475 lines
46 KiB
XML
475 lines
46 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.452.8152" ID-PMC="PMC4258635" ID-Pensoft-Pub="1313-2970-452-107" ID-Pensoft-UUID="FFDEFFD61800FFEEBA2CFFA1014FFF8A" ID-PubMed="25493056" ID-Zenodo-Dep="578587" ID-ZooBank="F7F5E4C32D764ACAA4C35B8DE5D0F90C" ModsDocID="1313-2970-452-107" checkinTime="1451245146091" checkinUser="pensoft" docAuthor="Pekarsky, Oleg" docDate="2014" docId="A7F00213DD9B54A2A159C87CA0A2E7AF" docLanguage="en" docName="ZooKeys 452: 107-129" docOrigin="ZooKeys 452" docPubDate="2014-11-05" docSource="http://dx.doi.org/10.3897/zookeys.452.8152" docTitle="Lygephila lubrosa subsp. lubrosa lubrosa (Staudinger 1901" docType="treatment" docVersion="3" id="FFDEFFD61800FFEEBA2CFFA1014FFF8A" lastPageNumber="114" masterDocId="FFDEFFD61800FFEEBA2CFFA1014FFF8A" masterDocTitle="Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae)" masterLastPageNumber="129" masterPageNumber="107" pageNumber="110" updateTime="1668159553656" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pekarsky, Oleg</mods:namePart>
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<mods:affiliation>H- 1068, Budapest, Felsoerdosor u. 16 - 18, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">opbp@t-online.hu</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-11-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>452</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>107</mods:start>
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<mods:end>129</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.452.8152</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.452.8152</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-452-107</mods:identifier>
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<mods:identifier type="ZooBank">F7F5E4C32D764ACAA4C35B8DE5D0F90C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFDEFFD61800FFEEBA2CFFA1014FFF8A</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578587</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182234915" LSID="urn:lsid:plazi:treatment:A7F00213DD9B54A2A159C87CA0A2E7AF" httpUri="http://treatment.plazi.org/id/A7F00213DD9B54A2A159C87CA0A2E7AF" lastPageId="7" lastPageNumber="114" pageId="3" pageNumber="110">
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<subSubSection pageId="3" pageNumber="110" type="nomenclature">
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<paragraph pageId="3" pageNumber="110">
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<taxonomicName LSID="A7F00213-DD9B-54A2-A159-C87CA0A2E7AF" authority="lubrosa (Staudinger, 1901)" authorityName="lubrosa (Staudinger" authorityYear="1901" baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa subsp. lubrosa" order="Lepidoptera" pageId="3" pageNumber="110" phylum="Arthropoda" rank="subSpecies" species="lubrosa" subSpecies="lubrosa">Lygephila lubrosa lubrosa (Staudinger, 1901)</taxonomicName>
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<figureCitation captionStart="Figures 11–20" captionStartId="F2" captionText="Figures 11 - 20. Adults. 11 - 16 Lygephila kazachkaratavika 11 neotype, ♂, Balamurum 12 ♀, Kazakhstan, Baigacum 13 ♂, Kazakhstan, Taldy-Kurgan reg. 14 ♀, Kazakstan, Baigacum 15 ♂, Kyrgyzstan, Issyk Kul 16 ♀, Kyrgyzstan, Issyk Kul 17, 18 Lygephila lubrosa lubrosa 17 neotype, ♂, Kazakhstan, Ili river 18 ♀, Kazakhstan, Baigacum 19, 20 Lygephila lubrosa orbonaria 19 neotype, ♂, Tajikistan, Pianj river 20 ♂, Tajikistan, Pianj river." pageId="3" pageNumber="110">Figs 17</figureCitation>
