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<document id="27B2B9956A4ACAED2D15307250028B80" ID-DOI="10.5281/zenodo.195947" ID-GBIF-Dataset="42a82f84-c379-48f0-812c-52c6c244d62a" ID-ISSN="1175-5326" ID-Zenodo-Dep="195947" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460236227298" checkinUser="plazi" docAuthor="Queney, Pierre" docDate="2010" docId="05042952FFFEFFBD96E6A6EDFEA343A3" docLanguage="en" docName="zt02506p058.pdf" docOrigin="Zootaxa 2506" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Berosus spiniger Queney, 2010, sp. n." docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="F93D512AFFFAFFBA9671A220FFB6427F" masterDocTitle="Two new species of Berosus Leach from Brazil (Coleoptera: Hydrophilidae: Berosini)" masterLastPageNumber="58" masterPageNumber="51" pageNumber="55" updateTime="1698239631079" updateUser="plazi">
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<mods:title id="53CA3191E1E234BC6E8ABE475BFEEC23">Two new species of Berosus Leach from Brazil (Coleoptera: Hydrophilidae: Berosini)</mods:title>
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<mods:namePart id="DD338AF77ED9F7FC3D5382E826AFED2D">Queney, Pierre</mods:namePart>
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<mods:date id="EC189345815A0B404C92EBAEFDEBC4F5">2010</mods:date>
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<treatment id="05042952FFFEFFBD96E6A6EDFEA343A3" ID-DOI="http://doi.org/10.5281/zenodo.6201419" ID-GBIF-Taxon="119388279" ID-Zenodo-Dep="6201419" LSID="urn:lsid:plazi:treatment:05042952FFFEFFBD96E6A6EDFEA343A3" httpUri="http://treatment.plazi.org/id/05042952FFFEFFBD96E6A6EDFEA343A3" lastPageId="7" lastPageNumber="58" pageId="4" pageNumber="55">
<subSubSection id="C5B7CBCFFFFEFFBE96E6A6EDFED64775" pageId="4" pageNumber="55" type="nomenclature">
<paragraph id="8D129844FFFEFFBE96E6A6EDFE1D4698" blockId="4.[151,427,1229,1290]" box="[151,427,1229,1255]" pageId="4" pageNumber="55">
<heading id="D65A2F28FFFEFFBE96E6A6EDFE1D4698" bold="true" box="[151,427,1229,1255]" fontSize="11" level="1" pageId="4" pageNumber="55" reason="1">
<emphasis id="BFD94456FFFEFFBE96E6A6EDFE1D4698" bold="true" box="[151,427,1229,1255]" pageId="4" pageNumber="55">
<taxonomicName id="4AADE3C7FFFEFFBE96E6A6EDFED44698" ID-CoL="LMDG" box="[151,354,1229,1255]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="BFD94456FFFEFFBE96E6A6EDFED44698" bold="true" box="[151,354,1229,1255]" italics="true" pageId="4" pageNumber="55">Berosus spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4EAF92DFFFEFFBE9719A6EDFE1D4698" box="[360,427,1229,1255]" pageId="4" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8D129844FFFEFFBE96E6A6D0FED64775" blockId="4.[151,427,1229,1290]" box="[151,352,1264,1290]" pageId="4" pageNumber="55">
(
<figureCitation id="159684C1FFFEFFBE96EEA6D0FF454775" box="[159,243,1264,1290]" captionStart="FIGURES 1 2" captionStartId="1.[151,269,1054,1078]" captionTargetBox="[318,1288,422,1015]" captionTargetId="figure@1.[290,1296,421,1029]" captionTargetPageId="1" captionText="FIGURES 1 2. Dorsal habitus: 1) Berosus degallieri (3.3 mm long); 2) Berosus spiniger (6.6 mm long)." httpUri="https://zenodo.org/record/195948/files/figure.png" pageId="4" pageNumber="55">Figs. 2</figureCitation>
,
<figureCitation id="159684C1FFFEFFBE968EA6D0FEB94775" box="[255,271,1264,1290]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="4" pageNumber="55">7</figureCitation>
,
<figureCitation id="159684C1FFFEFFBE976DA6D0FEE04775" box="[284,342,1264,1290]" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="4" pageNumber="55">811</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C5B7CBCFFFFEFFBE96E6A71AFD2A4705" pageId="4" pageNumber="55" type="materials_examined">
<paragraph id="8D129844FFFEFFBE96E6A71AFD2A4705" blockId="4.[151,1437,1338,2036]" pageId="4" pageNumber="55">
<emphasis id="BFD94456FFFEFFBE96E6A71AFEE2472B" bold="true" box="[151,340,1338,1364]" pageId="4" pageNumber="55">
<typeStatus id="521626E6FFFEFFBE96E6A71AFF63472B" box="[151,213,1338,1364]" pageId="4" pageNumber="55">Type</typeStatus>
material.
