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<document id="A4914E979A34B631DFD17112F755DFF0" ID-CLB-Dataset="7214" ID-DOI="10.11646/zootaxa.4963.1.2" ID-GBIF-Dataset="78e78beb-b1ed-43a4-84c7-8180e61463c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4711249" ID-ZooBank="E93AB178-3419-498B-9563-A704ABAE5EB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618560083953" checkinUser="plazi" docAuthor="Bálint, Zsolt, Boyer, Pierre, Cerdeña, José, Larico, Jackie Farfán, Brudecka, Jadwiga Lorenc-, Prieto, Carlos &amp; Pyrcz, Tomasz W." docDate="2021" docId="D5008F50FFE6B31CFF76FBC99221FE1C" docLanguage="en" docName="zootaxa.4963.1.2.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Johnsonita johnbanksi Balint 2003" docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="2939F728FFE9B30CFFE1FFA69629FFCD" masterDocTitle="Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)" masterLastPageNumber="57" masterPageNumber="11" pageNumber="26" updateTime="1698937134920" updateUser="ExternalLinkService">
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<mods:title id="D91B181F4C969143553125F335496B44">Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar &amp; Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)</mods:title>
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<mods:name id="7AF7FCF981A39835056AF3ABBF77F144" type="personal">
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<mods:namePart id="20BE2E25E2190FF8EA687B774B3A44E9">Bálint, Zsolt</mods:namePart>
<mods:nameIdentifier id="3D2BF684A13DB0ABF877CD7062358671" type="ORCID">0000-0001-8174-878X</mods:nameIdentifier>
<mods:affiliation id="529FAAF8A3718E667746547E76F447F4">Hungarian Natural History Museum, Department of Zoology, H- 1088 Budapest, Baross utca 13, Hungary. balint. zsolt @ nhmus. hu; https: // orcid. org / 0000 - 0001 - 8174 - 878 X</mods:affiliation>
<mods:nameIdentifier id="F423B7AB50006233249592DF3B9942FD" type="email">balint.zsolt@nhmus.hu</mods:nameIdentifier>
</mods:name>
<mods:name id="24EB7EC45FFE88ECFE5EE1A60D0D5259" type="personal">
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<mods:namePart id="DD33CFEB709521FF36C1578011FBC9B3">Boyer, Pierre</mods:namePart>
<mods:affiliation id="530E4F9F3A09627DB4C42B086BA8DAD5">Lotissement l’Horizon, 13610 Le Puy Sainte Réparade, France.</mods:affiliation>
</mods:name>
<mods:name id="61107484D3C81186133CEF716C376001" type="personal">
<mods:role id="29A9791348496A604A974C78145205FB">
<mods:roleTerm id="775E2957949031B12DCD6801E9BFDEC3">Author</mods:roleTerm>
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<mods:namePart id="B131479C231DFF2BB1D072A893C3926C">Cerdeña, José</mods:namePart>
<mods:nameIdentifier id="52136677F57E5C9C3717D6F96B0D78CF" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="96576F7B4FA5F8806EEF34D1A7E76969">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="A978F402414B1A5A417E5C73B69DC2BB" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
</mods:name>
<mods:name id="DDC4D3E4DB113F6D0C9F8F397F5F26EF" type="personal">
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<mods:roleTerm id="5D907D64B67BDAE31620DFE58BCFE767">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F11F41EB193FDA81715A454E70F9AA44">Larico, Jackie Farfán</mods:namePart>
<mods:nameIdentifier id="BC6935EAF18E547EBEA4BF37CC462DE3" type="ORCID">0000-0003-3930-6480</mods:nameIdentifier>
<mods:affiliation id="FCDF4F0F2FE5523F21F2BACFF40DD252">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
<mods:nameIdentifier id="CB6D7B7431000E865AA0756EBCB50BDB" type="email">cerdenajoseal@yahoo.es</mods:nameIdentifier>
</mods:name>
<mods:name id="3540A7C59AA269BFB5D35653E425A8DB" type="personal">
<mods:role id="4FBDBA65FDE7643710110E085781F371">