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<figureCitation captionStart="Figures 11–20" captionStartId="F2" captionText="Figures 11 - 20. Adults. 11 - 16 Lygephila kazachkaratavika 11 neotype, ♂, Balamurum 12 ♀, Kazakhstan, Baigacum 13 ♂, Kazakhstan, Taldy-Kurgan reg. 14 ♀, Kazakstan, Baigacum 15 ♂, Kyrgyzstan, Issyk Kul 16 ♀, Kyrgyzstan, Issyk Kul 17, 18 Lygephila lubrosa lubrosa 17 neotype, ♂, Kazakhstan, Ili river 18 ♀, Kazakhstan, Baigacum 19, 20 Lygephila lubrosa orbonaria 19 neotype, ♂, Tajikistan, Pianj river 20 ♂, Tajikistan, Pianj river." pageId="3" pageNumber="110">, 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="111" type="reference_group">
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<paragraph pageId="4" pageNumber="111">
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<taxonomicName class="Insecta" family="Erebidae" genus="Toxocampa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Toxocampa lubrica subsp. var. var. lubrosa" order="Lepidoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="variety" species="lubrica" subSpecies="var." variety="lubrosa">
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<pageBreakToken pageId="4" pageNumber="111" start="start">Toxocampa</pageBreakToken>
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lubrica var. lubrosa
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</taxonomicName>
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Staudinger, 1901, Catalog der Lepidopteren des Palaearctischen Faunengebietes. I: 252. (TL: [Kazakhstan], Ili, [Kyrgyzstan, Issyk Kul], "Iss. K.")
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="111" type="type material examined">
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<paragraph pageId="4" pageNumber="111">Type material examined.</paragraph>
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<paragraph pageId="4" pageNumber="111">
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<emphasis bold="true" pageId="4" pageNumber="111">Neotype</emphasis>
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(here designated) male, Kazakhstan, Ili river valley near bridge 23,4 km asimut 222 from Koktal, 600 m,
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<geoCoordinate degrees="43" direction="north" minutes="58" orientation="latitude" precision="15" seconds="004" value="43.967777">N43°58'004"</geoCoordinate>
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, E79°35'905", 04.07.2010, leg. S.K. Korb, slide No. OP2082m (coll. O. Pekarsky, deposited in HNHM Budapest).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="111" type="additional material examined">
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<paragraph pageId="4" pageNumber="111">Additional material examined.</paragraph>
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<paragraph pageId="4" pageNumber="111">
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1 ♂, with same data as neotype; 1 ♂ & 1 ♀, Kazakhstan, Ili river valley near Koktal, 506 m,
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<geoCoordinate degrees="43" direction="north" minutes="57" orientation="latitude" precision="1" seconds="57.50" value="43.965973">N43°57'57.50"</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="east" minutes="36" orientation="longitude" precision="1" seconds="1.06" value="79.600296">E79°36'1.06"</geoCoordinate>
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, 03.07.2010, leg. S.K. Korb, slide No. OP2489f (coll. O. Pekarsky); 1 ♀, [Kazakhstan], Syr-Daria, Baigacum, Koshantschikoff, 23.6.1913, 4/7, ex. coll.
