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<document id="C71EDD809BA0238489D6D7D1D1A241D2" ID-CLB-Dataset="7214" ID-DOI="10.11646/zootaxa.4963.1.2" ID-GBIF-Dataset="78e78beb-b1ed-43a4-84c7-8180e61463c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4711249" ID-ZooBank="E93AB178-3419-498B-9563-A704ABAE5EB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618560083953" checkinUser="plazi" docAuthor="Bálint, Zsolt, Boyer, Pierre, Cerdeña, José, Larico, Jackie Farfán, Brudecka, Jadwiga Lorenc-, Prieto, Carlos &amp; Pyrcz, Tomasz W." docDate="2021" docId="D5008F50FFFAB31FFF76FF31959FFBDC" docLanguage="en" docName="zootaxa.4963.1.2.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Johnsonita oxalida Balint, Boyer &amp; Pyrcz 2021, sp. n." docType="treatment" docVersion="6" lastPageNumber="30" masterDocId="2939F728FFE9B30CFFE1FFA69629FFCD" masterDocTitle="Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)" masterLastPageNumber="57" masterPageNumber="11" pageNumber="30" updateTime="1698937134920" updateUser="ExternalLinkService">
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<mods:title id="335EB27611446FFC2BD7446171AE6C50">Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)</mods:title>
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<mods:namePart id="96A9D423BE62919BBF7CF6CAFD5F5D53">Bálint, Zsolt</mods:namePart>
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<mods:affiliation id="8E743828D68FC18974A75CFF7FEFCCBE">Hungarian Natural History Museum, Department of Zoology, H- 1088 Budapest, Baross utca 13, Hungary. balint. zsolt @ nhmus. hu; https: // orcid. org / 0000 - 0001 - 8174 - 878 X</mods:affiliation>
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<mods:namePart id="E3D6F5EE877FC0D2661F0A42927701F6">Boyer, Pierre</mods:namePart>
<mods:affiliation id="9E4C511A05A24E791501378A0CBDDBF3">Lotissement lHorizon, 13610 Le Puy Sainte Réparade, France.</mods:affiliation>
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<mods:namePart id="AB58EF2754B9D7CD86548280ACEC4468">Cerdeña, José</mods:namePart>
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<mods:affiliation id="CA0ECC6F19084FA17AD45624CEEA79C4">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="C7202C67D4C747F6167FE48337446866" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
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<mods:namePart id="443313EDE4A63F8D2DB07DE098525470">Larico, Jackie Farfán</mods:namePart>
<mods:nameIdentifier id="3F0FDE68D6055C54EA877D85D4A72D45" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="3057C01FDEA0A50AAE52D0E0BD929A35">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="3DA1386A54B9A447664635405991640F" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
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<mods:namePart id="14C9728FBCA40C6D247CAB16AD41DCEC">Brudecka, Jadwiga Lorenc-</mods:namePart>
<mods:affiliation id="159109C38F7107C897311E3CFB51F626">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:namePart id="310B6F00AD08FDA0656F15A50E98EFEC">Prieto, Carlos</mods:namePart>
<mods:affiliation id="C76852F1C2832105B672AC4554C853A2">Corporación Universitaria Autónoma del Cauca, Popayán, Colombia.</mods:affiliation>
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<mods:namePart id="DFED3E7628F933FFA29E38AA9DE4B95C">Pyrcz, Tomasz W.</mods:namePart>
<mods:affiliation id="AEE6FE53959D9EE97C5B4CA0F3924001">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland.</mods:affiliation>
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<treatment id="D5008F50FFFAB31FFF76FF31959FFBDC" ID-DOI="http://doi.org/10.5281/zenodo.4696427" ID-GBIF-Taxon="181531935" ID-Zenodo-Dep="4696427" LSID="urn:lsid:plazi:treatment:D5008F50FFFAB31FFF76FF31959FFBDC" httpUri="http://treatment.plazi.org/id/D5008F50FFFAB31FFF76FF31959FFBDC" lastPageNumber="30" pageId="19" pageNumber="30">
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<heading id="065E892AFFFAB31FFF76FF3194C5FF7C" bold="true" box="[151,748,150,178]" fontSize="11" level="1" pageId="19" pageNumber="30" reason="1">
<emphasis id="6FDDE254FFFAB31FFF76FF3194C5FF7C" bold="true" box="[151,748,150,178]" pageId="19" pageNumber="30">
<taxonomicName id="9AA945C5FFFAB31FFF76FF3194B4FF7C" authority="Balint, Boyer &amp; Pyrcz" authorityName="Balint, Boyer &amp; Pyrcz" authorityYear="2021" box="[151,669,150,178]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="30" phylum="Arthropoda" rank="species" species="oxalida" status="sp. nov.">