<mods:roleTerm id="9264D786CFE8271CD362CD9AD7A5B2EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90C20078CADCAD4C62B74A06D61FDD2F">Brudecka, Jadwiga Lorenc-</mods:namePart>
<mods:affiliation id="A5A466B133006D460E71AA722191DC7E">Nature Education Centre, Jagiellonian University, Gronostajowa 5, 30 - 387 Kraków, Poland.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="907245E9FE8CBB4A786D49C3056AECB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27789F0727D79D7CECFCA0A6190E5A60">Prieto, Carlos</mods:namePart>
<mods:affiliation id="8F8DFF18FDEB5FDBB120A41BE12D7CE8">Corporación Universitaria Autónoma del Cauca, Popayán, Colombia.</mods:affiliation>
</mods:name>
<mods:name id="B83900327F4C653436312C422B31F7BA" type="personal">
<mods:role id="19B3756F81B6051A5E6E5BD801C38831">
<mods:roleTerm id="CED90C13D42B5C887DA851DCD6045C9B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA48BE3DA33B8ED2C099AEF0133AFEE1">Pyrcz, Tomasz W.</mods:namePart>
<mods:affiliation id="C86466308F60F2A25F40EBAF7081F9D1">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland.</mods:affiliation>
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<mods:title id="7CA2F6C20C4D40E5A07D8E09D09DF303">Zootaxa</mods:title>
</mods:titleInfo>
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<mods:date id="FD989DD7EFB63EFCC63B5CFD4A446830">2021</mods:date>
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<mods:number id="F671BC1C5946D9D8ED2B47E06993A727">2021-04-16</mods:number>
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<mods:number id="93B557C8511570A11A150D65CF5CC782">4963</mods:number>
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<mods:classification id="B63BA92B48CF337C5968CBB264534158">journal article</mods:classification>
<mods:identifier id="6549C074CBDA23BF81CC5C32D4CD4FED" type="CLB-Dataset">7214</mods:identifier>
<mods:identifier id="581F0A2253A09ACF93918449AFE00135" type="DOI">10.11646/zootaxa.4963.1.2</mods:identifier>
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<treatment id="D5008F50FFE6B31CFF76FBC99221FE1C" ID-DOI="http://doi.org/10.5281/zenodo.4696422" ID-GBIF-Taxon="181531945" ID-Zenodo-Dep="4696422" LSID="urn:lsid:plazi:treatment:D5008F50FFE6B31CFF76FBC99221FE1C" httpUri="http://treatment.plazi.org/id/D5008F50FFE6B31CFF76FBC99221FE1C" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26">
<subSubSection id="15B36DCDFFE6B303FF76FBC99462FB47" box="[151,587,1135,1162]" pageId="15" pageNumber="26" type="nomenclature">
<paragraph id="5D163E46FFE6B303FF76FBC99462FB47" blockId="15.[151,587,1135,1198]" box="[151,587,1135,1162]" pageId="15" pageNumber="26">
<heading id="065E892AFFE6B303FF76FBC99462FB47" bold="true" box="[151,587,1135,1162]" fontSize="11" level="1" pageId="15" pageNumber="26" reason="1">
<taxonomicName id="9AA945C5FFE6B303FF76FBC99462FB47" authority="Balint, 2003" authorityName="Balint" authorityYear="2003" box="[151,587,1135,1162]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="johnbanksi">
<emphasis id="6FDDE254FFE6B303FF76FBC99462FB47" bold="true" box="[151,587,1135,1162]" pageId="15" pageNumber="26">
<emphasis id="6FDDE254FFE6B303FF76FBC99784FB47" bold="true" box="[151,429,1135,1162]" italics="true" pageId="15" pageNumber="26">Johnsonita johnbanksi</emphasis>
<bibRefCitation id="393843B7FFE6B303FE55FBC99462FB47" author="Balint" box="[436,587,1135,1162]" firstAuthor="Balint" journalOrPublisher="Folia entomologica hungarica" pageId="15" pageNumber="26" pagination="243 - 250" part="64" refId="ref21464" refString="Balint, Zs. (2003) Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae). Folia entomologica hungarica, 64, 243 - 250." title="Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae)" type="journal article" year="2003">Bálint, 2003</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="15B36DCDFFE6B303FF76FB329405FB63" box="[151,556,1172,1198]" pageId="15" pageNumber="26" type="description">
