. v.; 109, detail of penes, v. v.; 110, detail of pedestal, v. v. Abbreviations: ic = inner corner, ml = main lobe, p = parastylus, sl = secondary lobe." httpUri="https://zenodo.org/record/838460/files/figure.png" pageId="50" pageNumber="351">Figs. 105–106</figureCitation>
)............................. 
<taxonomicName id="4C2F6EAFEF45A2048FCAFACBFAC1FA91" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1240,1353,1295,1316]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="sinamari" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FCAFACBFAC1FA91" box="[1240,1353,1295,1316]" italics="true" pageId="50" pageNumber="351">C. sinamari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687445EF45A2048E5DFACBFA14FA90" box="[1359,1436,1295,1317]" pageId="50" pageNumber="351" rank="species">
<emphasis id="B95BC93EEF45A2048E5DFACBFA14FA90" bold="true" box="[1359,1436,1295,1317]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FAE9FA13FAC9" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FAE9FA13FAEA" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FAE9FA13FAEA" pageId="50" pageNumber="351">
19(13) Distal margin of sternum IX with a subtriangular distomedial projection, sometimes also with a small median triangular point (
<figureCitation id="131409A9EF45A2048BFAFA8EFECCFAEA" box="[232,324,1354,1375]" captionStart="FIGURES 1 – 12" captionStartId="4.[151,264,1834,1857]" captionTargetBox="[229,1357,377,1813]" captionTargetId="figure@4.[229,1357,373,1813]" captionTargetPageId="4" captionText="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." httpUri="https://zenodo.org/record/838440/files/figure.png" pageId="50" pageNumber="351">Figs. 1, 9</figureCitation>
, 
<figureCitation id="131409A9EF45A2048A5CFA8EFE03FAEA" box="[334,395,1354,1375]" captionStart="FIGURES 25 – 34" captionStartId="7.[151,264,1939,1962]" captionTargetBox="[212,1363,202,1896]" captionTargetId="figure@7.[212,1363,193,1896]" captionTargetPageId="7" captionText="FIGURES 25 – 34. Campsurus salobra sp. nov., adult: 25, male genitalia, v. v.; 26, same, l. v.; 27, female sternum VIII; 28, detail of female sockets. SEM photographs: 29, male genitalia, v. v. (with detail of microtrichiae); 30, apical portion of penes, v. v.; 31, female sterna VIII – X; 32, female sockets (with detail of microtrichia); 33, egg, convex side; 34, detail of capped pole." httpUri="https://zenodo.org/record/838444/files/figure.png" pageId="50" pageNumber="351">25, 29</figureCitation>
)....................................................................................20
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85FAA3FA13FAC9" blockId="50.[151,1436,151,1991]" box="[151,1435,1383,1404]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FAA3FA13FAC9" box="[151,1435,1383,1404]" pageId="50" pageNumber="351">
- Distal margin of sternum IX straight, slightly concave, or with a small truncated median projection (
<figureCitation id="131409A9EF45A2048FB3FAA3FAA4FAC9" box="[1185,1324,1383,1404]" captionStart="FIGURES 138 – 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 – 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="50" pageNumber="351">Figs. 138, 140</figureCitation>
)...... 2 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85FA41FA13FA61" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FA41FA13FA02" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FA41FA13FA02" pageId="50" pageNumber="351">
20(19) Main lobe of penes with a similar width along its length (Emmerich &amp; Molineri 2011: 
<figureCitation id="131409A9EF45A2048F0DFA41FB1AFA2F" box="[1055,1170,1413,1434]" captionStart-0="FIGURES 1 – 12" captionStart-1="FIGURES 13 – 24" captionStartId-0="4.[151,264,1834,1857]" captionStartId-1="5.[151,264,1882,1905]" captionTargetBox-0="[229,1357,377,1813]" captionTargetBox-1="[235,1359,202,1827]" captionTargetId-0="figure@4.[229,1357,373,1813]" captionTargetId-1="figure@5.[211,1367,193,1852]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." captionText-1="FIGURES 13 – 24. Campsurus latipennis, nymph. 13, head, d. v. (long setae omitted); 14, left mandible, d. v. (row of filtering setae indicated with dashed line); 15, right mandible, v. v. (filtering setae as in previous figure); 16, apex of right mandibular tusk; 17, foreleg, d. v. (filtering setae not shown); 18, foreleg, v. v. (only filtering setae are indicated, with dashed lines); 19, foretarsal claw; 20, middle leg, d. v.; 21, middle tarsal claw; 22, hind leg, d. v.; 23, hind tarsal claw; 24, paraprocts with spines indicated (arrow)." httpUri-0="https://zenodo.org/record/838440/files/figure.png" httpUri-1="https://zenodo.org/record/838442/files/figure.png" pageId="50" pageNumber="351">Figs. 11–13</figureCitation>