<emphasis id="6FDDE254FFFAB31FFF76FF3197A9FF7F" bold="true" box="[151,384,151,178]" italics="true" pageId="19" pageNumber="30">Johnsonita oxalida</emphasis>
Bálint, Boyer &amp; Pyrcz
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,
<taxonomicNameLabel id="74EE5F2FFFFAB31FFD48FF3194C5FF7C" box="[681,748,151,177]" pageId="19" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="5D163E46FFFAB31FFF76FF1D97FBFF18" blockId="19.[151,748,150,213]" box="[151,466,187,213]" pageId="19" pageNumber="30">
(
<figureCitation id="C59222C3FFFAB31FFF7EFF1D96DCFF18" box="[159,245,187,213]" captionStart="FIGURE 11" captionStartId="18.[151,250,1911,1935]" captionTargetBox="[214,1386,715,1874]" captionTargetId="figure-229@18.[151,1436,705,1888]" captionTargetPageId="18" captionText="FIGURE 11. Johnsonita oxalida Bálint, Boyer &amp; Pyrcz, sp. n. imagines (left column: recto, right column: verso). AB: holotype male (Peru: Pasco) (MUSM), CD: paratype female (Peru: Huánuco) (CEPMZUJ) (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711273" httpUri="https://zenodo.org/record/4711273/files/figure.png" pageId="19" pageNumber="30">
Figs
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</figureCitation>
,
<figureCitation id="C59222C3FFFAB31FFEE0FF1D9765FF18" box="[257,332,187,213]" captionStart="FIGURE 19" captionStartId="29.[151,250,1909,1933]" captionTargetBox="[313,1283,191,1883]" captionTargetId="figure-18@29.[200,1388,181,1885]" captionTargetPageId="29" captionText="FIGURE 19. Johnsonita genitalia armatures; entire male organs in lateral (left column) and ventral (right column) views.AB: J. subcunicula, Peru, Huánuco, San Pedro de Carpish; CD: J. turquisca, Ecuador, Napo, Papallacta, Chalpi Grande; EF: J. oxalida, Peru, Pasco, Oxapampa; GH: J. zubkova, Peru, Cuzco, Abra Acjanaco vers Pillcopata (CEPMZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711295" httpUri="https://zenodo.org/record/4711295/files/figure.png" pageId="19" pageNumber="30">19EF</figureCitation>
,
<figureCitation id="C59222C3FFFAB31FFEB6FF1D9788FF18" box="[343,417,187,213]" captionStart="FIGURE 22" captionStartId="32.[151,250,1909,1933]" captionTargetBox="[183,1405,181,1885]" captionTargetId="figure-23@32.[183,1405,181,1885]" captionTargetPageId="32" captionText="FIGURE 22. Entire Johnsonita female organs in lateral (left column), ductus in lateral (central column) and ventral (right column) views. AC: J. pardoa, Ecuador, Tungurahua, Laguna de San Borja; DF: J. turquisca, Ecuador, Sucumbios, El CalvarioLa Bonita; GI: J. oxalida, Peru, Huánuco, Paso Carpish; JL: J. zubkova, Peru, Puno, Carcel Punco (CEPMZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711303" httpUri="https://zenodo.org/record/4711303/files/figure.png" pageId="19" pageNumber="30">22GI</figureCitation>
,
<figureCitation id="C59222C3FFFAB31FFE4CFF1D97E3FF18" box="[429,458,187,213]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="19" pageNumber="30">30</figureCitation>
)
</paragraph>
<paragraph id="5D163E46FFFAB31FFF76FEA29312FEAB" blockId="19.[151,1437,259,1042]" pageId="19" pageNumber="30">
<emphasis id="6FDDE254FFFAB31FFF76FEA29710FED0" box="[151,313,259,285]" italics="true" pageId="19" pageNumber="30">Type material.</emphasis>
<materialsCitation id="EDC1341BFFFAB31FFEA1FEA59311FEAB" ID-GBIF-Occurrence="3083794304" collectingDate="2003-02-19" collectingDateMax="2009-10-20" collectingDateMin="2003-02-19" collectionCode="CB, CEP, MZUJ" collectorName="J. Lorenc" country="Peru" elevation="2350" location="Paso Carpish" pageId="19" pageNumber="30" specimenCount="2" stateProvince="Pasco" typeStatus="holotype">
<collectionCode id="3BB8A683FFFAB31FFEA1FEA5974CFED0" box="[320,357,259,285]" country="Australia" name="The CB Rhizobium Collection" pageId="19" pageNumber="30">CB</collectionCode>
<typeStatus id="821280E4FFFAB31FFE8DFEA597E7FED0" box="[364,462,259,285]" pageId="19" pageNumber="30" type="holotype">holotype</typeStatus>
male (will be deposited in
<collectionCode id="3BB8A683FFFAB31FFD1EFEA5951AFED0" box="[767,819,259,285]" pageId="19" pageNumber="30">CEP</collectionCode>
<collectionCode id="3BB8A683FFFAB31FFCA0FEA595A2FED0" box="[833,907,259,285]" pageId="19" pageNumber="30">MZUJ</collectionCode>
) in good condition, labelled as “