<paragraph id="5D163E46FFE6B303FF76FB329405FB63" blockId="15.[151,587,1135,1198]" box="[151,556,1172,1198]" pageId="15" pageNumber="26">
(
<figureCitation id="C59222C3FFE6B303FF7EFB3296C1FB63" box="[159,232,1172,1198]" captionStart="FIGURE 9" captionStartId="16.[151,250,1817,1841]" captionTargetBox="[151,1428,499,1777]" captionTargetId="figure-158@16.[151,1436,489,1794]" captionTargetPageId="16" captionText="FIGURE 9. Johnsonita johnbanksi Bálint, 2003 imagines (left column: recto, right column: verso). A–B: male (Peru: Amazonas), C–D: paratype female (Peru: Amazonas) (CEP–MZUJ specimens) (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711269" httpUri="https://zenodo.org/record/4711269/files/figure.png" pageId="15" pageNumber="26">Figs 9</figureCitation>
,
<figureCitation id="C59222C3FFE6B303FF15FB32976EFB63" box="[244,327,1172,1198]" captionStart="FIGURE 18" captionStartId="28.[151,249,1943,1967]" captionTargetBox="[227,1333,184,1900]" captionTargetId="figure-23@28.[175,1413,181,1920]" captionTargetPageId="28" captionText="FIGURE 18. Johnsonita genitalia armatures, entire male organs in lateral (left column) and ventral (right column) views. A–B: J. ianusca, Ecuador, Azuay, Cuenca; C–D: J. johnbanksi, Peru, Molinopampa; E–F: J. johnsoni, Ecuador, Riobamba-Macas via Atillo; G–H: J. pardoa, Venezuela, Táchira, P. N. Tama (CEP–MZUJ specimens) (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711293" httpUri="https://zenodo.org/record/4711293/files/figure.png" pageId="15" pageNumber="26">18C–D</figureCitation>
,
<figureCitation id="C59222C3FFE6B303FEB2FB3297B5FB63" box="[339,412,1172,1198]" captionStart="FIGURE 21" captionStartId="31.[151,250,1909,1933]" captionTargetBox="[188,1399,181,1885]" captionTargetId="figure-18@31.[188,1399,181,1885]" captionTargetPageId="31" captionText="FIGURE 21. Entire Johnsonita female organs in lateral (left column), ductus in lateral (central column) and ventral (right column) views. A–C: J. chlamydem, Peru, Amazonas, Rodriguez de Mendoza; D–F: J. iacinta, Peru, Pasco-Oxapampa, La Antena; G–I: J. johnbanksi, Peru, Mendoza–Llanuayco; J–L: J. johnsoni, Ecuador, Azuay, Gualaceo-Limón road (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711299" httpUri="https://zenodo.org/record/4711299/files/figure.png" pageId="15" pageNumber="26">21G–I</figureCitation>
,
<figureCitation id="C59222C3FFE6B303FE49FB3297D5FB63" box="[424,508,1172,1198]" captionStart="FIGURE 24" captionStartId="34.[151,250,1502,1526]" captionTargetBox="[400,1168,186,1466]" captionTargetId="figure-23@34.[396,1192,181,1478]" captionTargetPageId="34" captionText="FIGURE 24. Johnsonita mail traits in dorsal hindwing (left column) and ventral forewing subcubital patches (right column) indicated by arrow. A–B: J. oxalida, C–D: J. johnbanksi, E–F: J. subcunicula, G: J. auda." figureDoi="http://doi.org/10.5281/zenodo.4711307" httpUri="https://zenodo.org/record/4711307/files/figure.png" pageId="15" pageNumber="26">24C–D</figureCitation>
,
<figureCitation id="C59222C3FFE6B303FDE6FB32940DFB63" box="[519,548,1172,1198]" captionStart="FIGURE 32" captionStartId="41.[151,250,1937,1961]" captionTargetBox="[185,1436,207,1913]" captionTargetId="figure-18@41.[151,1436,181,1913]" captionTargetPageId="41" captionText="FIGURE 32. Johnsonita distributions based on museum material examinded. J. johnbanksi = yellow triangle, J. johnsoni = green square, J. pardoa = red circle." figureDoi="http://doi.org/10.5281/zenodo.4711329" httpUri="https://zenodo.org/record/4711329/files/figure.png" pageId="15" pageNumber="26">32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="15B36DCDFFE6B303FF76FB789460FB3B" box="[151,585,1244,1270]" pageId="15" pageNumber="26" type="reference_group">