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<normalizedToken originalValue="Püngeler">Puengeler</normalizedToken>
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, slide No. OP1979f (coll. MNHU); 1 ♂, [Kazakhstan], Aj-Darle, Syr-Darja, 25.V.1909, leg. Koshantshikoff, slide No. 0325Matov (coll. ZISP).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="113" pageId="4" pageNumber="111" type="taxonomy">
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<paragraph pageId="4" pageNumber="111">Taxonomy.</paragraph>
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<paragraph lastPageId="5" lastPageNumber="112" pageId="4" pageNumber="111">
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Described by Staudinger in 1901 as a variation of
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<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="lubrica">
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<emphasis italics="true" pageId="5" pageNumber="112">
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<pageBreakToken pageId="5" pageNumber="112" start="start">Lygephila</pageBreakToken>
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lubrica
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</emphasis>
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</taxonomicName>
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; with the type locality mentioned as [Kazakhstan], Ili [river] and [Kyrgyzstan], Issyk Kul [lake]. The original description stated that the forewings are pale grey (
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<normalizedToken originalValue="“cinereo-griseis”">"cinereo-griseis"</normalizedToken>
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) without dark outer part, and that the hindwings are ochreous with broad marginal fascia. This description corresponds exactly with the external appearance of the moths from Ili river in Kazakhstan, therefore the neotype is designated from this area. Moths from Issyk Kul show, however, marked differences in habitus, especially the brown coloration of most parts of the forewings. These two taxa are different in genital structures of both sexes, which are discussed in detail under
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<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="kazachkaratavika">
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<emphasis italics="true" pageId="5" pageNumber="112">Lygephila kazachkaratavika</emphasis>
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</taxonomicName>
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. Starting from the 1980's, Stshetkin YuL treated
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="lubrosa">
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<emphasis italics="true" pageId="5" pageNumber="112">Lygephila lubrosa</emphasis>
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</taxonomicName>
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in his publications as a distinct species (
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<bibRefCitation author="Stshetkin, YuL" editor="Abdusalamov, IA" journalOrPublisher="Donish, Dushanbe" pageId="12" pageNumber="119" pagination="31 - 44" refId="B14" refString="Stshetkin, YuL, Degtarova, VI, Stshetkin, YuYu, 1988. Lepidoptera. In: Abdusalamov, IA, Ed., Red Book of Tajik SSR. Donish, Dushanbe: 31 - 44" title="Lepidoptera." volumeTitle="Red Book of Tajik SSR." year="1988">Stshetkin et al. 1988</bibRefCitation>
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,
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<bibRefCitation author="Sviridov, AV" journalOrPublisher="Kiev" pageId="12" pageNumber="119" pagination="97 - 100" refId="B15" refString="Sviridov, AV, 1990. K tipovym materialam sovok, opisannyh F. Brikom iz Korei. In: Novosti faunistiki i sistematiki. Kiev: 97 - 100" title="K tipovym materialam sovok, opisannyh F. Brikom iz Korei." volumeTitle="Novosti faunistiki i sistematiki." year="1990">Stshetkin 1991</bibRefCitation>
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). The explanation of this act was given only in 1994 [1997] by Stshetkin YuL & Stshetkin YuYu. Their argumentation was based only on the original description of
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="lubrosa">
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<emphasis italics="true" pageId="5" pageNumber="112">Lygephila lubrosa</emphasis>
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</taxonomicName>
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, but neither the type material nor the genitalia of the syntypes were studied. Unfortunately, the authors evidently failed in their taxonomic interpretation of the species complex. They were correct to suppose
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="lubrosa">
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<emphasis italics="true" pageId="5" pageNumber="112">Lygephila lubrosa</emphasis>
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</taxonomicName>
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Staudinger, 1901 is a separate taxon different from
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<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="lubrica">
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<emphasis italics="true" pageId="5" pageNumber="112">Lygephila lubrica</emphasis>
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</taxonomicName>
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, but they failed to define this taxon, and did not recognize that the yellowish hindwinged populations include two different species.
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112">
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The main fault of the
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<normalizedToken originalValue="Stshetkins’">Stshetkins'</normalizedToken>
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work is the lack of definition of
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrosa">
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<emphasis italics="true" pageId="6" pageNumber="113">
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<pageBreakToken pageId="6" pageNumber="113" start="start">Lygephila</pageBreakToken>
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lubrosa
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</emphasis>
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</taxonomicName>
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Staudinger, 1901. In their article they provided the following description of the genitalia of
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrosa">
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<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa</emphasis>
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</taxonomicName>
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:
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<normalizedToken originalValue="самца">samtsa</normalizedToken>
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.
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<normalizedToken originalValue="изогнутый">izognutyi</normalizedToken>
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,
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,
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.
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.
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<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
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<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
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</taxonomicName>
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;
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<normalizedToken originalValue="конец">konets</normalizedToken>
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(
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<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
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<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
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</taxonomicName>
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).
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,
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<normalizedToken originalValue="у">u</normalizedToken>
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<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
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<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
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</taxonomicName>
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.