</emphasis>
<typeStatus id="521626E6FFFEFFBE972FA71AFE66472B" box="[350,464,1338,1364]" pageId="4" pageNumber="55" type="holotype">Holotype</typeStatus>
(male): &quot;
<collectingCountry id="F5BAD8D4FFFEFFBE9435A71AFD21472B" box="[580,663,1338,1364]" name="Brazil" pageId="4" pageNumber="55">Brazil</collectingCountry>
:
<collectingCountry id="F5BAD8D4FFFEFFBE94D5A71AFCB0472B" box="[676,774,1338,1364]" name="Brazil" pageId="4" pageNumber="55">Brasilia</collectingCountry>
,/ Distrito Federal,
<quantity id="4A5535A1FFFEFFBE9580A71AFBF9472B" box="[1009,1103,1338,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="4" pageNumber="55" unit="m" value="1138.0">1138 m</quantity>
,/
<date id="F913BE84FFFEFFBE9217A71AFB7D472B" box="[1126,1227,1338,1364]" pageId="4" pageNumber="55" value="1999-03">III.1999</date>
,/
<geoCoordinate id="E899FE83FFFEFFBE9295A71AFA21472B" box="[1252,1431,1338,1364]" direction="south" orientation="latitude" pageId="4" pageNumber="55" precision="1" value="-15.759997">15°45'35.99''S</geoCoordinate>
,
<geoCoordinate id="E899FE83FFFEFFBE96E6A740FEFE4705" box="[151,328,1376,1402]" direction="west" orientation="longitude" pageId="4" pageNumber="55" precision="1" value="-47.919777">47°55'11.19''W</geoCoordinate>
,/ N. Dégallier leg.&quot; (
<collectionCode id="EBBC0081FFFEFFBE944DA740FD394705" box="[572,655,1376,1402]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="4" pageNumber="55">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C5B7CBCFFFFEFFBE96B7A7A5FA3C444B" pageId="4" pageNumber="55" type="diagnosis">
<paragraph id="8D129844FFFEFFBE96B7A7A5FA3C444B" blockId="4.[151,1437,1338,2036]" pageId="4" pageNumber="55">
<emphasis id="BFD94456FFFEFFBE96B7A7A5FEF047E0" bold="true" box="[198,326,1413,1439]" pageId="4" pageNumber="55">Diagnosis.</emphasis>
Within the
<taxonomicName id="4AADE3C7FFFEFFBE97A1A7A5FD8347E1" box="[464,565,1413,1438]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFEFFBE97A1A7A5FD8347E1" box="[464,565,1413,1438]" italics="true" pageId="4" pageNumber="55">auriceps</emphasis>
</taxonomicName>
-complex,
<taxonomicName id="4AADE3C7FFFEFFBE94C3A7A5FCC047E1" box="[690,886,1413,1438]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="BFD94456FFFEFFBE94C3A7A5FCC047E1" box="[690,886,1413,1438]" italics="true" pageId="4" pageNumber="55">Berosus spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4EAF92DFFFEFFBE950FA7A5FC0B47E0" box="[894,957,1413,1439]" pageId="4" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
combines a mesosternal process entirely laminar (
<figureCitation id="159684C1FFFEFFBE9772A78AFEFA47BB" box="[259,332,1450,1476]" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="4" pageNumber="55">Fig. 8</figureCitation>
), black spots without metallic sheen on pronotum disc on both sides of a median testaceous band, micropunctures very rare on interstriae, femoral pubescence obliquely limited and rather extensive (
<figureCitation id="159684C1FFFEFFBE931DA7F0FF0F4470" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="4" pageNumber="55">Fig. 10</figureCitation>
), male genitalia with distal part beyond basal piece narrow in ventral and dorsal aspect, basal piece representing about 60% of their total length, parameres ending in two narrow and parallel apices (
<figureCitation id="159684C1FFFEFFBE9360A43AFAE1444B" box="[1297,1367,1562,1588]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="4" pageNumber="55">Fig. 7</figureCitation>
b,c).
</paragraph>
</subSubSection>
<subSubSection id="C5B7CBCFFFFEFFBC96B7A41FFE65455D" lastPageId="6" lastPageNumber="57" pageId="4" pageNumber="55" type="description">
<paragraph id="8D129844FFFEFFBE96B7A41FFC3144DB" blockId="4.[151,1437,1338,2036]" pageId="4" pageNumber="55">
<emphasis id="BFD94456FFFEFFBE96B7A41FFEE94426" bold="true" box="[198,351,1599,1625]" pageId="4" pageNumber="55">Description.</emphasis>
<typeStatus id="521626E6FFFEFFBE9718A460FE604425" box="[361,470,1600,1626]" pageId="4" pageNumber="55" type="holotype">Holotype</typeStatus>
(male). Total body length:
<quantity id="4A5535A1FFFEFFBE9559A460FC304425" box="[808,902,1600,1626]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="55" unit="mm" value="6.6">6.6 mm</quantity>
(fully extended); maximal width:
<quantity id="4A5535A1FFFEFFBE935AA460FAC44425" box="[1323,1394,1600,1626]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="55" unit="mm" value="3.0">3 mm</quantity>
(at metacoxae level); humeral width:
<quantity id="4A5535A1FFFEFFBE9442A445FD244400" box="[563,658,1637,1663]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="55" unit="mm" value="2.4">2.4 mm</quantity>
; maximal height:
<quantity id="4A5535A1FFFEFFBE951AA445FC714400" box="[875,967,1637,1663]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="55" unit="mm" value="2.2">2.2 mm</quantity>
(at metacoxae level); length (body not extended)/height: 2.7; length/maximal width-humeral width: 11.