<paragraph id="5D163E46FFE6B303FF76FB789460FB3B" blockId="15.[151,585,1244,1270]" box="[151,585,1244,1270]" pageId="15" pageNumber="26">
<treatmentCitationGroup id="7DB91968FFE6B303FF76FB789460FB3B" box="[151,585,1244,1270]" pageId="15" pageNumber="26">
<taxonomicName id="9AA945C5FFE6B303FF76FB789755FB3B" authorityName="Balint" authorityYear="2003" box="[151,380,1246,1270]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="johnbanksi">
<emphasis id="6FDDE254FFE6B303FF76FB789755FB3B" box="[151,380,1246,1270]" italics="true" pageId="15" pageNumber="26">Johnsonita johnbanksi</emphasis>
</taxonomicName>
–
<treatmentCitation id="DC081857FFE6B303FE76FB78946CFB3B" author="Balint" box="[407,581,1246,1270]" page="243" pageId="15" pageNumber="26" year="2003">
<bibRefCitation id="393843B7FFE6B303FE76FB78946CFB3B" author="Balint" box="[407,581,1246,1270]" firstAuthor="Balint" journalOrPublisher="Folia entomologica hungarica" page="243" pageId="15" pageNumber="26" pagination="243 - 250" part="64" refId="ref21464" refString="Balint, Zs. (2003) Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae). Folia entomologica hungarica, 64, 243 - 250." title="Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae)" type="journal article" year="2003">Bálint 2003: 243</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="15B36DCDFFE6B31CFF76FA839221FE1C" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26" type="materials_examined">
<paragraph id="5D163E46FFE6B303FF76FA8396E6FA3F" blockId="15.[151,1437,1316,2026]" pageId="15" pageNumber="26">
<emphasis id="6FDDE254FFE6B303FF76FA839712FAF3" box="[151,315,1316,1342]" italics="true" pageId="15" pageNumber="26">Type material.</emphasis>
<materialsCitation id="EDC1341BFFE6B303FEA2FA8296E2FA3F" ID-GBIF-Occurrence="3083794357" collectingDate="2001-12-08" collectionCode="MUSM, CEP-MZUJ, HNHM, NMW" collectorName="J. Lorenc" country="Peru" elevation="3100" location="Described" pageId="15" pageNumber="26" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<location id="5876689DFFE6B303FEA2FA82979CFAF3" LSID="urn:lsid:plazi:treatment:D5008F50FFE6B31CFF76FBC99221FE1C:5876689DFFE6B303FEA2FA82979CFAF3" box="[323,437,1316,1342]" country="Peru" name="Described" pageId="15" pageNumber="26">Described</location>
on the basis of the
<collectionCode id="3BB8A683FFE6B303FD7BFA8294D9FAF3" box="[666,752,1316,1342]" pageId="15" pageNumber="26">MUSM</collectionCode>
<typeStatus id="821280E4FFE6B303FD18FA829572FAF3" box="[761,859,1316,1342]" pageId="15" pageNumber="26" type="holotype">holotype</typeStatus>
male from Peru, set dorsally, in perfect condition, labelled as: “
<collectingCountry id="25BE7ED6FFE6B303FECBFAEE975CFAAF" box="[298,373,1352,1378]" name="Peru" pageId="15" pageNumber="26">PERU</collectingCountry>
[//] Amazonas, Chachapoyas [//] Molinopampa-Granada [//]
<date id="29171886FFE6B303FBCEFAEE9299FAAF" box="[1071,1200,1352,1378]" pageId="15" pageNumber="26" value="2001-12-08">
<collectingDate id="3953E16EFFE6B303FBCEFAEE9299FAAF" box="[1071,1200,1352,1378]" pageId="15" pageNumber="26" value="2001-12-08">12.08.2001</collectingDate>
</date>
,
<quantity id="9A5193A3FFE6B303FB5CFAEE933DFAAF" box="[1213,1300,1352,1378]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="26" unit="m" value="3100.0">
<elevation id="D684D975FFE6B303FB5CFAEE933DFAAF" box="[1213,1300,1352,1378]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="26" unit="m" value="3100.0">3100 m</elevation>
</quantity>
[//]
<collectorName id="F05C5B90FFE6B303FAA6FAEE96D7FA4B" pageId="15" pageNumber="26">Leg. B. Calderon</collectorName>
[//]
<collectorName id="F05C5B90FFE6B303FECFFACA97C3FA4B" box="[302,490,1388,1414]" pageId="15" pageNumber="26">Coll. M. Bollino</collectorName>
” (white label, printed).