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<normalizedToken originalValue="В">V</normalizedToken>
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<normalizedToken originalValue="оральной">oralnoi</normalizedToken>
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,
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|
||
<normalizedToken originalValue="вентральному">ventralnomu</normalizedToken>
|
||
<normalizedToken originalValue="краю">krayu</normalizedToken>
|
||
<normalizedToken originalValue="вальвы">valvy</normalizedToken>
|
||
<normalizedToken originalValue="и">i</normalizedToken>
|
||
<normalizedToken originalValue="при">pri</normalizedToken>
|
||
<normalizedToken originalValue="этом">etom</normalizedToken>
|
||
<normalizedToken originalValue="не">ne</normalizedToken>
|
||
<normalizedToken originalValue="образует">obrazuet</normalizedToken>
|
||
<normalizedToken originalValue="резкого">rezkogo</normalizedToken>
|
||
<normalizedToken originalValue="угла">ugla</normalizedToken>
|
||
<normalizedToken originalValue="с">s</normalizedToken>
|
||
<normalizedToken originalValue="бугорком-гарпой">bugorkom-garpoi</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="имеющегося">imeyushchegosya</normalizedToken>
|
||
<normalizedToken originalValue="у">u</normalizedToken>
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
Frr.
|
||
<normalizedToken originalValue="Нижняя">Nizhnyaya</normalizedToken>
|
||
<normalizedToken originalValue="фультура">fultura</normalizedToken>
|
||
<normalizedToken originalValue="под">pod</normalizedToken>
|
||
<normalizedToken originalValue="эдеагусом">edeagusom</normalizedToken>
|
||
<normalizedToken originalValue="без">bez</normalizedToken>
|
||
<normalizedToken originalValue="особого">osobogo</normalizedToken>
|
||
<normalizedToken originalValue="изгиба">izgiba</normalizedToken>
|
||
<normalizedToken originalValue="прямо">pryamo</normalizedToken>
|
||
<normalizedToken originalValue="идет">idet</normalizedToken>
|
||
<normalizedToken originalValue="в">v</normalizedToken>
|
||
<normalizedToken originalValue="сторону">storonu</normalizedToken>
|
||
<normalizedToken originalValue="саккуса">sakkusa</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="как">kak</normalizedToken>
|
||
<normalizedToken originalValue="у">u</normalizedToken>
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lusoria" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lusoria">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lusoria</emphasis>
|
||
</taxonomicName>
|
||
L." The translation of this text is as follows: "The male genitalia are symmetrical. Uncus slightly curved, dilated in the middle part, pointed. Valva elongated with slightly convex dorsal and ventral edges. Apex of clasper digitiform, longer than that of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
; its end close to the dorsal margin of the valva (as for
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
, the latter is far from reaching the margin). The end of the valva from the base of the clasper till its back end is noticeably longer than that of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
. In the oral [basal] third of the valva, the longitudinal chitinized fold of the clasper is directed orally [basally] and is slightly curved towards the ventral margin of the valva without forming an abrupt angle with the hump-harpe, which is typical for
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
Frr. Lower fultura [juxta] under aedeagus almost straight and directed towards the saccus as
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lusoria" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lusoria">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lusoria</emphasis>
|
||
</taxonomicName>
|
||
L." This description is contradictory as it includes characteristics of both yellow hindwinged species occurring in Central Asia. To be precise, "Uncus dilated in the middle part" is only typical for the moth (