</paragraph>
<paragraph id="8D129844FFFEFFBE96B4A490FA99458B" blockId="4.[151,1437,1338,2036]" pageId="4" pageNumber="55">
<emphasis id="BFD94456FFFEFFBE96B4A490FEC544B6" box="[197,371,1712,1737]" italics="true" pageId="4" pageNumber="55">Body and form</emphasis>
. General body form (
<figureCitation id="159684C1FFFEFFBE9403A490FD0F44B5" box="[626,697,1712,1738]" captionStart="FIGURES 1 2" captionStartId="1.[151,269,1054,1078]" captionTargetBox="[318,1288,422,1015]" captionTargetId="figure@1.[290,1296,421,1029]" captionTargetPageId="1" captionText="FIGURES 1 2. Dorsal habitus: 1) Berosus degallieri (3.3 mm long); 2) Berosus spiniger (6.6 mm long)." httpUri="https://zenodo.org/record/195948/files/figure.png" pageId="4" pageNumber="55">Fig. 2</figureCitation>
) rather elongated and convex, with elytra weakly widened and pronotum rather narrow. Head entirely metallic and shiny bronze green. Pronotum and elytra basically testaceous and rather shiny, without metallic sheen, with several black areas: on pronotum disc narrowly on both sides of a median testaceous band, on elytra with indistinct spots distributed along a broad V-shaped basal band, a median band and an apical band. Suture, scutellum, striae except 10th, elytral and pronotal punctures (the latter on disc) also blackened. Ground punctures on head and pronotum rather uniform-sized but irregularly spaced in some places, fairly dense (spaced by 12 times their diameter), a little finer on clypeus than on frons, concealed on labrum under a thick fringe of long pale hairs. Besides the main punctures there are rather uniform micropunctures. A narrow, nearly impunctate band on pronotal median line.
</paragraph>
<paragraph id="8D129844FFFFFFBF96B4A2B8FD9742A8" blockId="5.[151,1437,152,215]" pageId="5" pageNumber="56">
<emphasis id="BFD94456FFFFFFBF96B4A2B8FEBF42CE" box="[197,265,152,177]" italics="true" pageId="5" pageNumber="56">Head</emphasis>
. Frons with a very fine longitudinal median carina. Maxillary palpi testaceous. Eyes rather prominent, ocular index OI = 2,4.
</paragraph>
<caption id="D9D2C8CCFFFFFFBF96E6A132FAE84156" box="[151,1374,786,810]" httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="5" pageNumber="56" targetBox="[494,1087,279,760]" targetPageId="5">
<paragraph id="8D129844FFFFFFBF96E6A132FAE84156" blockId="5.[151,1374,786,810]" box="[151,1374,786,810]" pageId="5" pageNumber="56">
<emphasis id="BFD94456FFFFFFBF96E6A132FEAF4155" bold="true" box="[151,281,786,810]" pageId="5" pageNumber="56">FIGURE 7.</emphasis>
Male genitalia of
<taxonomicName id="4AADE3C7FFFFFFBF97ADA132FD394156" box="[476,655,786,809]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="BFD94456FFFFFFBF97ADA132FD394156" box="[476,655,786,809]" italics="true" pageId="5" pageNumber="56">Berosus spiniger</emphasis>
</taxonomicName>
(1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect.
</paragraph>
</caption>
<caption id="D9D2C8CCFFFFFFBF96E6A489FDEF449E" httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="5" pageNumber="56" targetBox="[207,1398,860,1678]" targetPageId="5">
<paragraph id="8D129844FFFFFFBF96E6A489FDEF449E" blockId="5.[151,1436,1705,1761]" pageId="5" pageNumber="56">
<emphasis id="BFD94456FFFFFFBF96E6A489FEF544BE" bold="true" box="[151,323,1705,1729]" pageId="5" pageNumber="56">FIGURE 811.</emphasis>
Ventral details of
<taxonomicName id="4AADE3C7FFFFFFBF947EA48AFD7344BE" box="[527,709,1706,1729]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="BFD94456FFFFFFBF947EA48AFD7344BE" box="[527,709,1706,1729]" italics="true" pageId="5" pageNumber="56">Berosus spiniger</emphasis>
</taxonomicName>
. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite.