<typeStatus id="821280E4FFE6B303FD14FACA954BFA4B" box="[757,866,1388,1414]" pageId="15" pageNumber="26" type="paratype">Paratypes</typeStatus>
(n=6) all from
<collectingCountry id="25BE7ED6FFE6B303FBEEFACA9272FA4B" box="[1039,1115,1388,1414]" name="Peru" pageId="15" pageNumber="26">PERU</collectingCountry>
: three
<collectionCode id="3BB8A683FFE6B303FB45FACA9303FA4B" box="[1188,1322,1388,1414]" pageId="15" pageNumber="26">CEP-MZUJ</collectionCode>
<typeStatus id="821280E4FFE6B303FAD0FACA93B5FA4B" box="[1329,1436,1388,1414]" pageId="15" pageNumber="26" type="paratype">paratypes</typeStatus>
(
<specimenCount id="4BAFF5CFFFE6B303FF7EFA36973BFA67" box="[159,274,1424,1450]" pageId="15" pageNumber="26" type="male">two males</specimenCount>
and “
<typeStatus id="821280E4FFE6B303FEB3FA369798FA67" box="[338,433,1424,1450]" pageId="15" pageNumber="26" type="allotype">allotype</typeStatus>
” female, dissected: male: gen. prep. 382/
<collectorName id="F05C5B90FFE6B303FC9EFA3695CEFA67" box="[895,999,1424,1450]" pageId="15" pageNumber="26">J. Lorenc</collectorName>
; female: gen. prep. 373/
<collectorName id="F05C5B90FFE6B303FB16FA369349FA67" box="[1271,1376,1424,1450]" pageId="15" pageNumber="26">J. Lorenc</collectorName>
), two
<collectionCode id="3BB8A683FFE6B303FF76FA1296C4FA03" box="[151,237,1460,1486]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="15" pageNumber="26">HNHM</collectionCode>
<typeStatus id="821280E4FFE6B303FF14FA129749FA03" box="[245,352,1460,1486]" pageId="15" pageNumber="26" type="paratype">paratypes</typeStatus>
– all from the type locality; and one
<collectionCode id="3BB8A683FFE6B303FCEFFA12957CFA03" box="[782,853,1460,1486]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="26">NMW</collectionCode>
<typeStatus id="821280E4FFE6B303FCBFFA129597FA03" box="[862,958,1460,1486]" pageId="15" pageNumber="26" type="paratype">paratype</typeStatus>
female from Pedro Ruiz Gallo; all examined
</materialsCitation>
.
</paragraph>
<paragraph id="5D163E46FFE6B303FF26FA5A941CF96A" blockId="15.[151,1437,1316,2026]" pageId="15" pageNumber="26">
<emphasis id="6FDDE254FFE6B303FF26FA5A97B6F9DB" box="[199,415,1532,1558]" italics="true" pageId="15" pageNumber="26">Material examined</emphasis>
(n=6),
<materialsCitation id="EDC1341BFFE6B303FE1AFA5A941CF96A" ID-GBIF-Occurrence="3083794320" collectingDate="1998-06-29" collectingDateMax="2006-07-23" collectingDateMin="1998-06-29" collectionCode="T, CEP, MZUJ, CB, HNHM" collectorName=" O. Duviols" country="Peru" county="Amazonas" elevation="2150" location="Rio Nieva" municipality="Molinopampa" pageId="15" pageNumber="26" specimenCount="2" specimenCount-male="2" stateProvince="San Martin">
all from
<collectingCountry id="25BE7ED6FFE6B303FD82FA5A9487F9DB" box="[611,686,1532,1558]" name="Peru" pageId="15" pageNumber="26">PERU</collectingCountry>
.