|
||
<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="kazachkaratavika">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila kazachkaratavika</emphasis>
|
||
</taxonomicName>
|
||
) from Kyrgyzstan (lake Issyk Kul) and Kazakhstan (city of Kizilorda and station Baigacum on the river Syr Darja) (Figs
|
||
<figureCitation captionStart="Figures 33–38" captionStartId="F5" captionText="Figures 33 - 38. Clasping apparatus. 33, 34 Lygephila lubrosa lubrosa 33 neotype 35, 36 Lygephila lubrosa orbonaria 35 neotype 37, 38 Lygephila kazachkaratavika 37 neotype." pageId="6" pageNumber="113">37</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 33–38" captionStartId="F5" captionText="Figures 33 - 38. Clasping apparatus. 33, 34 Lygephila lubrosa lubrosa 33 neotype 35, 36 Lygephila lubrosa orbonaria 35 neotype 37, 38 Lygephila kazachkaratavika 37 neotype." pageId="6" pageNumber="113">38</figureCitation>
|
||
); "Apex of clasper digitiform and longer than that of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
; its end close to the dorsal margin of the valva" is only typical for the moths (
|
||
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrosa">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa</emphasis>
|
||
</taxonomicName>
|
||
) from Kazakhstan (river Ili) and Tajikistan (river Pianj) (Figs
|
||
<figureCitation captionStart="Figures 33–38" captionStartId="F5" captionText="Figures 33 - 38. Clasping apparatus. 33, 34 Lygephila lubrosa lubrosa 33 neotype 35, 36 Lygephila lubrosa orbonaria 35 neotype 37, 38 Lygephila kazachkaratavika 37 neotype." pageId="6" pageNumber="113">33</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figures 33–38" captionStartId="F5" captionText="Figures 33 - 38. Clasping apparatus. 33, 34 Lygephila lubrosa lubrosa 33 neotype 35, 36 Lygephila lubrosa orbonaria 35 neotype 37, 38 Lygephila kazachkaratavika 37 neotype." pageId="6" pageNumber="113">36</figureCitation>
|
||
). The female genitalia are described as follows:
|
||
<normalizedToken originalValue="“В">"V</normalizedToken>
|
||
<normalizedToken originalValue="гениталиях">genitaliyakh</normalizedToken>
|
||
<normalizedToken originalValue="самки">samki</normalizedToken>
|
||
<normalizedToken originalValue="копулятивная">kopulyativnaya</normalizedToken>
|
||
<normalizedToken originalValue="сумка">sumka</normalizedToken>
|
||
<normalizedToken originalValue="мешковидная">meshkovidnaya</normalizedToken>
|
||
<normalizedToken originalValue="и">i</normalizedToken>
|
||
<normalizedToken originalValue="вместе">vmeste</normalizedToken>
|
||
<normalizedToken originalValue="с">s</normalizedToken>
|
||
<normalizedToken originalValue="едва">edva</normalizedToken>
|
||
<normalizedToken originalValue="заметным">zametnym</normalizedToken>
|
||
<normalizedToken originalValue="бугорком-буллой">bugorkom-bulloi</normalizedToken>
|
||
<normalizedToken originalValue="вся">vsya</normalizedToken>
|
||
<normalizedToken originalValue="перепончатая">pereponchataya</normalizedToken>
|
||
(
|
||
<normalizedToken originalValue="у">u</normalizedToken>
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="булла">bulla</normalizedToken>
|
||
<normalizedToken originalValue="конусовидная">konusovidnaya</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="хитинизированная">khitinizirovannaya</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="как">kak</normalizedToken>
|
||
<normalizedToken originalValue="и">i</normalizedToken>
|
||
<normalizedToken originalValue="весь">ves</normalizedToken>
|
||
<normalizedToken originalValue="проток">protok</normalizedToken>
|
||
<normalizedToken originalValue="и">i</normalizedToken>
|
||
<normalizedToken originalValue="часть">chast</normalizedToken>
|
||
<normalizedToken originalValue="сумки">sumki</normalizedToken>
|
||
).