); secondary lobe variable................................................................................................... .. 21
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</paragraph>
<paragraph id="8B90152CEF45A2048B85FA7BFA13FA61" blockId="50.[151,1436,151,1991]" box="[151,1435,1471,1492]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FA7BFA13FA61" box="[151,1435,1471,1492]" pageId="50" pageNumber="351">
- Main lobe of penes subtriangular, wider at base; secondary lobe reduced in size (
<figureCitation id="131409A9EF45A20488F5FA7BFBA2FA61" box="[999,1066,1471,1492]" captionStart="FIGURES 1 – 12" captionStartId="4.[151,264,1834,1857]" captionTargetBox="[229,1357,377,1813]" captionTargetId="figure@4.[229,1357,373,1813]" captionTargetPageId="4" captionText="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." httpUri="https://zenodo.org/record/838440/files/figure.png" pageId="50" pageNumber="351">Figs. 3</figureCitation>
, 
<figureCitation id="131409A9EF45A2048F27FA7BFBD8FA61" box="[1077,1104,1471,1492]" captionStart="FIGURES 25 – 34" captionStartId="7.[151,264,1939,1962]" captionTargetBox="[212,1363,202,1896]" captionTargetId="figure@7.[212,1363,193,1896]" captionTargetPageId="7" captionText="FIGURES 25 – 34. Campsurus salobra sp. nov., adult: 25, male genitalia, v. v.; 26, same, l. v.; 27, female sternum VIII; 28, detail of female sockets. SEM photographs: 29, male genitalia, v. v. (with detail of microtrichiae); 30, apical portion of penes, v. v.; 31, female sterna VIII – X; 32, female sockets (with detail of microtrichia); 33, egg, convex side; 34, detail of capped pole." httpUri="https://zenodo.org/record/838444/files/figure.png" pageId="50" pageNumber="351">30</figureCitation>
).........................22
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</paragraph>
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<keyStep id="30DB0D89EF45A2048B85FA19FA14F999" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85FA19FA13F9BA" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85FA19FA13F9BA" pageId="50" pageNumber="351">
21(20) Penes with secondary lobe cylindrical and relatively long (Emmerich &amp; Molineri 2011: 
<figureCitation id="131409A9EF45A2048F28FA19FB26FA47" box="[1082,1198,1501,1522]" captionStart-0="FIGURES 1 – 12" captionStart-1="FIGURES 13 – 24" captionStartId-0="4.[151,264,1834,1857]" captionStartId-1="5.[151,264,1882,1905]" captionTargetBox-0="[229,1357,377,1813]" captionTargetBox-1="[235,1359,202,1827]" captionTargetId-0="figure@4.[229,1357,373,1813]" captionTargetId-1="figure@5.[211,1367,193,1852]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." captionText-1="FIGURES 13 – 24. Campsurus latipennis, nymph. 13, head, d. v. (long setae omitted); 14, left mandible, d. v. (row of filtering setae indicated with dashed line); 15, right mandible, v. v. (filtering setae as in previous figure); 16, apex of right mandibular tusk; 17, foreleg, d. v. (filtering setae not shown); 18, foreleg, v. v. (only filtering setae are indicated, with dashed lines); 19, foretarsal claw; 20, middle leg, d. v.; 21, middle tarsal claw; 22, hind leg, d. v.; 23, hind tarsal claw; 24, paraprocts with spines indicated (arrow)." httpUri-0="https://zenodo.org/record/838440/files/figure.png" httpUri-1="https://zenodo.org/record/838442/files/figure.png" pageId="50" pageNumber="351">Figs. 11–13</figureCitation>
); male fore wings 
<quantity id="4CD7B8C9EF45A2048E74FA19FEB2F9BA" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" metricValueMax="2.05" metricValueMin="1.59" pageId="50" pageNumber="351" unit="mm" value="18.2" valueMax="20.5" valueMin="15.9">15.9– 20.5 mm</quantity>
................................................................ 