<quantity id="9A5193A3FFFAB31FFB17FEA593BEFED3" box="[1270,1431,259,286]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="19" pageNumber="30" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">
<elevation id="D684D975FFFAB31FFB17FEA593BEFED3" box="[1270,1431,259,286]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="19" pageNumber="30" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">2200- 2500 m</elevation>
</quantity>
, [//] la Antena, [//] SE Oxapampa (
<collectingRegion id="9F6DF0A4FFFAB31FFDFEFE81944DFE8C" box="[543,612,295,321]" country="Peru" name="Pasco" pageId="19" pageNumber="30">Pasco</collectingRegion>
) [//] Pérou
<date id="29171886FFFAB31FFD0BFE819546FE8C" box="[746,879,295,321]" pageId="19" pageNumber="30" value="2009-10-20">
<collectingDate id="3953E16EFFFAB31FFD0BFE819546FE8C" box="[746,879,295,321]" pageId="19" pageNumber="30" value="2009-10-20">20-10-2009</collectingDate>
</date>
” (dissected. gen. prep. 893/
<collectorName id="F05C5B90FFFAB31FFB53FE819337FE8F" box="[1202,1310,295,322]" pageId="19" pageNumber="30">J. Lorenc</collectorName>
).
<typeStatus id="821280E4FFFAB31FFAD2FE8E93B0FE8F" box="[1331,1433,296,322]" pageId="19" pageNumber="30" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="25BE7ED6FFFAB31FFF76FEEA96E4FEAB" box="[151,205,332,358]" name="Peru" pageId="19" pageNumber="30">Peru</collectingCountry>
,
<collectingRegion id="9F6DF0A4FFFAB31FFF38FEED9768FEA8" box="[217,321,331,357]" country="Peru" name="Huanuco" pageId="19" pageNumber="30">Huánuco</collectingRegion>
,
<location id="5876689DFFFAB31FFEADFEEA97F7FEA8" LSID="urn:lsid:plazi:treatment:D5008F50FFFAB31FFF76FF31959FFBDC:5876689DFFFAB31FFEADFEEA97F7FEA8" box="[332,478,331,358]" country="Peru" name="Paso Carpish" pageId="19" pageNumber="30" stateProvince="Pasco">Paso Carpish</location>
<quantity id="9A5193A3FFFAB31FFE07FEED9457FEAB" box="[486,638,331,358]" metricMagnitude="3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="19" pageNumber="30" unit="m" value="2900.0" valueMax="3000.0" valueMin="2800.0">
<elevation id="D684D975FFFAB31FFE07FEED9457FEAB" box="[486,638,331,358]" metricMagnitude="3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="19" pageNumber="30" unit="m" value="2900.0" valueMax="3000.0" valueMin="2800.0">2800-3000 m</elevation>
</quantity>
,
<date id="29171886FFFAB31FFD6BFEED952FFEA8" box="[650,774,331,357]" pageId="19" pageNumber="30" value="2003-02-19">
<collectingDate id="3953E16EFFFAB31FFD6BFEED952FFEA8" box="[650,774,331,357]" pageId="19" pageNumber="30" value="2003-02-19">2003.II.19.</collectingDate>
</date>
(
<collectionCode id="3BB8A683FFFAB31FFCF4FEED9516FEA8" box="[789,831,331,357]" country="Australia" name="The CB Rhizobium Collection" pageId="19" pageNumber="30">CB</collectionCode>
: female; dissected; gen. prep. 894/
<collectorName id="F05C5B90FFFAB31FFB24FEED9306FEAB" box="[1221,1327,331,358]" pageId="19" pageNumber="30">J. Lorenc</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5D163E46FFFAB31FFF26FED695C8FD00" blockId="19.[151,1437,259,1042]" pageId="19" pageNumber="30">
<emphasis id="6FDDE254FFFAB31FFF26FED6977AFE44" box="[199,339,368,393]" italics="true" pageId="19" pageNumber="30">Description.</emphasis>
Wings. FW costa length:
<quantity id="9A5193A3FFFAB31FFD9FFEC994D0FE47" box="[638,761,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="19" pageNumber="30" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">1415 mm</quantity>
(n=2), apical area of costa and outer margin is not straight resulting in a somewhat rounded shape; dorsal wing surface colouration pale violet blue, black border inner margin reaching the erection of vein M3; HW dorsal surface discal area structurally coloured; ventral FW submedial and medial pattern running parallel, submarginal area conspicuously lighter than medial and basal regions, antemargin area brown; HW medial pattern composed by ruptive band brown coloured bordered basally by a thin white line, submarginal area conspicuously lighter than medial and basal regions, antemargin area brown. Genitalia. Male valva thin with rounded costal margin flip and extremely pointed valval terminus (with valval length) and strong saccus. Female ductus thin with parallel edges but at 2/3 length divergent, no lower plate, upper plate symmetric and large with thinly sclerotized frame with a single spine. Androconia present in FW ventral surface basal are with cubital scent patch and HW dorsal wing surface medial area with black fur.