<collectingCounty id="B47746CAFFE6B303FD5AFA5A951DF9DB" box="[699,820,1532,1558]" pageId="15" pageNumber="26">Amazonas</collectingCounty>
,
<collectingMunicipality id="BD72A43CFFE6B303FCA2FA5A95CAF9DB" box="[835,995,1532,1558]" pageId="15" pageNumber="26">Molinopampa</collectingMunicipality>
,
<date id="29171886FFE6B303FC13FA5A925DF9DB" box="[1010,1140,1532,1558]" pageId="15" pageNumber="26" value="1998-06-29">
<collectingDate id="3953E16EFFE6B303FC13FA5A925DF9DB" box="[1010,1140,1532,1558]" pageId="15" pageNumber="26" value="1998-06-29">29.VI.1998</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE6B303FB62FA5A9338F9DB" box="[1155,1297,1532,1558]" pageId="15" pageNumber="26">J. Wojtusiak</collectorName>
&amp;
<collectionCode id="3BB8A683FFE6B303FADAFA5A9364F9DB" box="[1339,1357,1532,1558]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="26">T</collectionCode>
. Pyrcz (
<collectionCode id="3BB8A683FFE6B303FF7EF98696FDF9F7" box="[159,212,1568,1594]" pageId="15" pageNumber="26">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE6B303FF02F9869719F9F7" box="[227,304,1568,1594]" pageId="15" pageNumber="26">MZUJ</collectionCode>
: male); Amazonas, Pomacocha,
<quantity id="9A5193A3FFE6B303FD44F98694D6F9F7" box="[677,767,1568,1595]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="15" pageNumber="26" unit="m" value="2150.0">
<elevation id="D684D975FFE6B303FD44F98694D6F9F7" box="[677,767,1568,1595]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="15" pageNumber="26" unit="m" value="2150.0">2150 m</elevation>
</quantity>
,
<date id="29171886FFE6B303FCEFF98695B9F9F7" box="[782,912,1568,1594]" pageId="15" pageNumber="26" value="2000-06-13">
<collectingDate id="3953E16EFFE6B303FCEFF98695B9F9F7" box="[782,912,1568,1594]" pageId="15" pageNumber="26" value="2000-06-13">13.VI.2000</collectingDate>
</date>
, Calderon (
<collectionCode id="3BB8A683FFE6B303FBF9F986926BF9F7" box="[1048,1090,1568,1594]" country="Australia" name="The CB Rhizobium Collection" pageId="15" pageNumber="26">CB</collectionCode>
:
<specimenCount id="4BAFF5CFFFE6B303FBAEF9869282F9F7" box="[1103,1195,1568,1594]" pageId="15" pageNumber="26" type="male">2 males</specimenCount>
, female); Amazonas, Alto
<location id="5876689DFFE6B303FF32F9E29762F993" LSID="urn:lsid:plazi:treatment:D5008F50FFE6B31CFF76FBC99221FE1C:5876689DFFE6B303FF32F9E29762F993" box="[211,331,1604,1630]" country="Peru" county="Amazonas" municipality="Molinopampa" name="Rio Nieva" pageId="15" pageNumber="26" stateProvince="San Martin">Rio Nieva</location>
,
<quantity id="9A5193A3FFE6B303FEB8F9E297D0F993" box="[345,505,1604,1631]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="15" pageNumber="26" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">
<elevation id="D684D975FFE6B303FEB8F9E297D0F993" box="[345,505,1604,1631]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="15" pageNumber="26" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">2200–2500 m</elevation>
</quantity>
,
<date id="29171886FFE6B303FDE6F9E2944EF993" box="[519,615,1604,1630]" pageId="15" pageNumber="26" value="2002-06">
<collectingDate id="3953E16EFFE6B303FDE6F9E2944EF993" box="[519,615,1604,1630]" pageId="15" pageNumber="26" value="2002-06">VI.2002</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE6B303FD95F9E294D6F993" box="[628,767,1604,1630]" pageId="15" pageNumber="26">B. Calderon</collectorName>
(
<collectionCode id="3BB8A683FFE6B303FCF1F9E2956CF993" box="[784,837,1604,1630]" pageId="15" pageNumber="26">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE6B303FCB5F9E29588F993" box="[852,929,1604,1630]" pageId="15" pageNumber="26">MZUJ</collectionCode>
: female); Amazonas, Chachapoyas, Molinopampa,
<quantity id="9A5193A3FFE6B303FF0EF9CE976CF94F" box="[239,325,1640,1666]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="26" unit="m" value="2300.0">
<elevation id="D684D975FFE6B303FF0EF9CE976CF94F" box="[239,325,1640,1666]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="26" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="29171886FFE6B303FEB1F9CE9786F94F" box="[336,431,1640,1666]" pageId="15" pageNumber="26" value="2002-09">