</paragraph>
</caption>
<paragraph id="8D129844FFFFFFBC96B4A52AFDE34058" blockId="5.[151,1437,1802,2014]" lastBlockId="6.[151,1437,152,551]" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56">
<emphasis id="BFD94456FFFFFFBF96B4A52AFEAD455C" box="[197,283,1802,1827]" italics="true" pageId="5" pageNumber="56">Thorax</emphasis>
. Pronotum with lateral angles rounded, the posterior ones more broadly so. Scutellum punctate. Elytra with ten fine striae, all well impressed, with the inner ones more deeply impressed than the outer ones, their punctures spaced by 1 to 2.5 times their own width, more closely set along the inner striae. Punctures along inner striae subequal in size to those on pronotum, the punctures becoming larger on the outer striae. Interstriae wide and almost flat, with punctures rather fine and dense, forming somewhat irregular rows, sometimes two or three in number, as large as those of first inner striae. Punctures on interstriae largely absent on apical part. Micropunctures very rare and only on inner interstriae. Scutellar stria with about twenty punctures. Systematic punctures not larger on interstriae 3rd, 5th and 7th but bearing long hairs on apical area. Short spine-like hairs on apical third of outer elytral edge. Long hairs on interstria 11th. Sutural angle of each elytron narrowly rounded. Ventral surface glossy and pubescent, mostly punctate, blackened except prosternum testaceous. Mesosternal process laminar with a rounded tooth, behind this an irregular, not very depressed ridge (
<figureCitation id="159684C1FFFCFFBC972FA372FE124313" box="[350,420,338,364]" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="6" pageNumber="57">Fig. 8</figureCitation>
). Metasternal process (
<figureCitation id="159684C1FFFCFFBC94C4A372FD4D4313" box="[693,763,338,364]" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="6" pageNumber="57">Fig. 9</figureCitation>
) triangular, narrowed backwards, with the postero-lateral angles strongly raised, produced and rounded, under abundant hairs, the posterior angle carinate and lower than postero-lateral angles. Legs testaceous except pubescent and blackish part of femora. Femoral pubescence obliquely limited and covering about 52% of the hind and 44% of the middle femora (
<figureCitation id="159684C1FFFCFFBC9345A3E2FA3843A3" box="[1332,1422,450,476]" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="6" pageNumber="57">Fig. 10</figureCitation>
). Male protarsi a little modified with the first basal segment widened and bearing expanded pad. Claws fine and very long, curved and toothed at base.
</paragraph>
<caption id="D9D2C8CCFFFCFFBC96E6A077FEC740F0" pageId="6" pageNumber="57">
<paragraph id="8D129844FFFCFFBC96E6A077FEC740F0" blockId="6.[151,1436,599,887]" pageId="6" pageNumber="57">
<emphasis id="BFD94456FFFCFFBC96E6A077FEBD4010" bold="true" box="[151,267,599,623]" pageId="6" pageNumber="57">TABLE 2.</emphasis>
Character matrix for identification of
<taxonomicName id="4AADE3C7FFFCFFBC94DDA078FCB44010" box="[684,770,600,623]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="BFD94456FFFCFFBC94DDA078FCB44010" box="[684,770,600,623]" italics="true" pageId="6" pageNumber="57">Berosus</emphasis>
</taxonomicName>
species in the
<emphasis id="BFD94456FFFCFFBC95D2A078FBBE4010" box="[931,1032,600,623]" italics="true" pageId="6" pageNumber="57">auriceps-</emphasis>
complex (from Oliva 1989, 1993 and present description).
</paragraph>
</caption>
<paragraph id="8D129844FFFCFFBC96EFA08AFAFE40BE" blockId="6.[151,1436,599,887]" box="[158,1352,682,705]" pageId="6" pageNumber="57">
Diagnostic characters
<emphasis id="BFD94456FFFCFFBC949CA08AFAFE40BE" box="[749,1352,682,705]" italics="true" pageId="6" pageNumber="57">
<taxonomicName id="4AADE3C7FFFCFFBC949CA08AFC9140BE" box="[749,807,682,705]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="aulus">aulus</taxonomicName>
<taxonomicName id="4AADE3C7FFFCFFBC95EAA08AFC4140BE" box="[923,1015,682,705]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="auriceps">auriceps</taxonomicName>
<taxonomicName id="4AADE3C7FFFCFFBC9224A08AFB6640BE" box="[1109,1232,682,705]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="ethmonotus">ethmonotus</taxonomicName>
<taxonomicName id="4AADE3C7FFFCFFBC9280A08AFAFE40BE" box="[1265,1352,682,705]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="spiniger">spiniger</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8D129844FFFCFFBC94FDA0EAFC94417E" blockId="6.[151,1436,599,887]" pageId="6" pageNumber="57">
Species O r c h y m o n t, Boheman, 1859
<bibRefCitation id="E93CE5B5FFFCFFBC9224A0EAFB64409E" author="Oliva" box="[1109,1234,714,737]" pageId="6" pageNumber="58" refString="Oliva, A. (1989) El genero Berosus (Coleoptera: Hydrophilidae) en America del Sur. Revista del Museo argentino de Ciencias naturales &quot; Bernardino Rivadavia &quot;, Entomologia 6 (4), 57 - 236." type="journal article" year="1989">Oliva, 1989</bibRefCitation>
Queney, sp. n. 1941
</paragraph>
<paragraph id="8D129844FFFCFFBC96EFA135FCB64128" blockId="6.[151,1436,599,887]" pageId="6" pageNumber="57">Length in mm 5.7 56.7 5.16.6 6.6 Pronotum: large medial spot, generally with metallic X</paragraph>
<paragraph id="8D129844FFFCFFBC96EFA140FF6D4108" blockId="6.[151,1436,599,887]" box="[158,219,864,887]" pageId="6" pageNumber="57">sheen</paragraph>
<paragraph id="8D129844FFFCFFBC96EFA1AAFAB24613" pageId="6" pageNumber="57">
<table id="FFAD6AE4FFFC004596E6A1AAFA2A4613" box="[151,1436,906,1132]" gridcols="4" gridrows="5" pageId="6" pageNumber="57">
<tr id="339D9A06FFFC004596E6A1AAFA2A41BE" box="[151,1436,906,961]" gridrow="0" pageId="6" pageNumber="57">
<th id="704CF37AFFFC004596E6A1AAFCB641BE" box="[151,768,906,961]" gridcol="0" gridrow="0" pageId="6" pageNumber="57">Pronotum: 2 small paramedial longitudinal spots without metallic sheen</th>
<th id="704CF37AFFFC004595EAA1AAFC1841BE" box="[923,942,906,961]" gridcol="1" gridrow="0" pageId="6" pageNumber="57">X</th>
<th id="704CF37AFFFC00459224A1AAFBDE41BE" box="[1109,1128,906,961]" gridcol="2" gridrow="0" pageId="6" pageNumber="57">X</th>
<th id="704CF37AFFFC00459280A1AAFA2A41BE" box="[1265,1436,906,961]" gridcol="3" gridrow="0" pageId="6" pageNumber="57">X</th>
</tr>
<tr id="339D9A06FFFC004596E6A1F5FA2A4193" box="[151,1436,981,1004]" gridrow="1" pageId="6" pageNumber="57" rowspan-1="1" rowspan-3="1">
<th id="704CF37AFFFC004596E6A1F5FCB64193" box="[151,768,981,1004]" gridcol="0" gridrow="1" pageId="6" pageNumber="57">Elytral interstriae: micropunctures well apparent X</th>
<td id="704CF37AFFFC00459224A1F5FBDE4193" box="[1109,1128,981,1004]" gridcol="2" gridrow="1" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596E6A620FA2A4668" box="[151,1436,1024,1047]" gridrow="2" pageId="6" pageNumber="57" rowspan-2="1">
<th id="704CF37AFFFC004596E6A620FCB64668" box="[151,768,1024,1047]" gridcol="0" gridrow="2" pageId="6" pageNumber="57">Elytral interstriae: micropunctures absent or very rare</th>
<td id="704CF37AFFFC004595EAA620FC184668" box="[923,942,1024,1047]" gridcol="1" gridrow="2" pageId="6" pageNumber="57">X</td>
<td id="704CF37AFFFC00459280A620FA2A4668" box="[1265,1436,1024,1047]" gridcol="3" gridrow="2" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596E6A60AFA2A463E" box="[151,1436,1066,1089]" gridrow="3" pageId="6" pageNumber="57" rowspan-2="1" rowspan-3="1">
<th id="704CF37AFFFC004596E6A60AFCB6463E" box="[151,768,1066,1089]" gridcol="0" gridrow="3" pageId="6" pageNumber="57">Mesosternal process: anterior tooth swollen</th>
<td id="704CF37AFFFC004595EAA60AFC18463E" box="[923,942,1066,1089]" gridcol="1" gridrow="3" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596E6A675FA2A4613" box="[151,1436,1109,1132]" gridrow="4" pageId="6" pageNumber="57" rowspan-1="1">
<th id="704CF37AFFFC004596E6A675FCB64613" box="[151,768,1109,1132]" gridcol="0" gridrow="4" pageId="6" pageNumber="57">Mesosternal process: entirely laminar X</th>
<td id="704CF37AFFFC00459224A675FBDE4613" box="[1109,1128,1109,1132]" gridcol="2" gridrow="4" pageId="6" pageNumber="57">X</td>
<td id="704CF37AFFFC00459280A675FA2A4613" box="[1265,1436,1109,1132]" gridcol="3" gridrow="4" pageId="6" pageNumber="57">X</td>
</tr>
</table>
</paragraph>
<paragraph id="8D129844FFFCFFBC96EFA6A0FAB247E8" pageId="6" pageNumber="57">
<table id="FFAD6AE4FFFC004596EFA6A0FAB247E8" box="[158,1284,1152,1431]" gridcols="5" gridrows="7" pageId="6" pageNumber="57">
<tr id="339D9A06FFFC004596EFA6A0FAB246E8" box="[158,1284,1152,1175]" gridrow="0" pageId="6" pageNumber="57" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="704CF37AFFFC004596EFA6A0FD9046E8" box="[158,550,1152,1175]" gridcol="0" gridrow="0" pageId="6" pageNumber="57">Basal piece/male genitalia:</th>
</tr>
<tr id="339D9A06FFFC004596EFA68AFAB246BE" box="[158,1284,1194,1217]" gridrow="1" pageId="6" pageNumber="57" rowspan-2="1" rowspan-4="1">
<th id="704CF37AFFFC004596EFA68AFD9046BE" box="[158,550,1194,1217]" gridcol="0" gridrow="1" pageId="6" pageNumber="57">-about 2/5</th>
<td id="704CF37AFFFC0045949CA68AFCB646BE" box="[749,768,1194,1217]" gridcol="1" gridrow="1" pageId="6" pageNumber="57">X</td>
<td id="704CF37AFFFC00459224A68AFBDE46BE" box="[1109,1128,1194,1217]" gridcol="3" gridrow="1" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596EFA6F5FAB24693" box="[158,1284,1237,1260]" gridrow="2" pageId="6" pageNumber="57" rowspan-1="1" rowspan-3="1">
<th id="704CF37AFFFC004596EFA6F5FD904693" box="[158,550,1237,1260]" gridcol="0" gridrow="2" pageId="6" pageNumber="57">-more than 1/2</th>
<td id="704CF37AFFFC004595EAA6F5FC184693" box="[923,942,1237,1260]" gridcol="2" gridrow="2" pageId="6" pageNumber="57">X</td>
<td id="704CF37AFFFC00459280A6F5FAB24693" box="[1265,1284,1237,1260]" gridcol="4" gridrow="2" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596EFA720FAB24768" box="[158,1284,1280,1303]" gridrow="3" pageId="6" pageNumber="57" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="704CF37AFFFC004596EFA720FD904768" box="[158,550,1280,1303]" gridcol="0" gridrow="3" pageId="6" pageNumber="57">Parameres (dorsal or ventral aspect):</th>
</tr>
<tr id="339D9A06FFFC004596EFA70AFAB2473E" box="[158,1284,1322,1345]" gridrow="4" pageId="6" pageNumber="57" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<th id="704CF37AFFFC004596EFA70AFD90473E" box="[158,550,1322,1345]" gridcol="0" gridrow="4" pageId="6" pageNumber="57">-apices divergent</th>
<td id="704CF37AFFFC00459224A70AFBDE473E" box="[1109,1128,1322,1345]" gridcol="3" gridrow="4" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596EFA775FAB24713" box="[158,1284,1365,1388]" gridrow="5" pageId="6" pageNumber="57" rowspan-3="1" rowspan-4="1">
<th id="704CF37AFFFC004596EFA775FD904713" box="[158,550,1365,1388]" gridcol="0" gridrow="5" pageId="6" pageNumber="57">-apices convergent</th>
<td id="704CF37AFFFC0045949CA775FCB64713" box="[749,768,1365,1388]" gridcol="1" gridrow="5" pageId="6" pageNumber="57">X</td>
<td id="704CF37AFFFC004595EAA775FC184713" box="[923,942,1365,1388]" gridcol="2" gridrow="5" pageId="6" pageNumber="57">X</td>
</tr>
<tr id="339D9A06FFFC004596EFA7A0FAB247E8" box="[158,1284,1408,1431]" gridrow="6" pageId="6" pageNumber="57" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="704CF37AFFFC004596EFA7A0FD9047E8" box="[158,550,1408,1431]" gridcol="0" gridrow="6" pageId="6" pageNumber="57">-apices parallel</th>
<td id="704CF37AFFFC00459280A7A0FAB247E8" box="[1265,1284,1408,1431]" gridcol="4" gridrow="6" pageId="6" pageNumber="57">X</td>
</tr>
</table>
</paragraph>
<paragraph id="8D129844FFFCFFBC96B4A7FDFE65455D" blockId="6.[151,1437,1501,2012]" pageId="6" pageNumber="57">
<emphasis id="BFD94456FFFCFFBC96B4A7FDFE824789" box="[197,308,1501,1526]" italics="true" pageId="6" pageNumber="57">Abdomen</emphasis>
. Lateral edges of ventrites evenly serrate. First ventrite without lateral depressions, with median carina on anterior half. Fifth ventrite with an apical notch, the lateral edges of which are not convergent, bottom produced into a tooth with a bidentate apex (
<figureCitation id="159684C1FFFCFFBC948FA408FCE2443D" box="[766,852,1576,1602]" captionStart="FIGURE 8 11" captionStartId="5.[151,256,1705,1729]" captionTargetBox="[207,1398,860,1678]" captionTargetId="figure@5.[188,1398,860,1684]" captionTargetPageId="5" captionText="FIGURE 8 11. Ventral details of Berosus spiniger. 8) Mesosternal process; 9) Meso- and metasternal processes; 10) Meso- and metafemora; 11) Fifth ventrite." httpUri="https://zenodo.org/record/195952/files/figure.png" pageId="6" pageNumber="57">Fig. 11</figureCitation>
). Male genitalia (
<figureCitation id="159684C1FFFCFFBC9255A408FBDC443D" box="[1060,1130,1576,1602]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
) subcylindrical and thick, ventral face somewhat concave and dorsal face strongly convex in lateral aspect (
<figureCitation id="159684C1FFFCFFBC922AA46DFB154418" box="[1115,1187,1613,1639]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
a), distal part beyond basal piece very narrow in dorsal and ventral aspect. Total length:
<quantity id="4A5535A1FFFCFFBC95E9A452FC4344F3" box="[920,1013,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="57" unit="mm" value="1.6">1.6 mm</quantity>
, basal piece representing about 60% of it. Parameres broad at base in lateral aspect, strongly and evenly narrowed, with apex curved towards ventral face (
<figureCitation id="159684C1FFFCFFBC9742A49DFECD44A8" box="[307,379,1725,1751]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
a). Median lobe much shorter than parameres, bent towards dorsal face which it overlaps with its turned-up apex (
<figureCitation id="159684C1FFFCFFBC97C6A4C2FE494483" box="[439,511,1762,1788]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="6" pageNumber="57">Fig. 7</figureCitation>
a). In dorsal and ventral aspects, parameres rounded up inwards, ending in two narrow and parallel apices.
</paragraph>
</subSubSection>
<subSubSection id="C5B7CBCFFFFCFFBD96B7A50DFD524313" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57" type="discussion">
<paragraph id="8D129844FFFCFFBD96B7A50DFD524313" blockId="6.[151,1437,1501,2012]" lastBlockId="7.[151,1436,152,476]" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
<emphasis id="BFD94456FFFCFFBC96B7A50DFEE74538" bold="true" box="[198,337,1837,1863]" pageId="6" pageNumber="57">Discussion.</emphasis>
As shown in
<tableCitation id="C02FADFFFFFCFFBC9472A50DFDE94538" box="[515,607,1837,1863]" captionStart="TABLE 2" captionStartId="6.[151,240,599,623]" captionText="TABLE 2. Character matrix for identification of Berosus species in the auriceps- complex (from Oliva 1989, 1993 and present description)." pageId="6" pageNumber="57">Table 2</tableCitation>
, examination of male genitalia allows
<taxonomicName id="4AADE3C7FFFCFFBC924DA50DFB764539" box="[1084,1216,1837,1862]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="BFD94456FFFCFFBC924DA50DFB764539" box="[1084,1216,1837,1862]" italics="true" pageId="6" pageNumber="57">B. spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4EAF92DFFFCFFBC92BDA50DFAB54538" box="[1228,1283,1837,1863]" pageId="6" pageNumber="57" rank="species">sp.n.</taxonomicNameLabel>
to be easily distinguished from the three other species already known within the
<taxonomicName id="4AADE3C7FFFCFFBC95B1A573FB934513" box="[960,1061,1875,1900]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFCFFBC95B1A573FB934513" box="[960,1061,1875,1900]" italics="true" pageId="6" pageNumber="57">auriceps</emphasis>
</taxonomicName>
-complex. The basal piece of
<taxonomicName id="4AADE3C7FFFCFFBC93F2A573FE0645ED" authority="Orchymont, 1941" authorityName="Orchymont" authorityYear="1941" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="aulus">
<emphasis id="BFD94456FFFCFFBC93F2A573FF6145EE" italics="true" pageId="6" pageNumber="57">B. aulus</emphasis>
Orchymont, 1941
</taxonomicName>
and
<taxonomicName id="4AADE3C7FFFCFFBC979DA558FC9D45ED" authority="Oliva, 1989" authorityName="Oliva" authorityYear="1989" box="[492,811,1912,1938]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="ethmonotus">
<emphasis id="BFD94456FFFCFFBC979DA558FE4845EE" box="[492,510,1912,1937]" italics="true" pageId="6" pageNumber="57">B</emphasis>
.
<emphasis id="BFD94456FFFCFFBC947CA558FD2045EE" box="[525,662,1912,1937]" italics="true" pageId="6" pageNumber="57">ethmonotus</emphasis>
<bibRefCitation id="E93CE5B5FFFCFFBC94EFA558FC9D45ED" author="Oliva" box="[670,811,1912,1938]" pageId="6" pageNumber="58" refString="Oliva, A. (1989) El genero Berosus (Coleoptera: Hydrophilidae) en America del Sur. Revista del Museo argentino de Ciencias naturales &quot; Bernardino Rivadavia &quot;, Entomologia 6 (4), 57 - 236." type="journal article" year="1989">Oliva, 1989</bibRefCitation>
</taxonomicName>
is short and represents less than 45% of total length against nearly 60% in
<taxonomicName id="4AADE3C7FFFCFFBC97EAA5BDFDAA45C9" box="[411,540,1949,1974]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="BFD94456FFFCFFBC97EAA5BDFDAA45C9" box="[411,540,1949,1974]" italics="true" pageId="6" pageNumber="57">B. spiniger</emphasis>
</taxonomicName>
and
<taxonomicName id="4AADE3C7FFFCFFBC9425A5BDFC2A45C8" authority="Boheman, 1859" authorityName="Boheman" authorityYear="1859" box="[596,924,1949,1975]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFCFFBC9425A5BDFD6F45C9" box="[596,729,1949,1974]" italics="true" pageId="6" pageNumber="57">B. auriceps</emphasis>
Boheman, 1859
</taxonomicName>
. In lateral aspect the parameres are not very different in the last two species but, in dorsal aspect, they are quite distinctive: in
<taxonomicName id="4AADE3C7FFFCFFBC9208A5E3FAB545A3" box="[1145,1283,1987,2012]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFCFFBC9208A5E3FAB545A3" box="[1145,1283,1987,2012]" italics="true" pageId="6" pageNumber="57">B. auriceps</emphasis>
</taxonomicName>
they narrow gradually towards the apex and curve regularly inwards while in
<taxonomicName id="4AADE3C7FFFDFFBD95D9A2B8FB9A42CE" box="[936,1068,152,177]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="BFD94456FFFDFFBD95D9A2B8FB9A42CE" box="[936,1068,152,177]" italics="true" pageId="7" pageNumber="58">B. spiniger</emphasis>
</taxonomicName>
(
<figureCitation id="159684C1FFFDFFBD924CA2B8FB3342CD" box="[1085,1157,152,178]" captionStart="FIGURE 7" captionStartId="5.[151,255,786,810]" captionTargetBox="[494,1087,279,760]" captionTargetId="figure@5.[493,1095,271,761]" captionTargetPageId="5" captionText="FIGURE 7. Male genitalia of Berosus spiniger (1.6 mm long). a) lateral aspect; b) dorsal aspect; c) ventral aspect." httpUri="https://zenodo.org/record/195951/files/figure.png" pageId="7" pageNumber="58">Fig. 7</figureCitation>
b) the curved outline is interrupted by an emargination before the apex, with the apices parallel. Other differential characters include: the shape of anterior tooth of mesosternal process (swollen in
<taxonomicName id="4AADE3C7FFFDFFBD9516A2C2FC5A4284" box="[871,1004,226,251]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFDFFBD9516A2C2FC5A4284" box="[871,1004,226,251]" italics="true" pageId="7" pageNumber="58">B. auriceps</emphasis>
</taxonomicName>
, laminar in the other species), elytral micropunctures well apparent in
<taxonomicName id="4AADE3C7FFFDFFBD946DA328FDC8435E" box="[540,638,264,289]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="aulus">
<emphasis id="BFD94456FFFDFFBD946DA328FDC8435E" box="[540,638,264,289]" italics="true" pageId="7" pageNumber="58">B. aulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4AADE3C7FFFDFFBD94C9A328FCD6435E" box="[696,864,264,289]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="ethmonotus">
<emphasis id="BFD94456FFFDFFBD94C9A328FCD6435E" box="[696,864,264,289]" italics="true" pageId="7" pageNumber="58">B. ethmonotus</emphasis>
</taxonomicName>
but not in
<taxonomicName id="4AADE3C7FFFDFFBD9592A328FBDF435E" box="[995,1129,264,289]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFDFFBD9592A328FBDF435E" box="[995,1129,264,289]" italics="true" pageId="7" pageNumber="58">B. auriceps</emphasis>
</taxonomicName>
and
<taxonomicName id="4AADE3C7FFFDFFBD92D5A328FA92435E" box="[1188,1316,264,289]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="BFD94456FFFDFFBD92D5A328FA92435E" box="[1188,1316,264,289]" italics="true" pageId="7" pageNumber="58">B. spiniger</emphasis>
</taxonomicName>
, extent of femoral pubescence, larger in
<taxonomicName id="4AADE3C7FFFDFFBD9789A30DFDCE4339" box="[504,632,301,326]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="BFD94456FFFDFFBD9789A30DFDCE4339" box="[504,632,301,326]" italics="true" pageId="7" pageNumber="58">B. spiniger</emphasis>
</taxonomicName>
than in
<taxonomicName id="4AADE3C7FFFDFFBD94A6A30DFCEA4339" box="[727,860,301,326]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="auriceps">
<emphasis id="BFD94456FFFDFFBD94A6A30DFCEA4339" box="[727,860,301,326]" italics="true" pageId="7" pageNumber="58">B. auriceps</emphasis>
</taxonomicName>
and
<taxonomicName id="4AADE3C7FFFDFFBD95E4A30DFB8A4339" box="[917,1084,301,326]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="ethmonotus">
<emphasis id="BFD94456FFFDFFBD95E4A30DFB8A4339" box="[917,1084,301,326]" italics="true" pageId="7" pageNumber="58">B. ethmonotus</emphasis>
</taxonomicName>
, and color of the pronotum of
<taxonomicName id="4AADE3C7FFFDFFBD96E6A372FEAC4314" box="[151,282,338,363]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="BFD94456FFFDFFBD96E6A372FEAC4314" box="[151,282,338,363]" italics="true" pageId="7" pageNumber="58">B. Spiniger</emphasis>
</taxonomicName>
without the metallic sheen of
<taxonomicName id="4AADE3C7FFFDFFBD940CA372FD6B4314" box="[637,733,338,363]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="aulus">
<emphasis id="BFD94456FFFDFFBD940CA372FD6B4314" box="[637,733,338,363]" italics="true" pageId="7" pageNumber="58">B. aulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C5B7CBCFFFFDFFBD96B7A357FBDD43ED" box="[198,1131,375,402]" pageId="7" pageNumber="58" type="etymology">
<paragraph id="8D129844FFFDFFBD96B7A357FBDD43ED" blockId="7.[151,1436,152,476]" box="[198,1131,375,402]" pageId="7" pageNumber="58">
<emphasis id="BFD94456FFFDFFBD96B7A357FEE543EE" bold="true" box="[198,339,375,401]" pageId="7" pageNumber="58">Etymology.</emphasis>
Named for the spine-like hairs on apical third of outer elytral edge.
</paragraph>
</subSubSection>
<subSubSection id="C5B7CBCFFFFDFFBD96B7A3BDFEA343A3" pageId="7" pageNumber="58" type="distribution">
<paragraph id="8D129844FFFDFFBD96B7A3BDFEA343A3" blockId="7.[151,1436,152,476]" pageId="7" pageNumber="58">
<emphasis id="BFD94456FFFDFFBD96B7A3BDFED143C8" bold="true" box="[198,359,413,439]" pageId="7" pageNumber="58">Distribution.</emphasis>
This species is only known from
<collectingCountry id="F5BAD8D4FFFDFFBD948EA3BDFCEC43C8" box="[767,858,413,439]" name="Brazil" pageId="7" pageNumber="58">Brasilia</collectingCountry>
(
<collectingCountry id="F5BAD8D4FFFDFFBD951DA3BDFC0143C8" box="[876,951,413,439]" name="Brazil" pageId="7" pageNumber="58">Brazil</collectingCountry>
) where it was collected at a light trap in &quot;Cerrado&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>