<taxonomicName id="4C2F6EAFEF45A2048F2AFA3EFA13F9BA" ID-CoL="QF4R" authority="Emmerich &amp; Molineri" authorityName="Emmerich &amp; Molineri ." authorityYear="2011" box="[1080,1435,1530,1551]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="vulturorum">
<emphasis id="B95BC93EEF45A2048F2AFA3EFB37F9BA" box="[1080,1215,1530,1551]" italics="true" pageId="50" pageNumber="351">C. vulturorum</emphasis>
Emmerich &amp; Molineri
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</paragraph>
<paragraph id="8B90152CEF45A2048B85F9D3FA14F999" blockId="50.[151,1436,151,1991]" box="[151,1436,1559,1580]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F9D3FA14F999" box="[151,1436,1559,1580]" pageId="50" pageNumber="351">
- Secondary lobe of penes apparently absent (
<bibRefCitation id="EFBE68DDEF45A2048994F9D3FC1EF999" author="Eaton" box="[646,918,1559,1580]" pageId="50" pageNumber="351" refString="Eaton, A. E. (1871) A Monograph of the Ephemeridae. Transactions of the Entomological Society of London, 1871, 1 - 164." type="journal article" year="1871">
Eaton 1871: Plate 4, 
<figureCitation id="131409A9EF45A2048842F9D3FC1EF999" box="[848,918,1559,1580]" captionStart="FIGURES 1 – 12" captionStartId="4.[151,264,1834,1857]" captionTargetBox="[229,1357,377,1813]" captionTargetId="figure@4.[229,1357,373,1813]" captionTargetPageId="4" captionText="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." httpUri="https://zenodo.org/record/838440/files/figure.png" pageId="50" pageNumber="351">Fig. 12</figureCitation>
</bibRefCitation>
), size unknown............. 
<taxonomicName id="4C2F6EAFEF45A2048FC9F9D3FA14F999" ID-CoL="QF3R" authority="Eaton" authorityName="Eaton" authorityYear="1871" box="[1243,1436,1559,1580]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B95BC93EEF45A2048FC9F9D3FAD5F999" box="[1243,1373,1559,1580]" italics="true" pageId="50" pageNumber="351">C. cuspidatus</emphasis>
Eaton
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85F9F1FA14F9D2" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85F9F1FA14F9FF" blockId="50.[151,1436,151,1991]" box="[151,1436,1588,1610]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F9F1FA14F9FF" box="[151,1436,1588,1610]" pageId="50" pageNumber="351">
22(20) Penes with dorsal margin smoothly curved in lateral view (
<figureCitation id="131409A9EF45A2048804F9F1FCD6F9FF" box="[790,862,1589,1610]" captionStart="FIGURES 25 – 34" captionStartId="7.[151,264,1939,1962]" captionTargetBox="[212,1363,202,1896]" captionTargetId="figure@7.[212,1363,193,1896]" captionTargetPageId="7" captionText="FIGURES 25 – 34. Campsurus salobra sp. nov., adult: 25, male genitalia, v. v.; 26, same, l. v.; 27, female sternum VIII; 28, detail of female sockets. SEM photographs: 29, male genitalia, v. v. (with detail of microtrichiae); 30, apical portion of penes, v. v.; 31, female sterna VIII – X; 32, female sockets (with detail of microtrichia); 33, egg, convex side; 34, detail of capped pole." httpUri="https://zenodo.org/record/838444/files/figure.png" pageId="50" pageNumber="351">Fig. 26</figureCitation>
)................................ 