<collectingDate id="3953E16EFFE6B303FEB1F9CE9786F94F" box="[336,431,1640,1666]" pageId="15" pageNumber="26" value="2002-09">IX.2002</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE6B303FE5BF9CE9469F94F" box="[442,576,1640,1666]" pageId="15" pageNumber="26">B. Calderón</collectorName>
(
<collectionCode id="3BB8A683FFE6B303FDAFF9CE9480F94F" box="[590,681,1640,1666]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="15" pageNumber="26">HNHM</collectionCode>
: male);
<collectingRegion id="9F6DF0A4FFE6B303FCE0F9CE95A9F94F" box="[769,896,1640,1666]" country="Peru" name="San Martin" pageId="15" pageNumber="26">San Martín</collectingRegion>
, Jorge Chavez,
<location id="5876689DFFE6B303FBCCF9CE92D8F94F" LSID="urn:lsid:plazi:treatment:D5008F50FFE6B31CFF76FBC99221FE1C:5876689DFFE6B303FBCCF9CE92D8F94F" box="[1069,1265,1640,1666]" country="Peru" county="Amazonas" municipality="Molinopampa" name="Serranoyacu area" pageId="15" pageNumber="26" stateProvince="San Martin">Serranoyacu area</location>
,
<quantity id="9A5193A3FFE6B303FB1AF9CE93BEF94F" box="[1275,1431,1640,1667]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="15" pageNumber="26" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">
<elevation id="D684D975FFE6B303FB1AF9CE93BEF94F" box="[1275,1431,1640,1667]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="15" pageNumber="26" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500–1600 m</elevation>
</quantity>
,
<date id="29171886FFE6B303FF76F92A9703F96B" box="[151,298,1676,1702]" pageId="15" pageNumber="26" value="2006-07-23">
<collectingDate id="3953E16EFFE6B303FF76F92A9703F96B" box="[151,298,1676,1702]" pageId="15" pageNumber="26" value="2006-07-23">23.VII. 2006</collectingDate>
</date>
,
<collectorName id="F05C5B90FFE6B303FED4F92A9799F96B" box="[309,432,1676,1702]" pageId="15" pageNumber="26">O. Duviols</collectorName>
(
<collectionCode id="3BB8A683FFE6B303FE21F92A97C3F96B" box="[448,490,1676,1702]" country="Australia" name="The CB Rhizobium Collection" pageId="15" pageNumber="26">CB</collectionCode>
: male)
</materialsCitation>
</paragraph>
<paragraph id="5D163E46FFE6B31CFF26F9169364FF18" blockId="15.[151,1437,1316,2026]" lastBlockId="16.[151,1437,151,465]" lastPageId="16" lastPageNumber="27" pageId="15" pageNumber="26">
<emphasis id="6FDDE254FFE6B303FF26F9169743F907" box="[199,362,1712,1738]" italics="true" pageId="15" pageNumber="26">Redescription.</emphasis>
Wings. FW costa length:
<quantity id="9A5193A3FFE6B303FD6CF916952FF907" box="[653,774,1712,1739]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="15" pageNumber="26" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
(n=5,
<collectionCode id="3BB8A683FFE6B303FCB1F91695AAF907" box="[848,899,1712,1738]" pageId="15" pageNumber="26">CEP</collectionCode>
–
<collectionCode id="3BB8A683FFE6B303FC70F91695F3F907" box="[913,986,1712,1738]" pageId="15" pageNumber="26">MZUJ</collectionCode>
material), apical area of costa and outer margin slightly convex, resulting in a somewhat rounded shape; dorsal wing surface colouration pale azure, black border inner margin reaching the erection of vein M3; HW dorsal surface discal area structurally coloured, and with patch of hairy androconial scales, ventral FW submedial and medial pattern running parallel, HW medial pattern composed of light ruptive dashes supplemented distally with dark spots, postmedial line complete and running from costa to tornus as a faint band; ventral HW submedial area at costa with discal area colour, postmedial pattern composed by conspicuous brown arrow-head markings, inner margin medial area with brown scaling. Androconia present in FW ventral surface postbasal area below the Cubitus and in the HW dorsal wing surface on the discalis where there is a fur and amongst them there are specialized scales. Genitalia. Male valva with rounded costal margin flip and extremely pointed and long valval terminus. Female ductus thin with mostly parallel edges but at 2/3 length divergent, no lower plate, upper plate symmetric and large with thinly sclerotized frame with a single spine.
</paragraph>
<paragraph id="5D163E46FFF9B31CFF26FF7995D4FED0" blockId="16.[151,1437,151,465]" pageId="16" pageNumber="27">
<emphasis id="6FDDE254FFF9B31CFF26FF799769FF35" box="[199,320,223,248]" italics="true" pageId="16" pageNumber="27">Diagnosis.</emphasis>
This is another species easy to distinguish from congeners on the basis of the dorsal HW androconia and the ventral HW surface with such contrasted pattern, unique in the genus.
</paragraph>
<paragraph id="5D163E46FFF9B31CFF26FE81954DFEA8" blockId="16.[151,1437,151,465]" pageId="16" pageNumber="27">
<emphasis id="6FDDE254FFF9B31CFF26FE819771FE8C" box="[199,344,295,321]" italics="true" pageId="16" pageNumber="27">Distribution.</emphasis>
Geographical:
<collectingCountry id="25BE7ED6FFF9B31CFDE3FE819463FE8C" box="[514,586,295,321]" name="Peru" pageId="16" pageNumber="27">PERU</collectingCountry>
(Amazonas) (
<figureCitation id="C59222C3FFF9B31CFD06FE819512FE8C" box="[743,827,295,321]" captionStart="FIGURE 32" captionStartId="41.[151,250,1937,1961]" captionTargetBox="[185,1436,207,1913]" captionTargetId="figure-18@41.[151,1436,181,1913]" captionTargetPageId="41" captionText="FIGURE 32. Johnsonita distributions based on museum material examinded. J. johnbanksi = yellow triangle, J. johnsoni = green square, J. pardoa = red circle." figureDoi="http://doi.org/10.5281/zenodo.4711329" httpUri="https://zenodo.org/record/4711329/files/figure.png" pageId="16" pageNumber="27">Fig. 32</figureCitation>
); spatial: recorded between
<quantity id="9A5193A3FFF9B31CFB90FE809325FE8C" box="[1137,1292,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="3.1" metricValueMin="1.5" pageId="16" pageNumber="27" unit="m" value="2300.0" valueMax="3100.0" valueMin="1500.0">1500–3100 m</quantity>
a.s.l; temporal: recorded in January–February, June and August–September.
</paragraph>
<paragraph id="5D163E46FFF9B31CFF26FEC99221FE1C" blockId="16.[151,1437,151,465]" pageId="16" pageNumber="27">
<emphasis id="6FDDE254FFF9B31CFF26FEC9972DFE45" box="[199,260,367,392]" italics="true" pageId="16" pageNumber="27">Note.</emphasis>
Dorsal HW surface discal area in female specimens is covered by long hair-like scales but amongst them there are no specialized scales. The species was dedicated to John Banks (London,
<collectingCountry id="25BE7ED6FFF9B31CFBDCFE35924EFE60" box="[1085,1127,403,429]" name="United Kingdom" pageId="16" pageNumber="27">UK</collectingCountry>
) who produced many films on butterflies, including South American fauna released in 1996 (
<bibRefCitation id="393843B7FFF9B31CFC93FE1195D5FE1C" author="Balint" box="[882,1020,439,465]" firstAuthor="Balint" journalOrPublisher="Boletin Cientifico Centro De Museos, Museo de Historia Natural" pageId="16" pageNumber="27" pagination="276" part="13" refId="ref21601" refString="Balint, Zs. (2009) Revisiones Bibliograficas: &quot; O Que Passa? What's Going On Here? &quot; - An Introduction to the Butterflies of the Amazon Forest. Boletin Cientifico Centro De Museos, Museo de Historia Natural, 13 (2), 276." title="Revisiones Bibliograficas: &quot; O Que Passa? What's Going On Here? &quot; - An Introduction to the Butterflies of the Amazon Forest" type="journal article" year="2009">Bálint 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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