</paragraph>
<paragraph id="5D163E46FFFAB31FFF26FD7E9551FC90" blockId="19.[151,1437,259,1042]" pageId="19" pageNumber="30">
<emphasis id="6FDDE254FFFAB31FFF26FD7E9769FD3C" box="[199,320,728,753]" italics="true" pageId="19" pageNumber="30">Diagnosis.</emphasis>
On the basis of the dorsal HW surface “fur” and the ventral HW surface zigzagged pattern contrasted it is easy to distinguish the species from the congeners. The similar species
<taxonomicName id="9AA945C5FFFAB31FFC37FD5A9243FCD8" authorityName="Balint" authorityYear="2003" box="[982,1130,763,789]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="30" phylum="Arthropoda" rank="species" species="johnbanksi">
<emphasis id="6FDDE254FFFAB31FFC37FD5A9243FCD8" box="[982,1130,763,789]" italics="true" pageId="19" pageNumber="30">J. johnbanksi</emphasis>
</taxonomicName>
is more vivid blue dorsally, the ventral wing surface submarginal areas are not so light, the medial zigzagged pattern is more ruptive and the submedian and basal area have a gleaming white scales suffusion.
</paragraph>
<paragraph id="5D163E46FFFAB31FFF26FCC194A2FC68" blockId="19.[151,1437,259,1042]" pageId="19" pageNumber="30">
<emphasis id="6FDDE254FFFAB31FFF26FCC19771FC4C" box="[199,344,871,897]" italics="true" pageId="19" pageNumber="30">Distribution.</emphasis>
Geographical:
<collectingCountry id="25BE7ED6FFFAB31FFDE7FCCE9467FC4F" box="[518,590,872,898]" name="Peru" pageId="19" pageNumber="30">PERU</collectingCountry>
(
<collectingRegion id="9F6DF0A4FFFAB31FFDBFFCC194E1FC4C" box="[606,712,871,897]" country="Peru" name="Huanuco" pageId="19" pageNumber="30">Huánuco</collectingRegion>
,
<collectingRegion id="9F6DF0A4FFFAB31FFD35FCCE953EFC4F" box="[724,791,872,898]" country="Peru" name="Pasco" pageId="19" pageNumber="30">Pasco</collectingRegion>
) (
<figureCitation id="C59222C3FFFAB31FFCCEFCC195ACFC4C" box="[815,901,871,897]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="19" pageNumber="30">Fig. 30</figureCitation>
); spatial: recorded between
<quantity id="9A5193A3FFFAB31FFB22FCC19376FC4F" box="[1219,1375,871,898]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="19" pageNumber="30" unit="m" value="2600.0" valueMax="3000.0" valueMin="2200.0">22003000 m</quantity>
a.s.l; temporal: recorded in February and October.
</paragraph>
<paragraph id="5D163E46FFFAB31FFF26FC09959FFBDC" blockId="19.[151,1437,259,1042]" pageId="19" pageNumber="30">
<emphasis id="6FDDE254FFFAB31FFF26FC09976CFC04" box="[199,325,943,969]" italics="true" pageId="19" pageNumber="30">Etymology.</emphasis>
The name “oxalida” is used to emphasize the
<typeStatus id="821280E4FFFAB31FFCA3FC16955BFC07" box="[834,882,944,970]" pageId="19" pageNumber="30">type</typeStatus>
locality “Oxapampa” and the observation that the species is lighter (= “pallida” in Latin) than the putative sister species
<taxonomicName id="9AA945C5FFFAB31FFC46FC729214FC20" authorityName="Balint" authorityYear="2003" box="[935,1085,979,1005]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="30" phylum="Arthropoda" rank="species" species="johnbanksi">
<emphasis id="6FDDE254FFFAB31FFC46FC729214FC20" box="[935,1085,979,1005]" italics="true" pageId="19" pageNumber="30">J. johnbanksi</emphasis>
</taxonomicName>
. The species name “oxalida” is freely formed Latin name and is treated as a female noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>