<taxonomicName id="4C2F6EAFEF45A2048FF1F9F1FAC1F9FF" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1251,1353,1589,1610]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="salobra" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FF1F9F1FAC1F9FF" box="[1251,1353,1589,1610]" italics="true" pageId="50" pageNumber="351">C. salobra</emphasis>
</taxonomicName>
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<emphasis id="B95BC93EEF45A2048E5DF9F0FA14F9FF" bold="true" box="[1359,1436,1588,1610]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B90152CEF45A2048B85F996FA14F9D2" blockId="50.[151,1436,151,1991]" box="[151,1436,1618,1639]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F996FA14F9D2" box="[151,1436,1618,1639]" pageId="50" pageNumber="351">
- Penes with indented or sinuous dorsal margin in lateral view (
<figureCitation id="131409A9EF45A2048823F996FC07F9D2" box="[817,911,1618,1639]" captionStart="FIGURES 1 – 12" captionStartId="4.[151,264,1834,1857]" captionTargetBox="[229,1357,377,1813]" captionTargetId="figure@4.[229,1357,373,1813]" captionTargetPageId="4" captionText="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." httpUri="https://zenodo.org/record/838440/files/figure.png" pageId="50" pageNumber="351">Figs. 4–5</figureCitation>
)......................................... 23
</keyLead>
</paragraph>
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<keyStep id="30DB0D89EF45A2048B85F9ABFA13F90A" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85F9ABFA13F931" blockId="50.[151,1436,151,1991]" box="[151,1435,1647,1668]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F9ABFA13F931" box="[151,1435,1647,1668]" pageId="50" pageNumber="351">
23(22) Sternum IX projected medially but without an acute point (
<figureCitation id="131409A9EF45A2048805F9ABFCF9F931" box="[791,881,1647,1668]" captionStart="FIGURES 1 – 12" captionStartId="4.[151,264,1834,1857]" captionTargetBox="[229,1357,377,1813]" captionTargetId="figure@4.[229,1357,373,1813]" captionTargetPageId="4" captionText="FIGURES 1 – 12. Campsurus latipennis, adult. Male imago: 1, genitalia, ventral view (v. v.); 2, details of pedestal, v. v.; 3, detail of penes and pedestals, v. v.; 4 – 5, penes, lateral view (l. v.); 6, genitalia, l. v. SEM photographs: 9, genitalia, v. v.; 10, detail of microtrichia on sternum IX; 11, pedestal, v. v.; 12, detail of penes, v. v. Female adult: 7, abdominal sternum VIII; 8, detail of sockets. Abbreviations: bp = base of penes, ml = main lobe of penes, mp = median projection of sternum IX, ps = parastylus, sl = secondary lobe of penes." httpUri="https://zenodo.org/record/838440/files/figure.png" pageId="50" pageNumber="351">Figs. 1, 9</figureCitation>
)........................... 
<taxonomicName id="4C2F6EAFEF45A2048FD4F9ABFA13F931" ID-CoL="QF45" authority="Walker" authorityName="Walker" authorityYear="1853" box="[1222,1435,1647,1668]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="B95BC93EEF45A2048FD4F9ABFAC8F931" box="[1222,1344,1647,1668]" italics="true" pageId="50" pageNumber="351">C. latipennis</emphasis>
(Walker)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B90152CEF45A2048B85F949FA13F90A" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F949FA13F90A" pageId="50" pageNumber="351">
- Sternum IX projected medially and with an acute triangular point (Emmerich &amp; Molineri 2011: 
<figureCitation id="131409A9EF45A2048F6AF949FB78F917" box="[1144,1264,1677,1698]" captionStart="FIGURES 25 – 34" captionStartId="7.[151,264,1939,1962]" captionTargetBox="[212,1363,202,1896]" captionTargetId="figure@7.[212,1363,193,1896]" captionTargetPageId="7" captionText="FIGURES 25 – 34. Campsurus salobra sp. nov., adult: 25, male genitalia, v. v.; 26, same, l. v.; 27, female sternum VIII; 28, detail of female sockets. SEM photographs: 29, male genitalia, v. v. (with detail of microtrichiae); 30, apical portion of penes, v. v.; 31, female sterna VIII – X; 32, female sockets (with detail of microtrichia); 33, egg, convex side; 34, detail of capped pole." httpUri="https://zenodo.org/record/838444/files/figure.png" pageId="50" pageNumber="351">Figs. 33, 34</figureCitation>
)........................................................................................ 
<taxonomicName id="4C2F6EAFEF45A2048F4FF96EFA13F90A" ID-CoL="QF3Z" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[1117,1435,1706,1727]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="evanidus">
<emphasis id="B95BC93EEF45A2048F4FF96EFB45F90A" box="[1117,1229,1706,1727]" italics="true" pageId="50" pageNumber="351">C. evanidus</emphasis>
Needham &amp; Murphy
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85F903FA13F881" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85F903FA14F94F" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F903FA14F94F" pageId="50" pageNumber="351">
24(19) Main lobe of penes slightly curved, conical; secondary lobe short, less than half the length of main lobe (Emmerich &amp; Molineri 2011: 
<figureCitation id="131409A9EF45A2048A0EF921FEE9F94F" box="[284,353,1765,1786]" captionStart="FIGURES 35 – 50" captionStartId="10.[151,264,1859,1882]" captionTargetBox="[218,1378,208,1822]" captionTargetId="figure@10.[212,1381,207,1822]" captionTargetPageId="10" captionText="FIGURES 35 – 50. Campsurus essequibo, imago. Holotype male genitalia: 35, general v. v.; 36, detail of penes, v. v. Female adult: 37, abdominal sternum VIII; 38, detail of sockets. Male imago: 39 – 42, genitalia, variations, v. v.; 43, genitalia after treatment with KOH; 44 – 47, genitalia, variations, l. v.; 48, male mesofurcasternum. SEM photographs: 49, genitalia, v. v. (with detail of microlepides); 50, detail of penes, v. v." httpUri="https://zenodo.org/record/838446/files/figure.png" pageId="50" pageNumber="351">Fig. 50</figureCitation>
)............................................................ 
<taxonomicName id="4C2F6EAFEF45A2048F5EF921FA14F94F" ID-CoL="QF44" authority="Esben-Petersen 1912" authorityName="Esben-Petersen" authorityYear="1912" box="[1100,1436,1765,1786]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="jorgenseni">
<emphasis id="B95BC93EEF45A2048F5EF921FB43F94F" box="[1100,1227,1765,1786]" italics="true" pageId="50" pageNumber="351">C. jorgenseni</emphasis>
<bibRefCitation id="EFBE68DDEF45A2048FC0F921FA14F94F" author="Esben-Petersen" box="[1234,1436,1765,1786]" pageId="50" pageNumber="351" refString="Esben-Petersen, T. (1912) New and little-known species of Ephemerida from Argentine (Neuropt.). Deutsche Entomologische Zeitschrift, 1912, 333 - 342." type="journal article" year="1912">Esben-Petersen 1912</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8B90152CEF45A2048B85F8C6FA13F881" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F8C6FA13F881" pageId="50" pageNumber="351">
- Main lobe of penes strongly curved (
<figureCitation id="131409A9EF45A204895AF8C6FC8FF8A2" box="[584,775,1794,1815]" captionStart="FIGURES 138 – 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 – 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="50" pageNumber="351">Figs. 139, 142, 143</figureCitation>
) and cylindrical; secondary lobe long (more than 0.6× the length of main lobe)..........................................................................................25
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</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85F8F9FA14F8DA" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85F8F9FA14F8E7" blockId="50.[151,1436,151,1991]" box="[151,1436,1852,1874]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F8F9FA14F8E7" box="[151,1436,1852,1874]" pageId="50" pageNumber="351">
25(24) Main and secondary lobes of penes arising from a large membranous base (
<figureCitation id="131409A9EF45A20488A1F8F9FBB6F8E7" box="[947,1086,1853,1874]" captionStart="FIGURES 138 – 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 – 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="50" pageNumber="351">Figs. 142, 143</figureCitation>
)............. 
<taxonomicName id="4C2F6EAFEF45A2048FFCF8F9FAC0F8E7" authority="Molineri &amp; Salles, 2017" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[1262,1352,1853,1874]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="cotaxe" status="sp. nov.">
<emphasis id="B95BC93EEF45A2048FFCF8F9FAC0F8E7" box="[1262,1352,1853,1874]" italics="true" pageId="50" pageNumber="351">C. cotaxe</emphasis>
</taxonomicName>
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<emphasis id="B95BC93EEF45A2048E5DF8F8FA14F8E7" bold="true" box="[1359,1436,1852,1874]" pageId="50" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B90152CEF45A2048B85F89EFA14F8DA" blockId="50.[151,1436,151,1991]" box="[151,1436,1882,1903]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F89EFA14F8DA" box="[151,1436,1882,1903]" pageId="50" pageNumber="351">
- Membranous base of penes smaller; secondary lobe arising basally from main lobe (
<figureCitation id="131409A9EF45A2048F1AF89EFBD3F8DA" box="[1032,1115,1882,1903]" captionStart="FIGURES 138 – 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 – 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="50" pageNumber="351">Fig. 139</figureCitation>
)....................... 26
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</paragraph>
</keyStep>
<keyStep id="30DB0D89EF45A2048B85F8B3FA13F872" pageId="50" pageNumber="351">
<paragraph id="8B90152CEF45A2048B85F8B3FA1DF81F" blockId="50.[151,1436,151,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F8B3FA1DF81F" pageId="50" pageNumber="351">
26(25) Smaller size (male FW max. length 
<quantity id="4CD7B8C9EF45A2048928F8B3FD0FF839" box="[570,647,1911,1932]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="50" pageNumber="351" unit="mm" value="9.8">9.8 mm</quantity>
), abdomen darker (brownish to reddish, 
<figureCitation id="131409A9EF45A2048F19F8B3FB1DF839" box="[1035,1173,1911,1932]" captionStart="FIGURES 146 – 150" captionStartId="40.[151,264,1639,1662]" captionTargetBox="[226,1361,195,1618]" captionTargetId="figure@40.[226,1361,195,1618]" captionTargetPageId="40" captionText="FIGURES 146 – 150. C. segnis (specimens from Venezuela): 146, male imago, abdomen, laterodorsal view; 147, female imago, abdomen, d. v.; 148, female head and thorax, v. v. (fs = mesofurcasternum, im = inner margin, p = mesofurcasternal plates, L = middle legs); 149, abdominal sterna (g = gonopore and eggs, sk = sockets, roman numbers = segment number); 150, detail of sockets (arrows indicate socket´s openings)." httpUri="https://zenodo.org/record/838474/files/figure.png" pageId="50" pageNumber="351">Figs. 146, 147</figureCitation>
)............................................................................................... 
<taxonomicName id="4C2F6EAFEF45A2048F62F851FA1DF81F" ID-CoL="QF4L" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy ." authorityYear="1924" box="[1136,1429,1941,1962]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="segnis">
<emphasis id="B95BC93EEF45A2048F62F851FB40F81F" box="[1136,1224,1941,1962]" italics="true" pageId="50" pageNumber="351">C. segnis</emphasis>
Needham &amp; Murphy
</taxonomicName>
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</paragraph>
<paragraph id="8B90152CEF45A2048B85F876FA13F872" blockId="50.[151,1436,151,1991]" box="[151,1435,1970,1991]" pageId="50" pageNumber="351">
<keyLead id="30DEB619EF45A2048B85F876FA13F872" box="[151,1435,1970,1991]" pageId="50" pageNumber="351">
- Larger size (male FW length 
<quantity id="4CD7B8C9EF45A2048AE8F876FDC8F872" box="[506,576,1970,1991]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="50" pageNumber="351" unit="mm" value="11.0">11 mm</quantity>
), paler abdomen (whitish to yellowish)........................... 
<taxonomicName id="4C2F6EAFEF45A2048FEBF876FA13F872" ID-CoL="QF4P" authority="Traver" authorityName="Traver" authorityYear="1947" box="[1273,1435,1970,1991]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="351" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B95BC93EEF45A2048FEBF876FADEF872" box="[1273,1366,1970,1991]" italics="true" pageId="50" pageNumber="351">C. ulmeri</emphasis>
Traver
</taxonomicName>
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>