|
||
<normalizedToken originalValue="Проток">Protok</normalizedToken>
|
||
<normalizedToken originalValue="сумки">sumki</normalizedToken>
|
||
<normalizedToken originalValue="значительно">znachitelno</normalizedToken>
|
||
<normalizedToken originalValue="короче">koroche</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="его">ego</normalizedToken>
|
||
<normalizedToken originalValue="оральная">oralnaya</normalizedToken>
|
||
<normalizedToken originalValue="часть">chast</normalizedToken>
|
||
<normalizedToken originalValue="перепончатая”">pereponchataya"</normalizedToken>
|
||
. The translation is as follows: "In the female genitalia the copulative pouch [corpus bursae] is saccular and all membranous along with a barely noticeable bulla (while the bulla of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
is conical and chitinized as well as the whole antrum and part of the pouch [corpus bursae]). The antrum is significantly shorter with membranous oral [basal] part." The characteristics mentioned as "the antrum is significantly shorter than that of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
" partially corresponds to the moths from the Ili region. However, it is not diagnostic because in many specimens of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
the antrum has the same length. The antrum of the moths (
|
||
<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="kazachkaratavika">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila kazachkaratavika</emphasis>
|
||
</taxonomicName>
|
||
) from the Issyk Kul region and the river Syr-Darja is one and a half times longer than that of
|
||
<taxonomicName baseAuthorityName="Freyer" baseAuthorityYear="1842" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrica" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrica">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrica</emphasis>
|
||
</taxonomicName>
|
||
and two times longer than that of the moths from the valley of the river Ili (
|
||
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrosa">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa</emphasis>
|
||
</taxonomicName>
|
||
). The other characteristics mentioned by the authors are general, non-autapomorphic and unsuitable for determination.
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">
|
||
In the same work the authors described two subspecies of
|
||
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrosa">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa</emphasis>
|
||
</taxonomicName>
|
||
on the basis of external characteristics, admitting that the two subspecies do not differ in genitalia structure from the nominotypical subspecies. However, the moths from the Kazakh Karatau, station Balamurum collected by V. Kozhantshikov in 1909 (
|
||
<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="kazachkaratavika">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila kazachkaratavika</emphasis>
|
||
</taxonomicName>
|
||
) differ significantly in their genitalia structure from the moths from the valleys of the river Ili (
|
||
<taxonomicName authority="lubrosa" authorityName="lubrosa (Staudinger" authorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa subsp. lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="subSpecies" species="lubrosa" subSpecies="lubrosa">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa lubrosa</emphasis>
|
||
</taxonomicName>
|
||
) and the river Pianj (
|
||
<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa subsp. orbonaria" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="subSpecies" species="lubrosa" subSpecies="orbonaria">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa orbonaria</emphasis>
|
||
</taxonomicName>
|
||
). All above-mentioned data prove that the authors did not consider the subject of their research, which caused unsatisfactory results and added further difficulties for the clarification of this species-complex. A further difficulty is that the authors did not define holotypes or paratypes (or simply type series) for the newly described taxa. According to the information from the museum curators of ZISP and IZIP, they do not possess the aforesaid type specimens with the corresponding type labels.
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">
|
||
In order to correctly identify the taxa of this species complex, in view of complexity of the current taxonomic situation, and to give an accurate definition of
|
||
<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa" order="Lepidoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="lubrosa">
|
||
<emphasis italics="true" pageId="6" pageNumber="113">Lygephila lubrosa</emphasis>
|
||
</taxonomicName>
|
||
, it is necessary to designate a neotype of this taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="113" type="diagnosis">
|
||
<paragraph pageId="6" pageNumber="113">Diagnosis.</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">
|
||
Easily distinguishable from all other members of the species group by its unicolorous grey forewings. Comparing the genital structures of the taxa of the group, it differs from all related species by the narrow uncus without a real dilatation (only some slight thickening is present), the wide valva, and the subapically located, strong clasper with its tip reaching the valval edge (males); and by the funnel-shaped antrum bent dorsally at 45 degrees, being a unique female character for the whole genus (Fig.
|
||
<figureCitation captionStart="Figures 46–63" captionStartId="F8" captionText="Figures 46 - 63. Female genitalia. 46 - 57 Lygephila lubrica 58 - 61 Lygephila lubrosa lubrosa 58 ventral view 59 dorsal view 60 lateral view 61 lateral view 62, 63 Lygephila kazachkaratavika." pageId="6" pageNumber="113">60</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="114" pageId="6" pageNumber="113" type="description">
|
||
<paragraph pageId="6" pageNumber="113">Description.</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">Wingspan 42-46 mm. Head and body grey with some brownish scales; collar chocolate brown. Forewing almost unicolorous, wing pattern poorly developed; subbasal line hardly traceable, represented by groups of dark scales on veins; antemedial line semicircular; medial shade not expressed; reniform stigma small, indistinct, with one or two black dots basad; orbicular stigma small dot-like; postmedial and subterminal lines distinct; terminal line a row of black dots on veins. Hindwing pale ochreous; transverse line not discernible; outer dark third with sharply defined inner margin; fringes ochreous.</paragraph>
|
||
<paragraph lastPageId="7" lastPageNumber="114" pageId="6" pageNumber="113">
|
||
<emphasis bold="true" pageId="6" pageNumber="113">Male genitalia</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 33–38" captionStartId="F5" captionText="Figures 33 - 38. Clasping apparatus. 33, 34 Lygephila lubrosa lubrosa 33 neotype 35, 36 Lygephila lubrosa orbonaria 35 neotype 37, 38 Lygephila kazachkaratavika 37 neotype." pageId="6" pageNumber="113">33</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 33–38" captionStartId="F5" captionText="Figures 33 - 38. Clasping apparatus. 33, 34 Lygephila lubrosa lubrosa 33 neotype 35, 36 Lygephila lubrosa orbonaria 35 neotype 37, 38 Lygephila kazachkaratavika 37 neotype." pageId="6" pageNumber="113">34</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 42–45" captionStartId="F7" captionText="Figures 42 - 45. Vesica structure. 42, 43 Lygephila kazachkaratavika, neotype, Kazakhstan, Balamurum, slide No. OP 2009 m 42 lateral view 43 lateral view opposite side 44, 45 Lygephila lubrosa lubrosa, neotype, Kazakhstan, Ili river, slide No. OP 2082 m 44 lateral view 45 lateral view opposite side." pageId="6" pageNumber="113">44</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 42–45" captionStartId="F7" captionText="Figures 42 - 45. Vesica structure. 42, 43 Lygephila kazachkaratavika, neotype, Kazakhstan, Balamurum, slide No. OP 2009 m 42 lateral view 43 lateral view opposite side 44, 45 Lygephila lubrosa lubrosa, neotype, Kazakhstan, Ili river, slide No. OP 2082 m 44 lateral view 45 lateral view opposite side." pageId="6" pageNumber="113">45</figureCitation>
|
||
). Uncus long, stout, slightly thickened medially with skewed fine tip, anal tube membranous with oval hardening of tissue - scaphial crown on scaphium and sclerotized fig on subscaphium; valva elongated, wide, with parallel margins in the middle, valval apex rounded; clasper digitiform, strong, thickened with wide base, placed subapically, asymmetrical, left one shorter than the right one, both almost reach valval costa. Aedeagus tubular with narrow, long, acute sclerotised lamina on ventral side of carina. Vesica globular, multidiverticulate, membranous; basal diverticulum small; medial diverticulum large with small lateral hemispherical bulging; 1st terminal diverticulum large, more or less wedge shaped,
|
||
<pageBreakToken pageId="7" pageNumber="114" start="start">membranous</pageBreakToken>
|
||
with cauliflower-like part bearing numerous small pockets; 2nd terminal diverticulum large, cone shaped, scobinated on top; 3rd terminal diverticulum medium-sized, bifurcated, Y-shaped; 4th terminal diverticulum large, bilobate, located oppositely to the 3rd medial diverticulum; terminal tube membranous as long as aedeagus, opening point of terminal tube located near to carina.
|
||
</paragraph>
|
||
<paragraph pageId="7" pageNumber="114">
|
||
<emphasis bold="true" pageId="7" pageNumber="114">Female genitalia</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 46–63" captionStartId="F8" captionText="Figures 46 - 63. Female genitalia. 46 - 57 Lygephila lubrica 58 - 61 Lygephila lubrosa lubrosa 58 ventral view 59 dorsal view 60 lateral view 61 lateral view 62, 63 Lygephila kazachkaratavika." pageId="7" pageNumber="114">58</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figures 46–63" captionStartId="F8" captionText="Figures 46 - 63. Female genitalia. 46 - 57 Lygephila lubrica 58 - 61 Lygephila lubrosa lubrosa 58 ventral view 59 dorsal view 60 lateral view 61 lateral view 62, 63 Lygephila kazachkaratavika." pageId="7" pageNumber="114">61</figureCitation>
|
||
). Ovipositor relatively large, broad, papillae anales hairy with long setae on apical edges. Apophyses anteriores stout, apophyses posteriores thin, longer than apophyses anteriores. Antrum funnel shaped, bent dorsally at 45 degrees, boomerang shaped from lateral view; ostium bursae broad, posterior margin U-shaped; ductus bursae medium sized; appendix bursae small; corpus bursae membranous, bevelled oval.
|
||
</paragraph>
|
||
<caption pageId="7" pageNumber="114" start="Figures 11–20" startId="F2">
|
||
<paragraph pageId="7" pageNumber="114">
|
||
<emphasis bold="true" pageId="7" pageNumber="114">Figures 11-20.</emphasis>
|
||
Adults.
|
||
<emphasis bold="true" pageId="7" pageNumber="114">11-16</emphasis>
|
||
<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" baseAuthorityName="Pekarsky" baseAuthorityYear="2014" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila kazachkaratavika" order="Lepidoptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="kazachkaratavika">
|
||
<emphasis italics="true" pageId="7" pageNumber="114">Lygephila kazachkaratavika</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="7" pageNumber="114">11</emphasis>
|
||
neotype, ♂, Balamurum
|
||
<emphasis bold="true" pageId="7" pageNumber="114">12</emphasis>
|
||
♀, Kazakhstan, Baigacum
|
||
<emphasis bold="true" pageId="7" pageNumber="114">13</emphasis>
|
||
♂, Kazakhstan, Taldy-Kurgan reg.
|
||
<emphasis bold="true" pageId="7" pageNumber="114">14</emphasis>
|
||
♀, Kazakstan, Baigacum
|
||
<emphasis bold="true" pageId="7" pageNumber="114">15</emphasis>
|
||
♂, Kyrgyzstan, Issyk Kul
|
||
<emphasis bold="true" pageId="7" pageNumber="114">16</emphasis>
|
||
♀, Kyrgyzstan, Issyk Kul
|
||
<emphasis bold="true" pageId="7" pageNumber="114">17, 18</emphasis>
|
||
<taxonomicName authority="lubrosa" authorityName="lubrosa (Staudinger" authorityYear="1901" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa subsp. lubrosa" order="Lepidoptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="lubrosa" subSpecies="lubrosa">
|
||
<emphasis italics="true" pageId="7" pageNumber="114">Lygephila lubrosa lubrosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="7" pageNumber="114">17</emphasis>
|
||
neotype, ♂, Kazakhstan, Ili river
|
||
<emphasis bold="true" pageId="7" pageNumber="114">18</emphasis>
|
||
♀, Kazakhstan, Baigacum
|
||
<emphasis bold="true" pageId="7" pageNumber="114">19, 20</emphasis>
|
||
<taxonomicName authorityName="Stshetkin YuL & Stshetkin YuYu, 1994" authorityYear="1997" class="Insecta" family="Erebidae" genus="Lygephila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygephila lubrosa subsp. orbonaria" order="Lepidoptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="lubrosa" subSpecies="orbonaria">
|
||
<emphasis italics="true" pageId="7" pageNumber="114">Lygephila lubrosa orbonaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="7" pageNumber="114">19</emphasis>
|
||
neotype, ♂, Tajikistan, Pianj river
|
||
<emphasis bold="true" pageId="7" pageNumber="114">20</emphasis>
|
||
♂, Tajikistan, Pianj river.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="114" type="destribution">
|
||
<paragraph pageId="7" pageNumber="114">Destribution.</paragraph>
|
||
<paragraph pageId="7" pageNumber="114">Kazakhstan, valley of the river Ili.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |