treatments-xml/data/A7/2C/87/A72C87FBFFC0FFE20EC80B4B08969114.xml

<document id="B19A3F57D8F1DF6CA297D592717AC083" ID-CLB-Dataset="22193" ID-DOI="10.5852/ejt.2018.467" ID-GBIF-Dataset="7a1afb26-e06c-43fc-bc68-fff2a45859c0" ID-Zenodo-Dep="3824775" ID-ZooBank="8D27AAB8-B7F2-424C-B1A6-66FEFA66EDFF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589386207548" checkinUser="valdenar" docAuthor="Cupello, Mario &amp; Vaz-De, Fernando Z." docDate="2018" docId="A72C87FBFFC0FFE20EC80B4B08969114" docLanguage="en" docName="ejt-467_cupello_vaz-de-mello.pdf.imf" docOrigin="European Journal of Taxonomy 467" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sylvicanthon aequinoctialis Cupello &amp; Vaz-De 2018, comb. nov." docType="treatment" docVersion="7" lastPageNumber="95" masterDocId="5B15FF83FF94FFBD0F1B0C210B2E905D" masterDocTitle="A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter &amp; Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of ‘ Canthon sensu lato’" masterLastPageNumber="205" masterPageNumber="1" pageNumber="85" updateTime="1698839428571" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E14472D013FB9D730685FC40F33C0471" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FBA1B96862E8D623788350D8391EE902">
<mods:title id="D68C9148B4C65E0D0A616FB7212A1AC9">A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter &amp; Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of ‘ Canthon sensu lato’</mods:title>
</mods:titleInfo>
<mods:name id="0CD7326CF4F66DB58B40FDD5C47F6090" type="personal">
<mods:role id="32EE7B7A610099E9F82FF8A52EF4C354">
<mods:roleTerm id="0FAF824098962D98FEDC522E75C13EB7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CE878015F363189C7F1CC0918999C160">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="E9400242B3AAF105153F9C6B9FAAF317" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D&amp;2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="39083F28DFBFD6DB3C6AE5C734546C8A">Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, UFRJ, Quinta da Boa Vista, São Cristóvão, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil.</mods:affiliation>
<mods:nameIdentifier id="379C30DFE7DB646C856C0B9779A5CB1D" type="email">mcupello@hotmail.com&amp;vazdemello@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="FA9D9E6CC0F33787B3D7DDD97ADB5121" type="personal">
<mods:role id="DC828CDEC8690A52504526EA5D7D915A">
<mods:roleTerm id="17A5A9DA97DFC4916411C4ABEFD3FAB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D1602178EB7668889FF2BE7523C5F786">Vaz-De, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="343EF97AE04240D12440C1022C73B9C6">text</mods:typeOfResource>
<mods:relatedItem id="41A97C291BBF3ED37ECF9DCEB3FCBC1A" type="host">
<mods:titleInfo id="F4A7757DE08E0BB237A4E79106617599">
<mods:title id="2881054CAFCBE7E772F889C91D60BC9F">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7775919ED9B517CEC446459230C0DB2E">
<mods:date id="EEBCF47C4B9045DD4B62522ACD746593">2018</mods:date>
<mods:detail id="069958BDFBE4F56599C1FBE17D737A60" type="pubDate">
<mods:number id="C62E59B4918ADAF4AB8C0064B94230F7">2018-10-16</mods:number>
</mods:detail>
<mods:detail id="5C42E1C44534B1C175169394338FA222" type="volume">
<mods:number id="BF54F4368847EFC922C46C588E452484">467</mods:number>
</mods:detail>
<mods:extent id="188906D5AF508C742E9FF522300B0CE1" unit="page">
<mods:start id="D652D036B961E7EBDBFCC0184012807C">1</mods:start>
<mods:end id="44F777AF5110C3827F0EAB07C59BAE12">205</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="743BE44B4EFB25D33338B3AE87AC102C">journal article</mods:classification>
<mods:identifier id="02DF80655903F76D488458EC90C431D9" type="CLB-Dataset">22193</mods:identifier>
<mods:identifier id="21B9279CB49F1F41BD8D89EA2996CD01" type="DOI">10.5852/ejt.2018.467</mods:identifier>
<mods:identifier id="C18C1D4E05F3E82090BC9055B51FFFDD" type="GBIF-Dataset">7a1afb26-e06c-43fc-bc68-fff2a45859c0</mods:identifier>
<mods:identifier id="6800D9DA809D82FD90B5172F178EC551" type="Zenodo-Dep">3824775</mods:identifier>
<mods:identifier id="993E354102588BC699456BD538DAD236" type="ZooBank">8D27AAB8-B7F2-424C-B1A6-66FEFA66EDFF</mods:identifier>
</mods:mods>
<treatment id="A72C87FBFFC0FFE20EC80B4B08969114" ID-DOI="http://doi.org/10.5281/zenodo.3846309" ID-GBIF-Taxon="164129494" ID-Zenodo-Dep="3846309" LSID="urn:lsid:plazi:treatment:A72C87FBFFC0FFE20EC80B4B08969114" httpUri="http://treatment.plazi.org/id/A72C87FBFFC0FFE20EC80B4B08969114" lastPageId="95" lastPageNumber="95" pageId="84" pageNumber="85">
<subSubSection id="679F6566FFC0FFE90EC80B4B0F4F97D8" box="[467,1121,1898,1925]" pageId="84" pageNumber="85" type="nomenclature">
<paragraph id="2F3A36EDFFC0FFE90EC80B4B0F4F97D8" blockId="84.[464,1125,1898,1961]" box="[467,1121,1898,1925]" pageId="84" pageNumber="85">
<heading id="74728181FFC0FFE90EC80B4B0F4F97D8" box="[467,1121,1898,1925]" centered="true" fontSize="11" level="2" pageId="84" pageNumber="85" reason="2">
<taxonomicName id="E8854D6EFFC0FFE90EC80B4B08F997D8" ID-CoL="53N4T" authority="(Harold, 1868)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[467,983,1898,1925]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="aequinoctialis" status="comb. nov.">
<emphasis id="1DF1EAFFFFC0FFE90EC80B4B080D97D8" bold="true" box="[467,803,1898,1925]" italics="true" pageId="84" pageNumber="85">Sylvicanthon aequinoctialis</emphasis>
(Harold, 1868)
</taxonomicName>
<taxonomicNameLabel id="06C25784FFC0FFE90CC40B4B0F4F97D8" box="[991,1121,1898,1925]" pageId="84" pageNumber="85" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="679F6566FFC0FFE90ECB0BAF0F4B97F5" box="[464,1125,1934,1961]" pageId="84" pageNumber="85" type="description">
<paragraph id="2F3A36EDFFC0FFE90ECB0BAF0F4B97F5" blockId="84.[464,1125,1898,1961]" box="[464,1125,1934,1961]" pageId="84" pageNumber="85">
<figureCitation id="B7BE2A68FFC0FFE90ECB0BAF091F97F5" box="[464,561,1934,1960]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,372,1844]" captionTargetId="figure@21.[189,1398,372,1849]" captionTargetPageId="21" captionText="Fig. 7. Pterothorax, ventrolateral views. A. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov. B. S. obscurus (Schmidt, 1920). C. Canthon xanthopus Blanchard, 1846. Abbreviations: MsC = mesocoxa; MMsC = external margin of mesocoxa; CMtE = centre of metaventrite; LMtE = lateral region of metaventrite; MtC = metacoxa." figureDoi="http://doi.org/10.5281/zenodo.3824793" httpUri="https://zenodo.org/record/3824793/files/figure.png" pageId="84" pageNumber="85">Figs 7A</figureCitation>
, 
<figureCitation id="B7BE2A68FFC0FFE90D260BAF094097F5" box="[573,622,1934,1960]" captionStart="Fig" captionStartId="25.[189,232,1787,1813]" captionTargetBox="[196,1391,645,1731]" captionTargetId="figure@25.[189,1399,645,1755]" captionTargetPageId="25" captionText="Fig. 11. Morphological diversity in the protibiae of Sylvicanthon Halffter &amp; Martínez, 1977. A. S. enkerlini (Martínez et al., 1964) comb. nov. B. S. aequinoctialis (Harold, 1869) comb. nov. C. S. securus (Schmidt, 1920) comb. nov. D. S. obscurus (Schmidt, 1920). E. S. furvus (Schmidt, 1920). F. S. bridarollii (Martínez, 1949) (southern populations). G. S. bridarollii (northern populations). H. S. attenboroughi sp. nov. I. S. edmondsi sp. nov. J. S. candezei (Harold, 1869). Note the variation related to the presence and degree of development of an expansion on the internal edge of protibiae and to the number, size and position of the external teeth." figureDoi="http://doi.org/10.5281/zenodo.3824801" httpUri="https://zenodo.org/record/3824801/files/figure.png" pageId="84" pageNumber="85">11B</figureCitation>
, 
<figureCitation id="B7BE2A68FFC0FFE90D600BAF09BB97F4" box="[635,661,1934,1961]" captionStart="Fig" captionStartId="30.[189,232,1790,1816]" captionTargetBox="[196,1383,292,1753]" captionTargetId="figure@30.[196,1383,289,1754]" captionTargetPageId="30" captionText="Fig. 15. Morphological variation and sexual dimorphism in the protibial spur of Sylvicanthon. A. S. enkerlini (Martínez et al., 1964) comb. nov., ♂. B. S. foveiventris (Schmidt, 1920), ♂. C. S. candezei (Harold, 1869), ♂. D. S. genieri sp. nov., ♂. E. S. aequinoctialis (Harold, 1868), ♂. F. S. aequinoctialis comb. nov., ♀. G. S. proseni (Martínez, 1949) stat. et comb. nov., ♂. H. S. proseni stat. et comb. nov., ♀. I. S. bridarollii (Martínez, 1949), ♂. J. S. seag sp. nov., ♂. K. S. edmondsi sp. nov.,♂. L. S. attenboroughi sp. nov., ♂. M. S. obscurus (Schmidt, 1920), ♂. N. S. furvus (Schmidt, 1920), ♂. O. S. securus (Schmidt, 1920) comb. nov., ♂." figureDoi="http://doi.org/10.5281/zenodo.3824809" httpUri="https://zenodo.org/record/3824809/files/figure.png" pageId="84" pageNumber="85">15</figureCitation>
E–F, 17E, 20, 28A–B, 29A–B, 30, 31A
</paragraph>
</subSubSection>
<subSubSection id="679F6566FFC0FFE80FA60BF00FE5955F" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="reference_group">
<paragraph id="2F3A36EDFFC0FFE90FA60BF0087A97B6" blockId="84.[189,852,2000,2027]" box="[189,852,2000,2027]" pageId="84" pageNumber="85">
<treatmentCitationGroup id="0F9511C3FFC0FFE90FA60BF0087A97B6" box="[189,852,2000,2027]" pageId="84" pageNumber="85">
<treatmentCitation id="AE2410FCFFC0FFE90FA60BF009B197B6" author="Harold E." box="[189,671,2001,2027]" page="2" pageId="84" pageNumber="85" year="1868">
<taxonomicName id="E8854D6EFFC0FFE90FA60BF009B197B6" ID-CoL="QPB6" authority="Harold, 1868 a: 2" authorityName="Harold" authorityPageNumber="2" authorityYear="1868" box="[189,671,2001,2027]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC0FFE90FA60BF00AE197B6" box="[189,463,2001,2027]" italics="true" pageId="84" pageNumber="85">Canthon aequinoctialis</emphasis>
<bibRefCitation id="4B144B1CFFC0FFE90ECC0BF009B197B6" author="Harold E." box="[471,671,2001,2027]" pageId="84" pageNumber="85" pagination="1 - 144" refId="ref136501" refString="Harold E. 1868 a. Monographie der Gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120104" type="journal article" year="1868">Harold, 1868a: 2</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 5, 14, 79, 141.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60D2A09DB9178" blockId="85.[189,1399,267,1283]" box="[189,757,267,294]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60D2A09DB9178" box="[189,757,267,294]" pageId="85" pageNumber="86">
“ 
<taxonomicName id="E8854D6EFFC1FFE80FD10D2A0ADC9178" box="[202,498,267,293]" class="Insecta" family="Scarabaeidae" genus="Coprobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80FD10D2A0ADC9178" box="[202,498,267,293]" italics="true" pageId="85" pageNumber="86">Coprobius aequinoctialis</emphasis>
</taxonomicName>
” – 
<treatmentCitation id="AE2410FCFFC1FFE80D060D2A09DE9178" author="Dejean P. F. M. A." box="[541,752,267,293]" page="136" pageId="85" pageNumber="86" year="1833">
<bibRefCitation id="4B144B1CFFC1FFE80D060D2A09DE9178" author="Dejean P. F. M. A." box="[541,752,267,293]" pageId="85" pageNumber="86" pagination="1833 - 1836" refId="ref133982" refString="Dejean P. F. M. A. 1833. [Livraison 2.]. In: Dejean P. F. M. A. 1833 - 1836. Catalogue des Coleopteres de la collection de M. le Comte Dejean: 97 - 176. Mequignon-Marvis Pere et Fils, Paris. [Publication date according to Madge 1988.] Available from http: // biodiversitylibrary. org / page / 9322478 [accessed 30 May 2018]." type="journal article" year="1833">Dejean 1833: 136</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60D0E0A829130" blockId="85.[189,1399,267,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60D0E0A829130" pageId="85" pageNumber="86">
“ 
<taxonomicName id="E8854D6EFFC1FFE80FD10D0E0ADF9114" box="[202,497,303,329]" class="Insecta" family="Scarabaeidae" genus="Coprobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80FD10D0E0ADF9114" box="[202,497,303,329]" italics="true" pageId="85" pageNumber="86">Coprobius aequinoctialis</emphasis>
</taxonomicName>
” – Dejean 1836: 151. — 
<treatmentCitation id="AE2410FCFFC1FFE80C0D0D0E08CE9114" author="Harold E." box="[790,992,303,329]" page="79" pageId="85" pageNumber="86" year="1868">
<bibRefCitation id="4B144B1CFFC1FFE80C0D0D0E08CE9114" author="Harold E." box="[790,992,303,329]" pageId="85" pageNumber="86" pagination="1 - 144" refId="ref136501" refString="Harold E. 1868 a. Monographie der Gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120104" type="journal article" year="1868">Harold 1868a: 79</bibRefCitation>
</treatmentCitation>
(as synonym of 
<taxonomicName id="E8854D6EFFC1FFE80B850D0E0E4B9114" box="[1182,1381,303,329]" class="Insecta" family="Scarabaeidae" genus="Coprobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80B850D0E0E4B9114" box="[1182,1381,303,329]" italics="true" pageId="85" pageNumber="86">C. aequinoctialis</emphasis>
</taxonomicName>
). 
<emphasis id="1DF1EAFFFFC1FFE80FF10D730A8B9131" bold="true" box="[234,421,338,364]" pageId="85" pageNumber="86">Nomen nudum</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60D5409899365" blockId="85.[189,1399,267,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60D5409899365" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA60D540AFD91D2" ID-CoL="QPB6" authorityName="Harold" authorityYear="1868" box="[189,467,373,399]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80FA60D540AFD91D2" box="[189,467,373,399]" italics="true" pageId="85" pageNumber="86">Canthon aequinoctialis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80EE20D5409CD91D2" author="Harold E." box="[505,739,373,399]" page="989" pageId="85" pageNumber="86" year="1869">
<bibRefCitation id="4B144B1CFFC1FFE80EE20D5409CD91D2" author="Harold E." box="[505,739,373,399]" pageId="85" pageNumber="86" refId="ref136616" refString="Harold E. 1869 b. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae: 979 - 1346. Sumptu E. H. Gummi, Munich. Available from http: // biodiversitylibrary. org / page / 9641549 [accessed 30 May 2018]." type="url" year="1869">Harold 1869b: 989</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="AE2410FCFFC1FFE80DE80D54084F91D2" author="Harold E." box="[755,865,373,399]" page="16" pageId="85" pageNumber="86" year="1880">
<bibRefCitation id="4B144B1CFFC1FFE80DE80D54084F91D2" author="Harold E." box="[755,865,373,399]" pageId="85" pageNumber="86" pagination="13 - 46" refId="ref136720" refString="Harold E. 1880. Verzeichniss der von E. Steinheil in Neu-Granada gesammelten coprophagen Lamellicornien. Entomologischen Vereine zu Stettin 41: 13 - 46. Available from https: // biodiversitylibrary. org / page / 8989387 [accessed 30 May 2018]." type="journal article" year="1880">1880: 16</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80C810D570F7891D2" author="Bates H. W." box="[922,1110,373,400]" page="33" pageId="85" pageNumber="86" year="1887">
<bibRefCitation id="4B144B1CFFC1FFE80C810D570F7891D2" author="Bates H. W." box="[922,1110,373,400]" pageId="85" pageNumber="86" pagination="25 - 64" refId="ref131049" refString="Bates H. W. 1887. Tribe Lamellicornia In: Bates H. W. 1886 - 1890. Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor &amp; Francis, London. Available from http: // biodiversitylibrary. org / page / 578700 [accessed 30 May 2018]." type="book chapter" year="1887">Bates 1887: 33</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80B950D540E6491D2" author="Gillet J. J. E." box="[1166,1354,373,399]" page="27" pageId="85" pageNumber="86" year="1911">
<bibRefCitation id="4B144B1CFFC1FFE80B950D540E6491D2" author="Gillet J. J. E." box="[1166,1354,373,399]" pageId="85" pageNumber="86" refId="ref135729" refString="Gillet J. J. E. 1911. Coleopterorum catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin." type="book" year="1911">Gillet 1911: 27</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80FF10DB80AEC91EE" author="Schmidt A." box="[234,450,409,435]" page="64" pageId="85" pageNumber="86" year="1922">
<bibRefCitation id="4B144B1CFFC1FFE80FF10DB80AEC91EE" author="Schmidt A." box="[234,450,409,435]" pageId="85" pageNumber="86" pagination="61 - 103" refId="ref145305" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from http: // biodiversitylibrary. org / page / 13259701 [accessed 30 May 2018]." type="journal article" year="1922">Schmidt 1922: 64</bibRefCitation>
</treatmentCitation>
, 72. — 
<treatmentCitation id="AE2410FCFFC1FFE80D3D0DB8083491EE" author="Balthasar V." box="[550,794,409,435]" page="187" pageId="85" pageNumber="86" year="1939">
<bibRefCitation id="4B144B1CFFC1FFE80D3D0DB8083491EE" author="Balthasar V." box="[550,794,409,435]" pageId="85" pageNumber="86" pagination="179 - 238" refId="ref130824" refString="Balthasar V. 1939. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. Folia Zoologica et Hydrobiologica 9 (2): 179 - 238." type="journal article" year="1939">Balthasar 1939: 187</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="AE2410FCFFC1FFE80C330DB8088F91EE" author="Balthasar V." box="[808,929,409,435]" page="341" pageId="85" pageNumber="86" year="1941">
<bibRefCitation id="4B144B1CFFC1FFE80C330DB8088F91EE" author="Balthasar V." box="[808,929,409,435]" pageId="85" pageNumber="86" pagination="337 - 358" refId="ref130851" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrag Fauna Perus 1: 337 - 358." type="journal article" year="1941">1941: 341</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80B770DB80FCC91EE" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1132,1250,409,435]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80B770DB80FCC91EE" box="[1132,1250,409,435]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
); 1951: 326 (error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80EAA0D9D0907918B" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[433,553,444,470]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80EAA0D9D0907918B" box="[433,553,444,470]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80D750D9D0841918B" author="Martinez A." box="[622,879,444,470]" page="290" pageId="85" pageNumber="86" year="1949">
<bibRefCitation id="4B144B1CFFC1FFE80D750D9D0841918B" author="Martinez A." box="[622,879,444,470]" pageId="85" pageNumber="86" pagination="281 - 291" refId="ref139950" refString="Martinez A. 1949 a. Notas coleopterologicas III. Dos nuevas especies de Glaphyrocanthon Martinez, 1948, con clave para las especies del genero (Col. Scarab.). Anales de la Sociedad Cientifica Argentina 148: 281 - 291. Available from http: // biodiversitylibrary. org / page / 48991419 [accessed 30 May 2018]." type="journal article" year="1949">Martínez 1949a: 290</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80CBD0D9D0E3E918A" author="Vulcano M. A. &amp; Pereira F. S." box="[934,1296,444,471]" page="561" pageId="85" pageNumber="86" year="1967">
<bibRefCitation id="4B144B1CFFC1FFE80CBD0D9D0E3E918A" author="Vulcano M. A. &amp; Pereira F. S." box="[934,1296,444,471]" pageId="85" pageNumber="86" pagination="533 - 603" refId="ref147845" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5 (Zoologia): 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 561</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80A520D9D0A8E91A7" author="Gill B. D." page="225" pageId="85" pageNumber="86" year="1991">
<bibRefCitation id="4B144B1CFFC1FFE80A520D9D0A8E91A7" author="Gill B. D." pageId="85" pageNumber="86" pagination="211 - 229" refId="ref135689" refString="Gill B. D. 1991. Dung beetles in tropical American forests. In: Hanski I. &amp; Cambefort Y. (eds) Dung Beetle Ecology: 211 - 229. Princeton University Press, Princeton." type="book chapter" year="1991">Gill 1991: 225–226</bibRefCitation>
</treatmentCitation>
, 228. — 
<treatmentCitation id="AE2410FCFFC1FFE80D0A0DFE09D391A4" author="Escobar F." box="[529,765,479,505]" page="206" pageId="85" pageNumber="86" year="2000">
<bibRefCitation id="4B144B1CFFC1FFE80D0A0DFE09D391A4" author="Escobar F." box="[529,765,479,505]" pageId="85" pageNumber="86" pagination="197 - 210" refId="ref134781" refString="Escobar F. 2000 a. Diversidad y distribucion de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. Monografias Tercer Milenio 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="journal article" year="2000">Escobar 2000a: 206</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80CDC0DC10F1391A4" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[967,1085,480,506]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80CDC0DC10F1391A4" box="[967,1085,480,506]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
); 2000b: 114, 121 (mixed information with 
<taxonomicName id="E8854D6EFFC1FFE80EAD0E2209049240" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[438,554,515,541]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80EAD0E2209049240" box="[438,554,515,541]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80D7E0E2208559240" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[613,891,514,541]" page="135" pageId="85" pageNumber="86" year="2001">
<bibRefCitation id="4B144B1CFFC1FFE80D7E0E2208559240" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[613,891,514,541]" pageId="85" pageNumber="86" pagination="131 - 144" refId="ref141467" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina 
<emphasis id="1DF1EAFFFFC1FFE80DDD0E25082E9240" box="[710,768,515,541]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2001: 135
</bibRefCitation>
</treatmentCitation>
(mixed information with 
<taxonomicName id="E8854D6EFFC1FFE80BB30E220E359240" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1192,1307,515,541]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80BB30E220E359240" box="[1192,1307,515,541]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
); 2003: 27, 51, 57–58, figs 9, 164; 2012: 93, 115, 171, 192. — 
<treatmentCitation id="AE2410FCFFC1FFE80C9D0E070F4C921D" author="Ratcliffe B. C." box="[902,1122,550,576]" page="12" pageId="85" pageNumber="86" year="2002">
<bibRefCitation id="4B144B1CFFC1FFE80C9D0E070F4C921D" author="Ratcliffe B. C." box="[902,1122,550,576]" pageId="85" pageNumber="86" pagination="107 - 111" refId="ref144706" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1 - 48. Ratcliffe B. C. 2013. Best writing and curatorial practices for describing a new species of beetle: a primer. The Coleopterists Bulletin 67 (2): 107 - 111. https: // doi. org / 10.1649 / 0010 - 065 X- 67.2.107" type="journal article" year="2002">Ratcliffe 2002: 12</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80B8D0E070A68923E" author="Solis A. &amp; Kohlmann B." page="1" pageId="85" pageNumber="86" year="2002">
<bibRefCitation id="4B144B1CFFC1FFE80B8D0E070A68923E" author="Solis A. &amp; Kohlmann B." pageId="85" pageNumber="86" pagination="1 - 68" refId="ref146463" refString="Solis A. &amp; Kohlmann B. 2002. El genero Canthon (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia 10: 1 - 68." type="journal article" year="2002">Solís &amp; Kohlmann 2002: 1</bibRefCitation>
</treatmentCitation>
, 4, 6–8, 53–54, 64, figs 1f, 31; 2012: 3, 21. — 
<treatmentCitation id="AE2410FCFFC1FFE80C9D0E680F77923E" author="Halffter G." box="[902,1113,585,611]" page="43" pageId="85" pageNumber="86" year="2003">
<bibRefCitation id="4B144B1CFFC1FFE80C9D0E680F77923E" author="Halffter G." box="[902,1113,585,611]" pageId="85" pageNumber="86" pagination="21 - 43" refId="ref136039" refString="Halffter G. 2003. Tribu Scarabaeini, subtribu Canthonina. In: Moron M. A. (ed.) Atlas de los escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 21 - 43. Argania Editio, Barcelona." type="book chapter" year="2003">Halffter 2003: 43</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80B960E680E58923E" author="Celi J. &amp; Terneus E. &amp; Torres J. &amp; Ortega M." box="[1165,1398,585,611]" page="42" pageId="85" pageNumber="86" year="2004">
<bibRefCitation id="4B144B1CFFC1FFE80B960E680E58923E" author="Celi J. &amp; Terneus E. &amp; Torres J. &amp; Ortega M." box="[1165,1398,585,611]" pageId="85" pageNumber="86" pagination="37 - 52" refId="ref132879" refString="Celi J., Terneus E., Torres J. &amp; Ortega M. 2004. Dung beetles (Coleoptera: Scarabaeinae) diversity in an altitudinal gradient in the Cutucu Range, Morona Santiago, Ecuadorian Amazon. Lyonia 7 (2): 37 - 52." type="journal article" year="2004">
Celi 
<emphasis id="1DF1EAFFFFC1FFE80BDC0E6B0E2A923E" box="[1223,1284,585,611]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2004: 42
</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80EBE0E4C093792DB" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[421,537,621,647]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80EBE0E4C093792DB" box="[421,537,621,647]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80D4C0E4C08B592DA" author="Noriega-Alvarado J. A." box="[599,923,621,647]" page="40" pageId="85" pageNumber="86" year="2004">
<bibRefCitation id="4B144B1CFFC1FFE80D4C0E4C08B592DA" author="Noriega-Alvarado J. A." box="[599,923,621,647]" pageId="85" pageNumber="86" pagination="37 - 44" refId="ref142973" refString="Noriega-Alvarado J. A. 2004. Preliminary checklist of the scarab community (Coleoptera: Scarabaeidae) at CIEM, Tinigua National Park, Meta-Colombia. Field Studies of Fauna and Flora, La Macarena, Colombia 14: 37 - 44." type="journal article" year="2004">Noriega-Alvarado 2004: 40</bibRefCitation>
</treatmentCitation>
(idem). — 
<treatmentCitation id="AE2410FCFFC1FFE80B3F0E4C0E6B92DA" author="Kohlmann B. &amp; Solis A. &amp; Elle O. &amp; Soto X. &amp; Russo R." box="[1060,1349,621,647]" page="9" pageId="85" pageNumber="86" year="2007">
<bibRefCitation id="4B144B1CFFC1FFE80B3F0E4C0E6B92DA" author="Kohlmann B. &amp; Solis A. &amp; Elle O. &amp; Soto X. &amp; Russo R." box="[1060,1349,621,647]" pageId="85" pageNumber="86" pagination="1 - 34" refId="ref138694" refString="Kohlmann B., Solis A., Elle O., Soto X. &amp; Russo R. 2007. Biodiversity, conservation, and hotspot atlas of Costa Rica: a dung beetle perspective (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 1457: 1 - 34." type="journal article" year="2007">
Kohlmann 
<emphasis id="1DF1EAFFFFC1FFE80BBC0E4C0FCC92DA" box="[1191,1250,621,647]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2007: 9
</bibRefCitation>
</treatmentCitation>
, 28; 2010: 553. — 
<treatmentCitation id="AE2410FCFFC1FFE80E820EB109F492F7" author="Noriega J. A. &amp; Cubillos A. M. &amp; Castaneda C. &amp; Sanchez A. M." box="[409,730,656,682]" page="78" pageId="85" pageNumber="86" year="2008">
<bibRefCitation id="4B144B1CFFC1FFE80E820EB109F492F7" author="Noriega J. A. &amp; Cubillos A. M. &amp; Castaneda C. &amp; Sanchez A. M." box="[409,730,656,682]" pageId="85" pageNumber="86" pagination="75 - 86" refId="ref142833" refString="Noriega J. A., Cubillos A. M., Castaneda C. &amp; Sanchez A. M. 2008. Actividad diaria de colonizacion del recurso alimenticio en un ensamblaje de escarabajos coprofagos (Coleoptera: Scarabaeidae) en la amazonia colombiana. Acta Biologica Colombiana 13 (3): 75 - 86." type="journal article" year="2008">
Noriega 
<emphasis id="1DF1EAFFFFC1FFE80D1B0EB0091392F7" box="[512,573,656,682]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2008: 78–79
</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80CB80EB10F3792F7" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[931,1049,656,682]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80CB80EB10F3792F7" box="[931,1049,656,682]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80B420EB10E5C92F7" author="Carpio C. &amp; Donoso D. A. &amp; Ramon G. &amp; Dangles O." box="[1113,1394,656,682]" page="462" pageId="85" pageNumber="86" year="2009">
<bibRefCitation id="4B144B1CFFC1FFE80B420EB10E5C92F7" author="Carpio C. &amp; Donoso D. A. &amp; Ramon G. &amp; Dangles O." box="[1113,1394,656,682]" pageId="85" pageNumber="86" pagination="455 - 469" refId="ref132669" refString="Carpio C., Donoso D. A., Ramon G. &amp; Dangles O. 2009. Short term response of dung beetle communities to disturbance by road construction in the Ecuadorian Amazon. Annales de la Societe entomologique de France (N. S.) 45 (4): 455 - 469. https: // doi. org / 10.1080 / 00379271.2009.10697629" type="journal article" year="2009">
Carpio 
<emphasis id="1DF1EAFFFFC1FFE80BA90EB00FC092F7" box="[1202,1262,656,682]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2009: 462
</bibRefCitation>
</treatmentCitation>
, 464, 469 (
<emphasis id="1DF1EAFFFFC1FFE80E7E0E920AB09290" box="[357,414,691,717]" italics="true" pageId="85" pageNumber="86">idem</emphasis>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80EC40E92080E9293" author="Medina C. A. &amp; Pulido L. A." box="[479,800,691,718]" page="58" pageId="85" pageNumber="86" year="2009">
<bibRefCitation id="4B144B1CFFC1FFE80EC40E92080E9293" author="Medina C. A. &amp; Pulido L. A." box="[479,800,691,718]" pageId="85" pageNumber="86" pagination="55 - 62" refId="ref141434" refString="Medina C. A. &amp; Pulido L. A. 2009. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de la Orinoquia colombiana. Biota Colombiana 10: 55 - 62." type="journal article" year="2009">Medina &amp; Pulido 2009: 58</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80CF20E950F719290" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1001,1119,692,718]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80CF20E950F719290" box="[1001,1119,692,718]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80B850E920A4D92AD" author="Arango L. &amp; Montes J. M." page="262" pageId="85" pageNumber="86" year="2010">
<bibRefCitation id="4B144B1CFFC1FFE80B850E920A4D92AD" author="Arango L. &amp; Montes J. M." pageId="85" pageNumber="86" pagination="249 - 268" refId="ref130609" refString="Arango L. &amp; Montes J. M. 2010. Caracterizacion entomologica parcial de la cuenca del rio La Miel en el departamento de Caldas (Colombia). Boletin Cientifico, Centro de Museos, Museo de Historia Natural 13 (2): 249 - 268." type="journal article" year="2010">Arango &amp; Montes 2010: 262</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80E8B0EF6081092AC" author="Figueroa L. &amp; Alvarado M." box="[400,830,727,753]" page="210" pageId="85" pageNumber="86" year="2011">
<bibRefCitation id="4B144B1CFFC1FFE80E8B0EF6081092AC" author="Figueroa L. &amp; Alvarado M." box="[400,830,727,753]" pageId="85" pageNumber="86" pagination="209 - 212" refId="ref135346" refString="Figueroa L. &amp; Alvarado M. 2011. Coleopteros coprofagos (Scarabaeidae: Scarabaeinae) de la Reserva Nacional Tambopata, Madre de Dios, Peru. Revista Peruana de Biologia 18 (2): 209 - 212." type="journal article" year="2011">Figueroa &amp; Alvarado 2011: 210–211</bibRefCitation>
</treatmentCitation>
, fig. 1c (error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80B550EF60FEF92AD" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1102,1217,727,753]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80B550EF60FEF92AD" box="[1102,1217,727,753]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80BE70EF60A7D9349" author="Solis C. &amp; Noriega J. A. &amp; Herrera G." page="36" pageId="85" pageNumber="86" year="2011">
<bibRefCitation id="4B144B1CFFC1FFE80BE70EF60A7D9349" author="Solis C. &amp; Noriega J. A. &amp; Herrera G." pageId="85" pageNumber="86" pagination="33 - 41" refId="ref146525" refString="Solis C., Noriega J. A. &amp; Herrera G. 2011. Escarabajos coprofagos (Coleoptera: Scarabaeinae) en tres bosques secos del departamento del Atlatico-Colombia. Boletin del Museo de Entomologia de la Universidad del Valle 12 (1): 33 - 41." type="journal article" year="2011">
Solís 
<emphasis id="1DF1EAFFFFC1FFE80A260EF60E5992AC" box="[1341,1399,727,753]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2011: 36
</bibRefCitation>
</treatmentCitation>
, 38. — 
<treatmentCitation id="AE2410FCFFC1FFE80EB60EDB09589349" author="Krajcik M." box="[429,630,762,788]" page="63" pageId="85" pageNumber="86" year="2012">
<bibRefCitation id="4B144B1CFFC1FFE80EB60EDB09589349" author="Krajcik M." box="[429,630,762,788]" pageId="85" pageNumber="86" refId="ref138852" refString="Krajcik M. 2012. Animma. X. Supplement 5. Checklist of the World Scarabaeoidea. Privately published, Plzen." type="book" year="2012">Krajcik 2012: 63</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80DBF0EDB084C9349" author="Noriega J. A." box="[676,866,762,788]" page="4" pageId="85" pageNumber="86" year="2012">
<bibRefCitation id="4B144B1CFFC1FFE80DBF0EDB084C9349" author="Noriega J. A." box="[676,866,762,788]" pageId="85" pageNumber="86" pagination="1 - 6" refId="ref142684" refString="Noriega J. A. 2012. Dung beetles (Coleoptera: Scarabaeinae) attracted to Lagothrix lagotricha (Humboldt) and Alouatta seniculus (Linnaeus) (Primates: Atelidae) dung in a Colombian Amazon Forest. Psyche Article ID 437589: 1 - 6." type="journal article" year="2012">Noriega 2012: 4</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80B3A0EDB0FBA9349" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1057,1172,762,788]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80B3A0EDB0FBA9349" box="[1057,1172,762,788]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80BCB0EDB0A4E9365" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Jimenez-Ferbans L. &amp; Reyes-Castillo P." page="195" pageId="85" pageNumber="86" year="2015">
<bibRefCitation id="4B144B1CFFC1FFE80BCB0EDB0A4E9365" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Jimenez-Ferbans L. &amp; Reyes-Castillo P." pageId="85" pageNumber="86" pagination="186 - 207" refId="ref144842" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez-Ferbans L. &amp; Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Scarabaeoidea. Journal of the Kansas Entomological Society 88 (2): 186 - 207. https: // doi. org / 10.2317 / kent- 88 - 02 - 186 - 207.1" type="journal article" year="2015">
Ratcliffe 
<emphasis id="1DF1EAFFFFC1FFE80A270EDA0E599349" box="[1340,1399,762,788]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2015: 195
</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80D390F3F09B9936A" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[546,663,798,824]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80D390F3F09B9936A" box="[546,663,798,824]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60F6008F193FC" blockId="85.[189,1399,267,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60F6008F193FC" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA60F600AE69306" box="[189,456,833,859]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctiale">
<emphasis id="1DF1EAFFFFC1FFE80FA60F600AE69306" box="[189,456,833,859]" italics="true" pageId="85" pageNumber="86">Canthon aequinoctiale</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80EF90F6109DB9306" author="Blackwelder R. E." box="[482,757,832,859]" page="198" pageId="85" pageNumber="86" year="1944">
<bibRefCitation id="4B144B1CFFC1FFE80EF90F6109DB9306" author="Blackwelder R. E." box="[482,757,832,859]" pageId="85" pageNumber="86" pagination="189 - 341" refId="ref131382" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 198</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="AE2410FCFFC1FFE80C1B0F6008759306" author="Blackwelder R. E." box="[768,859,833,859]" page="2" pageId="85" pageNumber="86" year="1973">
<bibRefCitation id="4B144B1CFFC1FFE80C1B0F6008759306" author="Blackwelder R. E." box="[768,859,833,859]" pageId="85" pageNumber="86" refId="ref131432" refString="Blackwelder R. E. 1973. Checklist of the Scarabaeidae of Canada, United States, Mexico, Central America and the West Indies. Family no. 30, complete (Red Version). Biological Research Institute of America, Latham, New York." type="book" year="1973">1973: 2</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80C930F610F589306" author="Gacharna G. C." box="[904,1142,832,859]" page="221" pageId="85" pageNumber="86" year="1951">
<bibRefCitation id="4B144B1CFFC1FFE80C930F610F589306" author="Gacharna G. C." box="[904,1142,832,859]" pageId="85" pageNumber="86" pagination="221 - 229" refId="ref135535" refString="Gacharna G. C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 8: 221 - 229." type="journal article" year="1951">Gacharná 1951: 221</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80BBF0F610A769323" author="Howden H. F. &amp; Young O. P." page="20" pageId="85" pageNumber="86" year="1981">
<bibRefCitation id="4B144B1CFFC1FFE80BBF0F610A769323" author="Howden H. F. &amp; Young O. P." pageId="85" pageNumber="86" pagination="1 - 204" refId="ref137827" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young 1981: 20</bibRefCitation>
</treatmentCitation>
, 27–28, 153, 167, figs 36–37. — 
<treatmentCitation id="AE2410FCFFC1FFE80DEE0F450F8E9322" author="Vaz-de-Mello F. Z. &amp; Louzada J. N. C." box="[757,1184,868,895]" page="56" pageId="85" pageNumber="86" year="1997">
<bibRefCitation id="4B144B1CFFC1FFE80DEE0F450F8E9322" author="Vaz-de-Mello F. Z. &amp; Louzada J. N. C." box="[757,1184,868,895]" pageId="85" pageNumber="86" pagination="55 - 61" refId="ref147471" refString="Vaz-de-Mello F. Z. &amp; Louzada J. N. C. 1997. Consideracoes sobre forrageio arboreo por Scarabaeidae (Coleoptera, Scarabaeoidea), e dados sobre sua ocorrencia em floresta tropical do Brasil. Acta Zoologica Mexicana (n. s.) 72: 55 - 61." type="journal article" year="1997">Vaz-de-Mello &amp; Louzada 1997: 56</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80BCE0F450AB593FF" author="Vaz-de-Mello F. Z." page="449" pageId="85" pageNumber="86" year="1999">
<bibRefCitation id="4B144B1CFFC1FFE80BCE0F450AB593FF" author="Vaz-de-Mello F. Z." pageId="85" pageNumber="86" pagination="447 - 453" refId="ref147027" refString="Vaz-de-Mello F. Z. 1999. Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) de um fragmento de floresta amazonica no estado do Acre, Brasil. 1. Taxocenose. Anais da Sociedade Entomologica do Brasil 28 (3): 447 - 453." type="journal article" year="1999">Vaz-de-Mello 1999: 449–450</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80D460FA909FF93FC" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[605,721,904,930]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80D460FA909FF93FC" box="[605,721,904,930]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
). — 
<treatmentCitation id="AE2410FCFFC1FFE80C160FA908F293FC" author="Young O. P." box="[781,988,903,930]" page="322" pageId="85" pageNumber="86" year="2009">
<bibRefCitation id="4B144B1CFFC1FFE80C160FA908F293FC" author="Young O. P." box="[781,988,903,930]" pageId="85" pageNumber="86" pagination="319 - 324" refId="ref148410" refString="Young O. P. 2009. Observations on burrowing behavior by Panamanian Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 63 (3): 319 - 324. https: // doi. org / 10.1649 / 1164.1" type="journal article" year="2009">Young 2009: 322</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60F8A0FA89398" blockId="85.[189,1399,267,1283]" box="[189,1158,938,965]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60F8A0FA89398" box="[189,1158,938,965]" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA60F8A0ADB9398" ID-CoL="QPB6" authorityName="Harold" authorityYear="1868" box="[189,501,938,965]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80FA60F8A0A0C9398" box="[189,290,939,965]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
cf. 
<emphasis id="1DF1EAFFFFC1FFE80E540F8A0ADB9398" box="[335,501,939,965]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80D090F8B081D9398" author="Noriega J. A. &amp; Solis C. &amp; Escobar F. &amp; Realpe E." box="[530,819,938,965]" page="81" pageId="85" pageNumber="86" year="2007">
<bibRefCitation id="4B144B1CFFC1FFE80D090F8B081D9398" author="Noriega J. A. &amp; Solis C. &amp; Escobar F. &amp; Realpe E." box="[530,819,938,965]" pageId="85" pageNumber="86" pagination="77 - 86" refId="ref142784" refString="Noriega J. A., Solis C., Escobar F. &amp; Realpe E. 2007 b. Escarabajos coprofagos (Coleoptera: Scarabaeidae) de la provincia de la Sierra Nevada de Santa Marta. Biota Colombiana 8 (1): 77 - 86." type="journal article" year="2007">
Noriega 
<emphasis id="1DF1EAFFFFC1FFE80D630F8A099D9398" box="[632,691,939,965]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2007b: 81
</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80C7F0F8A0FAC9398" author="Martinez N. J. &amp; Canas L. M. &amp; Rangel J. L. &amp; Barraza J. M. &amp; Montes J. M. &amp; Blanco O. R." box="[868,1154,939,965]" page="24" pageId="85" pageNumber="86" year="2010">
<bibRefCitation id="4B144B1CFFC1FFE80C7F0F8A0FAC9398" author="Martinez N. J. &amp; Canas L. M. &amp; Rangel J. L. &amp; Barraza J. M. &amp; Montes J. M. &amp; Blanco O. R." box="[868,1154,939,965]" pageId="85" pageNumber="86" pagination="21 - 30" refId="ref140436" refString="Martinez N. J., Canas L. M., Rangel J. L., Barraza J. M., Montes J. M. &amp; Blanco O. R. 2010. Coleopteros coprofagos (Scarabaeidae: Scarabaeinae) en un fragmento de bosque seco tropical en el departamento del Atlantico, Colombia. Boletin del Museo de Entomologia de la Universidad del Valle 11 (1): 21 - 30." type="journal article" year="2010">
Martínez 
<emphasis id="1DF1EAFFFFC1FFE80CCE0F8A0F3E9398" box="[981,1040,939,965]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2010: 24
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60FEF09559451" blockId="85.[189,1399,267,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60FEF09559451" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA60FEF097893B5" ID-CoL="QPB6" authorityName="Harold" authorityYear="1868" box="[189,598,974,1000]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis" subGenus="Canthon">
<emphasis id="1DF1EAFFFFC1FFE80FA60FEF0A0C93B5" box="[189,290,974,1000]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
(
<emphasis id="1DF1EAFFFFC1FFE80E2D0FEF0AB593B5" box="[310,411,974,1000]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
) 
<emphasis id="1DF1EAFFFFC1FFE80EAB0FEF097893B5" box="[432,598,974,1000]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80D620FEF08CD93B5" author="Halffter G. &amp; Martinez A." box="[633,995,974,1000]" page="62" pageId="85" pageNumber="86" year="1977">
<bibRefCitation id="4B144B1CFFC1FFE80D620FEF08CD93B5" author="Halffter G. &amp; Martinez A." box="[633,995,974,1000]" pageId="85" pageNumber="86" pagination="29 - 107" refId="ref136250" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 62</bibRefCitation>
</treatmentCitation>
, 90–91. — 
<treatmentCitation id="AE2410FCFFC1FFE80B6E0FEF0A1D9451" author="Noriega J. A. &amp; Realpe E. &amp; Fagua G." page="54" pageId="85" pageNumber="86" year="2007">
<bibRefCitation id="4B144B1CFFC1FFE80B6E0FEF0A1D9451" author="Noriega J. A. &amp; Realpe E. &amp; Fagua G." pageId="85" pageNumber="86" pagination="51 - 63" refId="ref142732" refString="Noriega J. A., Realpe E. &amp; Fagua G. 2007 a. Diversidad de escarabajos coprofagos (Coleoptera: Scarabaeidae) en un bosque de galeria con tres estadios de alteracion. Revista de la Facultad de Ciencias, Edicion Especial I 12: 51 - 63." type="journal article" year="2007">
Noriega 
<emphasis id="1DF1EAFFFFC1FFE80BC50FEE0E3293B5" box="[1246,1308,974,1000]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
2007a: 54–55
</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80EEE0FD309449456" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[501,618,1010,1036]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80EEE0FD309449456" box="[501,618,1010,1036]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA608340FBE9472" blockId="85.[189,1399,267,1283]" box="[189,1168,1044,1071]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA608340FBE9472" box="[189,1168,1044,1071]" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA6083409679472" box="[189,585,1045,1071]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctiale" subGenus="Canthon">
<emphasis id="1DF1EAFFFFC1FFE80FA608340A0C9472" box="[189,290,1045,1071]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
(
<emphasis id="1DF1EAFFFFC1FFE80E2808340AB69472" box="[307,408,1045,1071]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
) 
<emphasis id="1DF1EAFFFFC1FFE80EB2083409679472" box="[425,585,1045,1071]" italics="true" pageId="85" pageNumber="86">aequinoctiale</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80D7D083508679472" box="[614,841,1044,1071]" pageId="85" pageNumber="86">
<bibRefCitation id="4B144B1CFFC1FFE80D7D083508679472" author="Vaz-de-Mello F. Z." box="[614,841,1044,1071]" pageId="85" pageNumber="86" pagination="183 - 195" refId="ref147076" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un proyecto CYTED para el inventario y estimacion de la diversidad entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. Monografias Tercer Milenio 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="journal article" year="2000">Vaz-de-Mello 2000</bibRefCitation>
</treatmentCitation>
(error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80B1008340F519473" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1035,1151,1045,1071]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80B1008340F519473" box="[1035,1151,1045,1071]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA608190804942B" blockId="85.[189,1399,267,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60819080B942B" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA608190962940F" box="[189,588,1080,1106]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinotialis" subGenus="Canthon">
<emphasis id="1DF1EAFFFFC1FFE80FA608190A0C940F" box="[189,290,1080,1106]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
(
<emphasis id="1DF1EAFFFFC1FFE80E2308190AB3940F" box="[312,413,1080,1106]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
) 
<emphasis id="1DF1EAFFFFC1FFE80EA808190962940F" box="[435,588,1080,1106]" italics="true" pageId="85" pageNumber="86">aequinotialis</emphasis>
</taxonomicName>
[sic] – 
<bibRefCitation id="4B144B1CFFC1FFE80DAF081908FD940F" author="Noriega-Alvarado J. A." box="[692,979,1080,1106]" pageId="85" pageNumber="86" pagination="83 - 89" refId="ref143017" refString="Noriega-Alvarado J. A. 2009. Analisis de la diversidad de escarabajos coprofagos (Coleoptera: Scarabaeidae) en el departamento de Cundinamarca. Revista de Tecnologia 8 (1): 83 - 89." type="journal article" year="2009">Noriega-Alvarado 2009</bibRefCitation>
. — 
<treatmentCitation id="AE2410FCFFC1FFE80B1608190FCA940F" box="[1037,1252,1080,1106]" pageId="85" pageNumber="86">
<bibRefCitation id="4B144B1CFFC1FFE80B1608190FCA940F" author="Culot L. &amp; Mann D. J. &amp; Lazo F. J. J. M. &amp; Huynen M. - C. &amp; Heymann E. W." box="[1037,1252,1080,1106]" pageId="85" pageNumber="86" pagination="84 - 92" refId="ref133602" refString="Culot L., Mann D. J., Lazo F. J. J. M., Huynen M. - C. &amp; Heymann E. W. 2011. Tamarins and dung beetles: an efficient diplochorous dispersal system in the Peruvian Amazonia. Biotropica 43 (1): 84 - 92. https: // doi. org / 10.1111 / j. 1744 - 7429.2010.00655. x" type="journal article" year="2011">
Culot 
<emphasis id="1DF1EAFFFFC1FFE80B4108180FBD940F" box="[1114,1171,1080,1106]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
. 2011
</bibRefCitation>
</treatmentCitation>
: Supporting information, table S1 (error: refers to 
<taxonomicName id="E8854D6EFFC1FFE80DBE087D08379428" ID-CoL="53N57" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[677,793,1116,1142]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC1FFE80DBE087D08379428" box="[677,793,1116,1142]" italics="true" pageId="85" pageNumber="86">S. proseni</emphasis>
</taxonomicName>
)
</treatmentCitationGroup>
.
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA6085E089094C4" blockId="85.[189,1399,267,1283]" box="[189,958,1150,1177]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA6085E089094C4" box="[189,958,1150,1177]" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA6085E099694C4" ID-CoL="QPB6" authorityName="Harold" authorityYear="1868" box="[189,696,1151,1177]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis" subGenus="Glaphyrocanthon">
<emphasis id="1DF1EAFFFFC1FFE80FA6085E0A0C94C4" box="[189,290,1151,1177]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
(
<emphasis id="1DF1EAFFFFC1FFE80E28085E092F94C4" box="[307,513,1151,1177]" italics="true" pageId="85" pageNumber="86">Glaphyrocanthon</emphasis>
) 
<emphasis id="1DF1EAFFFFC1FFE80D09085E099694C4" box="[530,696,1151,1177]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80DCD085F089494C4" author="Howden H. F." box="[726,954,1150,1177]" page="730" pageId="85" pageNumber="86" year="1966">
<bibRefCitation id="4B144B1CFFC1FFE80DCD085F089494C4" author="Howden H. F." box="[726,954,1150,1177]" pageId="85" pageNumber="86" pagination="725 - 741" refId="ref137718" refString="Howden H. F. 1966. Notes on Canthonini of the &quot; Biologia Centrali-Americana &quot; and Descriptions of new species (Coleoptera: Scarabaeidae). The Canadian Entomologist 98 (7): 725 - 741." type="journal article" year="1966">Howden 1966: 730</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60883088494BD" blockId="85.[189,1399,267,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA60883088494BD" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA60883080F94E1" box="[189,801,1186,1212]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis" subGenus="Glaphyrocanthon">
<emphasis id="1DF1EAFFFFC1FFE80FA608830AA594E1" box="[189,395,1186,1212]" italics="true" pageId="85" pageNumber="86">Glaphyrocanthon</emphasis>
(
<emphasis id="1DF1EAFFFFC1FFE80E870883094494E1" box="[412,618,1186,1212]" italics="true" pageId="85" pageNumber="86">Glaphyrocanthon</emphasis>
) 
<emphasis id="1DF1EAFFFFC1FFE80D600883080F94E1" box="[635,801,1186,1212]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="AE2410FCFFC1FFE80C2408830F8F94E1" author="Martinez A. &amp; Pereira F. S." box="[831,1185,1186,1213]" page="126" pageId="85" pageNumber="86" year="1956">
<bibRefCitation id="4B144B1CFFC1FFE80C2408830F8F94E1" author="Martinez A. &amp; Pereira F. S." box="[831,1185,1186,1213]" pageId="85" pageNumber="86" pagination="363 - 388" refId="ref140293" refString="Martinez A. &amp; Pereira F. S. 1956. Dois generos novos de Canthonini americanos (Col. Scarabaeoidea, Scarabaeidae). Papeis Avulsos do Departamento de Zoologia 12 (19): 363 - 388." type="journal article" year="1956">Pereira &amp; Martínez 1956: 126</bibRefCitation>
</treatmentCitation>
, 128. — 
<treatmentCitation id="AE2410FCFFC1FFE80A1608830AA89482" author="Martinez A. &amp; Halffter G. &amp; Halffter V." page="3" pageId="85" pageNumber="86" year="1964">
<bibRefCitation id="4B144B1CFFC1FFE80A1608830AA89482" author="Martinez A. &amp; Halffter G. &amp; Halffter V." pageId="85" pageNumber="86" pagination="1 - 42" refId="ref140396" refString="Martinez A., Halffter G. &amp; Halffter V. 1964. Notas sobre el genero Glaphyrocanthon (Coleopt., Scarab., Canthonina). Acta Zoologica Mexicana 7 (3): 1 - 42." type="journal article" year="1964">
Martínez 
<emphasis id="1DF1EAFFFFC1FFE80FF108E70A0B9482" box="[234,293,1221,1247]" italics="true" pageId="85" pageNumber="86">et al.</emphasis>
1964: 3
</bibRefCitation>
</treatmentCitation>
, 5, 8–10, 13. — 
<treatmentCitation id="AE2410FCFFC1FFE80D5008E4088B94BD" author="Vulcano M. A. &amp; Pereira F. S." box="[587,933,1221,1248]" page="561" pageId="85" pageNumber="86" year="1967">
<bibRefCitation id="4B144B1CFFC1FFE80D5008E4088B94BD" author="Vulcano M. A. &amp; Pereira F. S." box="[587,933,1221,1248]" pageId="85" pageNumber="86" pagination="533 - 603" refId="ref147845" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5 (Zoologia): 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 561</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA608C90FE5955F" blockId="85.[189,1399,267,1283]" box="[189,1227,1256,1283]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="0F9511C3FFC1FFE80FA608C90FE5955F" box="[189,1227,1256,1283]" pageId="85" pageNumber="86">
<taxonomicName id="E8854D6EFFC1FFE80FA608C9083A955F" box="[189,788,1256,1283]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinotialis" subGenus="Glaphyrocanthon">
<emphasis id="1DF1EAFFFFC1FFE80FA608C90AA5955F" box="[189,395,1256,1282]" italics="true" pageId="85" pageNumber="86">Glaphyrocanthon</emphasis>
(
<emphasis id="1DF1EAFFFFC1FFE80E8708C90944955F" box="[412,618,1256,1282]" italics="true" pageId="85" pageNumber="86">Glaphyrocanthon</emphasis>
) 
<emphasis id="1DF1EAFFFFC1FFE80D6008C9083A955F" box="[635,788,1256,1282]" italics="true" pageId="85" pageNumber="86">aequinotialis</emphasis>
</taxonomicName>
[sic] – 
<treatmentCitation id="AE2410FCFFC1FFE80C7708C90FE9955F" author="Vulcano M. A. &amp; Pereira F. S." box="[876,1223,1256,1283]" page="661" pageId="85" pageNumber="86" year="1964">
<bibRefCitation id="4B144B1CFFC1FFE80C7708C90FE9955F" author="Vulcano M. A. &amp; Pereira F. S." box="[876,1223,1256,1283]" pageId="85" pageNumber="86" pagination="570 - 685" refId="ref147746" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey 15 (2): 570 - 685. Available from https: // biodiversitylibrary. org / page / 45986646 [accessed 30 May 2018]." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 661</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="679F6566FFC1FFE80FA6090B09A895CA" pageId="85" pageNumber="86" type="etymology">
<paragraph id="2F3A36EDFFC1FFE80FA6090B0A6A9519" blockId="85.[189,324,1322,1348]" box="[189,324,1322,1348]" pageId="85" pageNumber="86">
<heading id="74728181FFC1FFE80FA6090B0A6A9519" bold="true" box="[189,324,1322,1348]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="1DF1EAFFFFC1FFE80FA6090B0A6A9519" bold="true" box="[189,324,1322,1348]" pageId="85" pageNumber="86">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA6097809A895CA" blockId="85.[189,1399,1369,1431]" pageId="85" pageNumber="86">
From the Latin ‘ 
<taxonomicName id="E8854D6EFFC1FFE80E900978091F952E" box="[395,561,1369,1395]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80E900978091F952E" box="[395,561,1369,1395]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
’, meaning ‘equinox’ or ‘equinoctial’. Probable reference to the 
<typeStatus id="F03E884FFFC1FFE80A5F09780E59952E" box="[1348,1399,1369,1395]" pageId="85" pageNumber="86">type</typeStatus>
locality, region crossed by the Equator.
</paragraph>
</subSubSection>
<subSubSection id="679F6566FFC1FFE40FA6099C0E0F976F" lastPageId="89" lastPageNumber="90" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="2F3A36EDFFC1FFE80FA6099C0A84958A" blockId="85.[189,426,1469,1495]" box="[189,426,1469,1495]" pageId="85" pageNumber="86">
<heading id="74728181FFC1FFE80FA6099C0A84958A" bold="true" box="[189,426,1469,1495]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
<emphasis id="1DF1EAFFFFC1FFE80FA6099C0A84958A" bold="true" box="[189,426,1469,1495]" pageId="85" pageNumber="86">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FF109CD0916965A" blockId="85.[189,1398,1516,1649]" box="[234,568,1516,1543]" pageId="85" pageNumber="86">
<materialsCitation id="9FED3CB0FFC1FFE80FF109CD0916965A" ID-GBIF-Occurrence="2614776319" box="[234,568,1516,1543]" pageId="85" pageNumber="86" typeStatus="lectotype">
<emphasis id="1DF1EAFFFFC1FFE80FF109CD0A48965B" bold="true" box="[234,358,1516,1542]" pageId="85" pageNumber="86">
<typeStatus id="F03E884FFFC1FFE80FF109CD0A48965B" box="[234,358,1516,1542]" pageId="85" pageNumber="86" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</materialsCitation>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60A3108C0962D" blockId="85.[189,1398,1516,1649]" pageId="85" pageNumber="86">
<collectingCountry id="5792767DFFC1FFE80FA60A310A709677" box="[189,350,1552,1578]" name="Colombia" pageId="85" pageNumber="86">COLOMBIA</collectingCountry>
: ♂, type locality cited by 
<bibRefCitation id="4B144B1CFFC1FFE80D970A3108119677" author="Harold E." box="[652,831,1551,1578]" pageId="85" pageNumber="86" pagination="1 - 144" refId="ref136501" refString="Harold E. 1868 a. Monographie der Gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120104" type="journal article" year="1868">Harold (1868a)</bibRefCitation>
as “ 
<emphasis id="1DF1EAFFFFC1FFE80C6F0A310F889677" box="[884,1190,1552,1578]" italics="true" pageId="85" pageNumber="86">Columbien, Neu-Granada</emphasis>
”, (“ 
<taxonomicName id="E8854D6EFFC1FFE80BC30A310E589677" box="[1240,1398,1552,1578]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80BC30A310E589677" box="[1240,1398,1552,1578]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFFC1FFE80FD40A150A389610" box="[207,278,1588,1613]" italics="true" pageId="85" pageNumber="86">t. Har</emphasis>
”, “ 
<emphasis id="1DF1EAFFFFC1FFE80E590A150ACB9610" box="[322,485,1587,1613]" italics="true" pageId="85" pageNumber="86">Nov. Grenade</emphasis>
”, “Ex-Musaeo / E. Harold”, “ 
<typeStatus id="F03E884FFFC1FFE80C5C0A1208C59610" box="[839,1003,1587,1613]" pageId="85" pageNumber="86" type="holotype">HOLOTYPE</typeStatus>
”, “ 
<emphasis id="1DF1EAFFFFC1FFE80B0B0A120F7F9610" box="[1040,1105,1587,1613]" italics="true" pageId="85" pageNumber="86">TIPO</emphasis>
/ 
<taxonomicName id="E8854D6EFFC1FFE80B770A120FFF9610" baseAuthorityName="Halffter &amp; Martinez" baseAuthorityYear="1977" box="[1132,1233,1587,1613]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFFC1FFE80B770A120FFF9610" box="[1132,1233,1587,1613]" italics="true" pageId="85" pageNumber="86">Canthon</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFFC1FFE80BF70A120E4B9610" box="[1260,1381,1587,1613]" italics="true" pageId="85" pageNumber="86">
(
<taxonomicName id="E8854D6EFFC1FFE80BEE0A120E729610" baseAuthorityName="Halffter &amp; Martinez" baseAuthorityYear="1977" box="[1269,1372,1587,1613]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">Canthon</taxonomicName>
)
</emphasis>
/ 
<taxonomicName id="E8854D6EFFC1FFE80FA60A770A4D962D" box="[189,355,1622,1648]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80FA60A770A4D962D" box="[189,355,1622,1648]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFFC1FFE80E610A770AE0962D" box="[378,462,1622,1648]" italics="true" pageId="85" pageNumber="86">Harold</emphasis>
/ 
<emphasis id="1DF1EAFFFFC1FFE80EFF0A7609F9962D" box="[484,727,1622,1649]" italics="true" pageId="85" pageNumber="86">
G. H. y A. 
<taxonomicName id="E8854D6EFFC1FFE80D7A0A760981962D" box="[609,687,1622,1648]" class="Insecta" family="Geotrupidae" genus="Mycotrupes" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="det">M. det</taxonomicName>
. 76
</emphasis>
”) (
<collectionCode id="4994AE28FFC1FFE80DE50A770875962D" box="[766,859,1622,1648]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="85" pageNumber="86">MNHN</collectionCode>
) (
<figureCitation id="B7BE2A68FFC1FFE80C680A7708CF962D" box="[883,993,1622,1648]" captionStart="Fig" captionStartId="88.[189,232,1929,1955]" captionTargetBox="[192,1399,370,1881]" captionTargetId="figure@88.[189,1399,364,1885]" captionTargetPageId="88" captionText="Fig. 29. Type material of the aequinoctialis subgroup. A–B. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov., type series. A. Lectotype and its labels. B. Paralectotype and its labels. C–E. S. proseni (Martínez, 1949) stat. et comb. nov., type series. C. Holotype and its labels. D. Paratype 1. E. Paratype 2." figureDoi="http://doi.org/10.5281/zenodo.3824837" httpUri="https://zenodo.org/record/3824837/files/figure.png" pageId="85" pageNumber="86">Fig. 29A</figureCitation>
).
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FF10AB90AB996EF" blockId="85.[189,1398,1688,1820]" box="[234,407,1688,1714]" pageId="85" pageNumber="86">
<emphasis id="1DF1EAFFFFC1FFE80FF10AB90AB996EF" bold="true" box="[234,407,1688,1714]" pageId="85" pageNumber="86">
<typeStatus id="F03E884FFFC1FFE80FF10AB90AB996EF" box="[234,407,1688,1714]" pageId="85" pageNumber="86" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FA60A9A0A809741" blockId="85.[189,1398,1688,1820]" pageId="85" pageNumber="86">
<collectingCountry id="5792767DFFC1FFE80FA60A9A0A709688" box="[189,350,1723,1749]" name="Colombia" pageId="85" pageNumber="86">COLOMBIA</collectingCountry>
: 
<specimenCount id="3983FD64FFC1FFE80E720A9D0A51968B" box="[361,383,1724,1750]" pageId="85" pageNumber="86" type="female">♀</specimenCount>
, (“26381”, “ 
<emphasis id="1DF1EAFFFFC1FFE80D0C0A9D09D7968B" box="[535,761,1723,1750]" italics="true" pageId="85" pageNumber="86">Nov Granad. Gom.</emphasis>
”, “Type”, “ 
<taxonomicName id="E8854D6EFFC1FFE80C9E0A9A0F059688" box="[901,1067,1723,1749]" class="Insecta" family="Scarabaeidae" genus="Glaphyrocanthon" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC1FFE80C9E0A9A0F059688" box="[901,1067,1723,1749]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFFC1FFE80B590A9A0FB89688" box="[1090,1174,1723,1749]" italics="true" pageId="85" pageNumber="86">Harold</emphasis>
* / 
<emphasis id="1DF1EAFFFFC1FFE80BA70A9D0E4C9688" box="[1212,1378,1723,1749]" italics="true" pageId="85" pageNumber="86">Nov. Granada</emphasis>
”, “ 
<typeStatus id="F03E884FFFC1FFE80FDD0AFE0A9296A4" box="[198,444,1759,1785]" pageId="85" pageNumber="86" type="paralectotype">PARALECTOTYPE</typeStatus>
/ 
<taxonomicName id="E8854D6EFFC1FFE80ECD0AFF09C796A5" authorityName="Harold" authorityYear="1868" box="[470,745,1758,1784]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="122" phylum="Arthropoda" rank="species" species="aequinoctialis">
Canthon 
<emphasis id="1DF1EAFFFFC1FFE80D580AFF09C796A5" box="[579,745,1758,1784]" italics="true" pageId="85" pageNumber="86">aequinoctialis</emphasis>
</taxonomicName>
/ Harold, 1868 / des. Cupello &amp; Vaz-de-Mello, 2015”) (
<collectionCode id="4994AE28FFC1FFE80FDE0B230A339741" box="[197,285,1794,1820]" pageId="85" pageNumber="86">ZMHB</collectionCode>
) (
<figureCitation id="B7BE2A68FFC1FFE80E2E0B230A8F9741" box="[309,417,1794,1820]" captionStart="Fig" captionStartId="88.[189,232,1929,1955]" captionTargetBox="[192,1399,370,1881]" captionTargetId="figure@88.[189,1399,364,1885]" captionTargetPageId="88" captionText="Fig. 29. Type material of the aequinoctialis subgroup. A–B. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov., type series. A. Lectotype and its labels. B. Paralectotype and its labels. C–E. S. proseni (Martínez, 1949) stat. et comb. nov., type series. C. Holotype and its labels. D. Paratype 1. E. Paratype 2." figureDoi="http://doi.org/10.5281/zenodo.3824837" httpUri="https://zenodo.org/record/3824837/files/figure.png" pageId="85" pageNumber="86">Fig. 29B</figureCitation>
).
</paragraph>
<paragraph id="2F3A36EDFFC1FFE80FF10B6208F19703" blockId="85.[189,1399,1858,2027]" box="[234,991,1858,1886]" pageId="85" pageNumber="86">
<emphasis id="1DF1EAFFFFC1FFE80FF10B620ACF9700" bold="true" box="[234,481,1859,1885]" pageId="85" pageNumber="86">Additional material</emphasis>
(282 ♂♂, 
<specimenCount id="3983FD64FFC1FFE80D7B0B6209EF9703" box="[608,705,1859,1886]" pageId="85" pageNumber="86" type="female">210 ♀♀</specimenCount>
, 81 unsexed specimens)
</paragraph>
<paragraph id="2F3A36EDFFC1FFEA0FA60B460F4D93AD" blockId="85.[189,1399,1858,2027]" lastBlockId="87.[189,1399,267,1008]" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86">
<collectingCountry id="5792767DFFC1FFE80FA60B460A7097DC" box="[189,350,1895,1921]" name="Colombia" pageId="85" pageNumber="86">COLOMBIA</collectingCountry>
: 
<specimenCount id="3983FD64FFC1FFE80E770B460AB297DC" box="[364,412,1895,1921]" pageId="85" pageNumber="86" type="female">1 ♀</specimenCount>
, “ 
<emphasis id="1DF1EAFFFFC1FFE80EA00B46092F97DC" box="[443,513,1895,1921]" italics="true" pageId="85" pageNumber="86">Lebas</emphasis>
” (collector or locality?) (
<collectionCode id="4994AE28FFC1FFE80C220B4608AE97DC" box="[825,896,1895,1921]" pageId="85" pageNumber="86">ISNB</collectionCode>
, Gillet collection); 1 ♂, illegible locality, 1921 (
<collectionCode id="4994AE28FFC1FFE80E130BAB0A7597F9" box="[264,347,1930,1956]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="85" pageNumber="86" type="Museum">MZSP</collectionCode>
); 1 ♂, “New Granada”, R. Bunch leg. (
<collectionCode id="4994AE28FFC1FFE80C340BAB08A297F9" box="[815,908,1930,1956]" pageId="85" pageNumber="86">BMNH</collectionCode>
, E.Y. Western coll.). – 
<emphasis id="1DF1EAFFFFC1FFE80B850BAB0E3297F9" bold="true" box="[1182,1308,1930,1956]" pageId="85" pageNumber="86">
<collectingRegion id="ED41F80FFFC1FFE80B850BAB0E3297F9" box="[1182,1308,1930,1956]" country="Colombia" name="Antioquia" pageId="85" pageNumber="86">Antioquia</collectingRegion>
</emphasis>
: 2 ♂♂, San Luis, Sonsón, Reserva Natural Cañon del Río Claro, 
<quantity id="E87D9B08FFC1FFE80C6B0B8C08E39795" box="[880,973,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="85" pageNumber="86" unit="m" value="1440.0">1440 m</quantity>
, 
<date id="5B3B102DFFC1FFE80CC00B8C0F5E979A" box="[987,1136,1965,1992]" pageId="85" pageNumber="86" value="1994-03-02">2 Mar. 1994</date>
, Harold M. Parra leg. (
<collectionCode id="4994AE28FFC1FFE80FDE0BF00A3397B6" box="[197,285,2001,2027]" pageId="85" pageNumber="86">CEMT</collectionCode>
); 2 ♂♂, San Luis, Sonsón, Reserva Natural Cañon del Río Claro, 
<quantity id="E87D9B08FFC1FFE80B340BF00FA597B6" box="[1071,1163,2001,2027]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="85" pageNumber="86" unit="m" value="1440.0">1440 m</quantity>
, 
<date id="5B3B102DFFC1FFE80B830BF00E0597B6" box="[1176,1323,2001,2027]" pageId="85" pageNumber="86" value="1994-03-03">3 Mar. 1994</date>
, H.M. Parra leg. (
<collectionCode id="4994AE28FFC3FFEA0E5F0D2A0AB29178" box="[324,412,267,293]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0EAF0D2A0ACA917B" box="[436,484,267,294]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, San Luis, Sonsón, Reserva Natural Cañon del Río Claro, 
<quantity id="E87D9B08FFC3FFEA0BAC0D2A0E38917B" box="[1207,1302,267,294]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="87" pageNumber="88" unit="m" value="1440.0">1440 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0A3D0D2A0BD49114" pageId="87" pageNumber="88" value="1994-03-03">3 Mar. 1994</date>
, H.M. Parra leg. (
<collectionCode id="4994AE28FFC3FFEA0ECF0D0E091E9114" box="[468,560,303,329]" country="Brazil" httpUri="http://grbio.org/cool/wyk1-wacz" name="Porto Alegre, Museu de Ciencias Naturais da Fundacao Zoo-Botanica do Rio Grande do Sul" pageId="87" pageNumber="88">MCNZ</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0D450D0E09899114" bold="true" box="[606,679,303,329]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0D450D0E09899114" box="[606,679,303,329]" country="Colombia" name="Cesar" pageId="87" pageNumber="88">Cesar</collectingRegion>
</emphasis>
: 1 ♂, 
<specimenCount id="3983FD64FFC3FFEA0DE30D0E08089117" box="[760,806,303,330]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Chimichagua, Finca Señor Reyes, 
<geoCoordinate id="4AB1502AFFC3FFEA0BCE0D0C0E5C9117" box="[1237,1394,301,330]" degrees="09" direction="north" minutes="21" orientation="latitude" pageId="87" pageNumber="88" precision="1" seconds="9.8" value="9.352722">09º21′9.8″ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA60D700A479130" box="[189,361,337,365]" degrees="73" direction="west" minutes="48" orientation="longitude" pageId="87" pageNumber="88" precision="1" seconds="22.2" value="-73.80617">73º48′22.2″ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0E6E0D720A9D9130" box="[373,435,339,365]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="87" pageNumber="88" unit="m" value="42.0">42 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0EA40D72097F9130" box="[447,593,339,365]" pageId="87" pageNumber="88" value="2010-03-07">7 Mar. 2010</date>
, pitfall with human faeces, P. Delgado leg. (
<collectionCode id="4994AE28FFC3FFEA0B4F0D720F829130" box="[1108,1196,339,365]" pageId="87" pageNumber="88">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0BCE0D730E0B9130" bold="true" box="[1237,1317,338,365]" pageId="87" pageNumber="88">Chocó</emphasis>
: 2 ♂♂ (1 dissected), 
<specimenCount id="3983FD64FFC3FFEA0E440D560AA491CC" box="[351,394,375,401]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Acandí, Capurganá, “caminho ‘Al Cielo’”, 
<quantity id="E87D9B08FFC3FFEA0C950D5608E491CC" box="[910,970,375,401]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="87" pageNumber="88" unit="m" value="10.0">10 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0CCD0D570F7291CC" box="[982,1116,374,401]" pageId="87" pageNumber="88" value="1999-01-06">6 Jan. 1999</date>
, A. Vitolo leg. (
<collectionCode id="4994AE28FFC3FFEA0A090D560E4491CC" box="[1298,1386,375,401]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 2 ♂♂ (1 dissected), Acandí, Capurganá, “Jardin”, 
<geoCoordinate id="4AB1502AFFC3FFEA0C0C0DB9088991E9" box="[791,935,408,436]" degrees="08" direction="south" minutes="37" orientation="latitude" pageId="87" pageNumber="88" precision="15" seconds="42" value="-8.628333">08º37′42″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0CAD0DB90F6091E8" box="[950,1102,408,437]" degrees="77" direction="west" minutes="21" orientation="longitude" pageId="87" pageNumber="88" precision="15" seconds="12" value="-77.35333">77º21′12″ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0B470DBB0FB591E8" box="[1116,1179,410,437]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="87" pageNumber="88" unit="m" value="30.0">30 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0BB10DBB0E5C91E9" box="[1194,1394,410,437]" pageId="87" pageNumber="88" value="2008-01-16" valueMax="2008-01-18" valueMin="2008-01-16">16–18 Jan. 2008</date>
, pitfall with human faeces, Arias 
<emphasis id="1DF1EAFFFFC3FFEA0D2F0D9E09409185" box="[564,622,446,472]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0DB50D9F08289185" box="[686,774,446,472]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FFC3FFEA0C4A0D9F08BD9185" box="[849,915,446,472]" pageId="87" pageNumber="88" type="female">4 ♀♀</specimenCount>
, Lloró, 
<geoCoordinate id="4AB1502AFFC3FFEA0CF50D9D0F799185" box="[1006,1111,444,472]" degrees="05" direction="north" minutes="31" orientation="latitude" pageId="87" pageNumber="88" precision="925" value="5.5166664">05º31′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0B7A0D9D0FE39185" box="[1121,1229,444,472]" degrees="76" direction="west" minutes="33" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="-76.55">76º33′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0BC30D9F0E3A9185" box="[1240,1300,446,472]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="87" pageNumber="88" unit="m" value="90.0">90 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0A3B0D9F0BD491A1" pageId="87" pageNumber="88" value="2003-02-20">20 Feb. 2003</date>
, Olaya and Mosquero leg. (
<collectionCode id="4994AE28FFC3FFEA0D570DC3098991A1" box="[588,679,482,508]" pageId="87" pageNumber="88">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0DA50DC309C391A1" box="[702,749,482,508]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Parque Nacional Natural Ensenada de Utría, 
<date id="5B3B102DFFC3FFEA0A040DC30BD49242" pageId="87" pageNumber="88" value="1997-06-19">19 Jun. 1997</date>
, pitfall with human faeces, Llanos-Jurado leg. (
<collectionCode id="4994AE28FFC3FFEA0C360E2408AB9242" box="[813,901,517,543]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 2 ♂♂, 
<specimenCount id="3983FD64FFC3FFEA0CF70E240F37927D" box="[1004,1049,517,544]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Quibdó, Tutunendó (“ 
<quantity id="E87D9B08FFC3FFEA0A320E240E589242" box="[1321,1398,517,543]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="87" pageNumber="88" unit="km" value="20.0">20 km</quantity>
NE Quibdó”), 
<quantity id="E87D9B08FFC3FFEA0E730E080A889219" box="[360,422,553,580]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="87" pageNumber="88" unit="m" value="60.0">60 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0EA90E08097C921E" box="[434,594,553,580]" pageId="87" pageNumber="88" value="2001-11-26">26 Nov. 2001</date>
, J.C. Neita leg. (
<collectionCode id="4994AE28FFC3FFEA0C080E080840921E" box="[787,878,553,579]" pageId="87" pageNumber="88">TAMU</collectionCode>
); 3 ♂♂, Quibdó, Estación Ambiental Pandó, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA60E6A0A60923A" box="[189,334,587,615]" degrees="05" direction="south" minutes="42" orientation="latitude" pageId="87" pageNumber="88" precision="15" seconds="43" value="-5.711944">05º42′43″ S</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0E450E6A0AD7923A" box="[350,505,587,615]" degrees="76" direction="west" minutes="37" orientation="longitude" pageId="87" pageNumber="88" precision="15" seconds="59" value="-76.63306">76º37′59″ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0D130E6C0967923A" box="[520,585,589,615]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="87" pageNumber="88" unit="m" value="43.0">43 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0D420E6C0839923A" box="[601,791,589,615]" pageId="87" pageNumber="88" value="2010-06-09" valueMax="2010-06-11" valueMin="2010-06-09">9–11 Jun. 2010</date>
, pitfall with human faeces, J. Arias leg. (
<collectionCode id="4994AE28FFC3FFEA0A0A0E6C0E47923A" box="[1297,1385,589,615]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 4 ♂♂, 
<specimenCount id="3983FD64FFC3FFEA0E160E510A6192D6" box="[269,335,624,651]" pageId="87" pageNumber="88" type="female">6 ♀♀</specimenCount>
, Quibdó, Pacurita, 
<geoCoordinate id="4AB1502AFFC3FFEA0D2A0E4E09B492D6" box="[561,666,623,651]" degrees="05" direction="north" minutes="41" orientation="latitude" pageId="87" pageNumber="88" precision="925" value="5.6833334">05º41′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0DBE0E4E083F92D6" box="[677,785,623,651]" degrees="76" direction="west" minutes="40" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="-76.666664">76º40′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0C070E51087492D6" box="[796,858,624,651]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="87" pageNumber="88" unit="m" value="53.0">53 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0C7D0E510F2B92D6" box="[870,1029,624,651]" pageId="87" pageNumber="88" value="2001-11-25">25 Nov. 2001</date>
, dung, J.C. Neita leg. (
<collectionCode id="4994AE28FFC3FFEA0A140E500E4492D6" box="[1295,1386,625,651]" pageId="87" pageNumber="88">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0FA60EB50A2C92F2" box="[189,258,660,687]" pageId="87" pageNumber="88" type="female">2 ♀♀</specimenCount>
, Quibdó, Pacurita, “Arriba de la quebrada Aguas Claras”, 
<geoCoordinate id="4AB1502AFFC3FFEA0CDB0EB30F0292F2" box="[960,1068,658,687]" degrees="05" direction="north" minutes="42" orientation="latitude" pageId="87" pageNumber="88" precision="925" value="5.7">05º42′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0B220EB30F8692F2" box="[1081,1192,658,687]" degrees="76" direction="west" minutes="40" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="-76.666664">76º40′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0BAD0EB50FDB92F2" box="[1206,1269,660,687]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="87" pageNumber="88" unit="m" value="43.0">43 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0A1F0EB50BD5928F" pageId="87" pageNumber="88" value="2010-06-09" valueMax="2010-06-11" valueMin="2010-06-09">9–11 Jun. 2010</date>
, pitfall with human faeces, J. Arias leg. (
<collectionCode id="4994AE28FFC3FFEA0C1B0E990876928F" box="[768,856,696,722]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 2 ♂♂, Unión Panamericana, 
<geoCoordinate id="4AB1502AFFC3FFEA0BC10E970E5C928F" box="[1242,1394,694,722]" degrees="05" direction="north" minutes="32" orientation="latitude" pageId="87" pageNumber="88" precision="15" seconds="45" value="5.545833">05º32′45″ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA60EFB0A7B92AB" box="[189,341,730,758]" degrees="76" direction="west" minutes="44" orientation="longitude" pageId="87" pageNumber="88" precision="15" seconds="33" value="-76.74249">76º44′33″ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0E780EFA0A9E92AB" box="[355,432,731,758]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="87" pageNumber="88" unit="m" value="115.0">115 m</quantity>
, J.C. Neita leg. (
<collectionCode id="4994AE28FFC3FFEA0D610EFD09FB92AB" box="[634,725,732,758]" pageId="87" pageNumber="88">TAMU</collectionCode>
); 3 ♂♂ (1 dissected), 
<specimenCount id="3983FD64FFC3FFEA0CC40EFD0F2292AB" box="[991,1036,732,758]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Unión Panamericana, Salero, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA60EDC0A099347" box="[189,295,765,794]" degrees="05" direction="north" minutes="32" orientation="latitude" pageId="87" pageNumber="88" precision="925" value="5.5333333">05º32′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0E2F0EDC0A8F9347" box="[308,417,765,794]" degrees="76" direction="west" minutes="44" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="-76.73333">76º44′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0EB60EDE0AD49347" box="[429,506,767,794]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="87" pageNumber="88" unit="m" value="120.0">120 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0D1C0EDE099F9344" box="[519,689,767,794]" pageId="87" pageNumber="88" value="2010-06-03" valueMax="2010-06-05" valueMin="2010-06-03">3–5 Jun. 2010</date>
, pitfall with human faeces, J. Arias leg. (
<collectionCode id="4994AE28FFC3FFEA0B8F0EDE0FC29344" box="[1172,1260,767,793]" pageId="87" pageNumber="88">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0A0D0EDE0A349360" bold="true" pageId="87" pageNumber="88">Distrito Capital</emphasis>
(?): 
<specimenCount id="3983FD64FFC3FFEA0E510F020AFC9360" box="[330,466,803,829]" pageId="87" pageNumber="88" type="generic">1 specimen</specimenCount>
, 
<collectingRegion id="ED41F80FFFC3FFEA0EC60F02091C9360" box="[477,562,803,829]" country="Colombia" name="Bogota" pageId="87" pageNumber="88">Bogotá</collectingRegion>
(
<collectionCode id="4994AE28FFC3FFEA0D590F02098F9360" box="[578,673,803,829]" pageId="87" pageNumber="88">BMNH</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0DD20F0208659360" bold="true" box="[713,843,803,829]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0DD20F0208659360" box="[713,843,803,829]" country="Colombia" name="Santander" pageId="87" pageNumber="88">Santander</collectingRegion>
</emphasis>
: 3 ♂♂, 
<specimenCount id="3983FD64FFC3FFEA0CB70F0208F69363" box="[940,984,803,830]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Serranía de las Quinchas, Reserva El Paujil, 
<date id="5B3B102DFFC3FFEA0E2B0F660AB0933C" box="[304,414,839,865]" pageId="87" pageNumber="88" value="2006-07">Jul. 2006</date>
, pitfall with 
<taxonomicName id="E8854D6EFFC3FFEA0D2B0F660826933C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[560,776,839,865]" class="Mammalia" family="Atelidae" genus="Alouatta" kingdom="Animalia" order="Primates" pageId="87" pageNumber="88" phylum="Chordata" rank="species" species="seniculus">
<emphasis id="1DF1EAFFFFC3FFEA0D2B0F660826933C" box="[560,776,839,865]" italics="true" pageId="87" pageNumber="88">Alouatta seniculus</emphasis>
</taxonomicName>
dung, Santos Zarate leg. (
<collectionCode id="4994AE28FFC3FFEA0B2C0F660FA1933C" box="[1079,1167,839,865]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FFC3FFEA0BC70F660E29933C" box="[1244,1287,839,865]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Serranía de las Quinchas, Reserva El Paujil, 
<date id="5B3B102DFFC3FFEA0D7E0F4A09F093D9" box="[613,734,874,901]" pageId="87" pageNumber="88" value="2006-09">Sep. 2006</date>
, pitfall with 
<taxonomicName id="E8854D6EFFC3FFEA0C6E0F4B0F6193D9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[885,1103,874,900]" class="Mammalia" family="Atelidae" genus="Alouatta" kingdom="Animalia" order="Primates" pageId="87" pageNumber="88" phylum="Chordata" rank="species" species="seniculus">
<emphasis id="1DF1EAFFFFC3FFEA0C6E0F4B0F6193D9" box="[885,1103,874,900]" italics="true" pageId="87" pageNumber="88">Alouatta seniculus</emphasis>
</taxonomicName>
dung, Santos Zarate leg. (
<collectionCode id="4994AE28FFC3FFEA0FDE0FAF0A3393F5" box="[197,285,910,936]" pageId="87" pageNumber="88">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0E5E0FAF0AB293F5" bold="true" box="[325,412,910,936]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0E5E0FAF0AB293F5" box="[325,412,910,936]" country="Colombia" name="Tolima" pageId="87" pageNumber="88">Tolima</collectingRegion>
</emphasis>
: 1 ♂, 
<quantity id="E87D9B08FFC3FFEA0EF80FAF090193F4" box="[483,559,910,937]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="87" pageNumber="88" unit="m" value="300.0">300 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0D210FAE099893F4" box="[570,694,910,937]" pageId="87" pageNumber="88" value="1995-11">Nov. 1995</date>
, human faeces, F. Escobar leg. (
<collectionCode id="4994AE28FFC3FFEA0B320FAF0FAF93F5" box="[1065,1153,910,936]" pageId="87" pageNumber="88">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0BB20FAF0E4093F5" bold="true" box="[1193,1390,910,936]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0BB20FAF0E4093F5" box="[1193,1390,910,936]" country="Colombia" name="Valle del Cauca" pageId="87" pageNumber="88">Valle Del Cauca</collectingRegion>
</emphasis>
: 1 ♂, Estación Agro-Forestal Bajo Calima, 
<quantity id="E87D9B08FFC3FFEA0DA80F9309DF9391" box="[691,753,946,973]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="87" pageNumber="88" unit="m" value="50.0">50 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0DE60F9308A49390" box="[765,906,946,973]" pageId="87" pageNumber="88" value="1995-10-02">2 Oct. 1995</date>
, A. Lopera leg. (
<collectionCode id="4994AE28FFC3FFEA0B540F930F899391" box="[1103,1191,946,972]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 1 ♂ (dissected), Dagua, Río Escalerete, 
<date id="5B3B102DFFC3FFEA0ECA0FF709E993AD" box="[465,711,982,1008]" pageId="87" pageNumber="88" value="1991-03-31" valueMax="1991-04-04" valueMin="1991-03-31">31 Mar.–4 Apr. 1991</date>
, L.C. Pardo Locarno leg. (
<collectionCode id="4994AE28FFC3FFEA0CE40FF70F7993AD" box="[1023,1111,982,1008]" pageId="87" pageNumber="88">CEMT</collectionCode>
).
</paragraph>
<caption id="7BFA6665FFC2FFEB0FA60B8C0F8797B6" ID-DOI="http://doi.org/10.5281/zenodo.3824835" ID-Zenodo-Dep="3824835" httpUri="https://zenodo.org/record/3824835/files/figure.png" pageId="86" pageNumber="87" startId="86.[189,232,1965,1991]" targetBox="[224,1361,268,1930]" targetPageId="86">
<paragraph id="2F3A36EDFFC2FFEB0FA60B8C0F8797B6" blockId="86.[189,1399,1965,2027]" pageId="86" pageNumber="87">
<emphasis id="1DF1EAFFFFC2FFEB0FA60B8C0A77979A" bold="true" box="[189,345,1965,1991]" pageId="86" pageNumber="87">Fig. 28. A–B</emphasis>
. 
<taxonomicName id="E8854D6EFFC2FFEB0E720B8F08489795" authority="(Harold, 1868)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[361,870,1965,1992]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="aequinoctialis" status="comb. nov.">
<emphasis id="1DF1EAFFFFC2FFEB0E720B8F09809795" box="[361,686,1966,1992]" italics="true" pageId="86" pageNumber="87">Sylvicanthon aequinoctialis</emphasis>
(Harold, 1868)
</taxonomicName>
<taxonomicNameLabel id="06C25784FFC2FFEB0C750B8F08DF9795" box="[878,1009,1966,1992]" pageId="86" pageNumber="87" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="1DF1EAFFFFC2FFEB0CEC0B8C0F22979A" bold="true" box="[1015,1036,1965,1991]" pageId="86" pageNumber="87">A</emphasis>
. Dorsal view. 
<emphasis id="1DF1EAFFFFC2FFEB0BA30B8F0FE29795" bold="true" box="[1208,1228,1966,1992]" pageId="86" pageNumber="87">B</emphasis>
. Ventral view. 
<emphasis id="1DF1EAFFFFC2FFEB0FA60BF00BD897B6" bold="true" box="[189,246,2001,2027]" pageId="86" pageNumber="87">C–D</emphasis>
. 
<taxonomicName id="E8854D6EFFC2FFEB0E1E0BF0096997B6" authority="(Martinez, 1949)" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[261,583,2001,2027]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC2FFEB0E1E0BF00A5797B6" box="[261,377,2001,2027]" italics="true" pageId="86" pageNumber="87">S. proseni</emphasis>
(Martínez, 1949)
</taxonomicName>
stat. et comb. nov. 
<emphasis id="1DF1EAFFFFC2FFEB0C300BF0086E97B6" bold="true" box="[811,832,2001,2027]" pageId="86" pageNumber="87">C</emphasis>
. Dorsal view. 
<emphasis id="1DF1EAFFFFC2FFEB0CF10BF008D197B6" bold="true" box="[1002,1023,2001,2027]" pageId="86" pageNumber="87">D</emphasis>
. Ventral view.
</paragraph>
</caption>
<paragraph id="2F3A36EDFFC3FFEA0FA60839094296A1" blockId="87.[189,1399,1047,1788]" pageId="87" pageNumber="88">
<collectingCountry id="5792767DFFC3FFEA0FA608390A5A946F" box="[189,372,1048,1074]" name="Costa Rica" pageId="87" pageNumber="88">COSTA RICA</collectingCountry>
: 
<emphasis id="1DF1EAFFFFC3FFEA0E9C08360935946C" bold="true" box="[391,539,1047,1073]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0E9C08360935946C" box="[391,539,1047,1073]" country="Costa Rica" name="Guanacaste" pageId="87" pageNumber="88">Guanacaste</collectingRegion>
</emphasis>
: Volcán Rincón de la Vieja, Hacienda Guachipelín, 
<date id="5B3B102DFFC3FFEA0BDB08390E5C946F" box="[1216,1394,1048,1074]" pageId="87" pageNumber="88" value="1999-08-13">13 Aug. 1999</date>
, trap with human faeces, J. L. Moreno &amp; T. Mestre leg. 1 ♂ (
<collectionCode id="4994AE28FFC3FFEA0CA0081A0F3A9408" box="[955,1044,1083,1109]" pageId="87" pageNumber="88">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0B5D081B0FB49408" bold="true" box="[1094,1178,1082,1109]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0B5D081B0FB49408" box="[1094,1178,1082,1109]" country="Costa Rica" name="Limon" pageId="87" pageNumber="88">Limón</collectingRegion>
</emphasis>
: Río Reventazón, Humburgfarm, 
<date id="5B3B102DFFC3FFEA0E63087E09359424" box="[376,539,1119,1146]" pageId="87" pageNumber="88" value="1923-04-15">15 Apr. 1923</date>
, without collector – 
<specimenCount id="3983FD64FFC3FFEA0C07087E08629424" box="[796,844,1119,1145]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
(
<collectionCode id="4994AE28FFC3FFEA0C44087E089D9424" box="[863,947,1119,1145]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="87" pageNumber="88" type="Museum">MZSP</collectionCode>
); Río Reventazón, Humburgfarm, 
<date id="5B3B102DFFC3FFEA0A42087E0A1194C0" pageId="87" pageNumber="88" value="1925-08-27">27 Aug. 1925</date>
, F. Nevermann leg. 1 ♂ (
<collectionCode id="4994AE28FFC3FFEA0D6E08A209E794C0" box="[629,713,1155,1181]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="87" pageNumber="88" type="Museum">MZSP</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0DEC08A208A994C0" bold="true" box="[759,903,1155,1181]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0DEC08A208A994C0" box="[759,903,1155,1181]" country="Costa Rica" name="Puntarenas" pageId="87" pageNumber="88">Puntarenas</collectingRegion>
</emphasis>
: Osa, 
<date id="5B3B102DFFC3FFEA0CC308A30F8694C1" box="[984,1192,1154,1181]" pageId="87" pageNumber="88" value="1966-08-10" valueMax="1966-08-15" valueMin="1966-08-10">10–15 Aug. 1966</date>
[trap with rotten flesh], S. Peck leg. 1 ♂ (
<collectionCode id="4994AE28FFC3FFEA0EF608870914949D" box="[493,570,1190,1216]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="87" pageNumber="88">CNCI</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0D49088709D8949D" box="[594,758,1190,1216]" pageId="87" pageNumber="88" type="generic">11 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0EB908E9091394B8" box="[418,573,1224,1253]" degrees="08" direction="north" minutes="30.259" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="8.504316">08º30.259′ N</geoCoordinate>
<geoCoordinate id="4AB1502AFFC3FFEA0D5E08E909C894B8" box="[581,742,1224,1253]" degrees="83" direction="west" minutes="35.958" orientation="longitude" pageId="87" pageNumber="88" precision="1" value="-83.5993">83º35.958′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0DE908EB081F94B8" box="[754,817,1226,1253]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="87" pageNumber="88" unit="m" value="18.0">18 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0C2608EB08D094B9" box="[829,1022,1226,1252]" pageId="87" pageNumber="88" value="2009-07-17" valueMax="2009-07-20" valueMin="2009-07-17">17–20 Jul. 2009</date>
, Mehrabi 
<emphasis id="1DF1EAFFFFC3FFEA0B6E08EA0F9F94B9" box="[1141,1201,1226,1252]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0BE808EB0E4794B9" box="[1267,1385,1226,1252]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0FA608CF0A4C9555" box="[189,354,1262,1288]" pageId="87" pageNumber="88" type="generic">20 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0BC908CD0E5C9555" box="[1234,1394,1260,1288]" degrees="08" direction="north" minutes="30.502" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="8.508367">08º30.502′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA609310A4F9571" box="[189,353,1296,1324]" degrees="83" direction="west" minutes="36.285" orientation="longitude" pageId="87" pageNumber="88" precision="1" value="-83.60475">83º36.285′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0E7409330A9F9571" box="[367,433,1298,1324]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="87" pageNumber="88" unit="m" value="29.0">29 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0EA4093309AB9571" box="[447,645,1297,1324]" pageId="87" pageNumber="88" value="2009-07-29" valueMax="2009-07-31" valueMin="2009-07-29">29–31 Jul. 2009</date>
, Mehrabi 
<emphasis id="1DF1EAFFFFC3FFEA0C1A093308119576" box="[769,831,1297,1323]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0C9C093308D39571" box="[903,1021,1298,1324]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 52 ♂♂ (1 dissected), 
<specimenCount id="3983FD64FFC3FFEA0A0009300E5E9571" box="[1307,1392,1297,1324]" pageId="87" pageNumber="88" type="female">46 ♀♀</specimenCount>
, Península de Osa, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0DC30912086D950D" box="[728,835,1331,1360]" degrees="08" direction="north" minutes="30" orientation="latitude" pageId="87" pageNumber="88" precision="925" value="8.5">08º30′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFC3FFEA0C4A091208EE950D" box="[849,960,1331,1360]" degrees="83" direction="west" minutes="36" orientation="longitude" pageId="87" pageNumber="88" precision="925" value="-83.6">83º36′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0CD609140F20950D" box="[973,1038,1333,1360]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="87" pageNumber="88" unit="m" value="30.0">30 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0B0009140FA59512" box="[1051,1163,1333,1359]" pageId="87" pageNumber="88" value="2009-07">Jul. 2009</date>
, swine dung, Mann leg. (
<collectionCode id="4994AE28FFC3FFEA0FE709780A7B952E" box="[252,341,1369,1395]" pageId="87" pageNumber="88">CEMT</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0E770978092A952E" box="[364,516,1369,1395]" pageId="87" pageNumber="88" type="generic">7 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA6095A0A7995CA" box="[189,343,1403,1431]" degrees="08" direction="north" minutes="29.891" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="8.498183">08º29.891′ N</geoCoordinate>
35º998′ W, 
<quantity id="E87D9B08FFC3FFEA0EFF095C090C95CA" box="[484,546,1405,1431]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="87" pageNumber="88" unit="m" value="31.0">31 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0D36095C09C595CA" box="[557,747,1404,1431]" pageId="87" pageNumber="88" value="2009-07-11" valueMax="2009-07-14" valueMin="2009-07-11">11–14 Jul. 2009</date>
, Mehrabi 
<emphasis id="1DF1EAFFFFC3FFEA0C7A095C08B595CA" box="[865,923,1405,1431]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0CC7095C0F7C95CA" box="[988,1106,1405,1431]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0B7E095C0FD795CB" box="[1125,1273,1404,1431]" pageId="87" pageNumber="88" type="generic">9 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0C9009BF0F3895E6" box="[907,1046,1438,1467]" degrees="08" direction="north" minutes="29.46" orientation="latitude" pageId="87" pageNumber="88" precision="9" value="8.491">08º29.46′ N</geoCoordinate>
<geoCoordinate id="4AB1502AFFC3FFEA0B0609BF0F8195E6" box="[1053,1199,1438,1467]" degrees="83" direction="west" minutes="35.57" orientation="longitude" pageId="87" pageNumber="88" precision="9" value="-83.592834">83º35.57′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0BA109810FD695E6" box="[1210,1272,1440,1467]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="87" pageNumber="88" unit="m" value="41.0">41 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0A1809810E5C95E7" box="[1283,1394,1440,1466]" pageId="87" pageNumber="88" value="2009-07">Jul. 2009</date>
, Mehrabi, Coals, Cowburn and Yu leg. (
<collectionCode id="4994AE28FFC3FFEA0D9309E509D09583" box="[648,766,1476,1502]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0C0A09E5088B9583" box="[785,933,1476,1502]" pageId="87" pageNumber="88" type="generic">2 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0D2209C709FA965F" box="[569,724,1510,1538]" degrees="08" direction="north" minutes="30.897" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="8.51495">08º30.897′ N</geoCoordinate>
<geoCoordinate id="4AB1502AFFC3FFEA0DC709C70853965F" box="[732,893,1510,1538]" degrees="83" direction="west" minutes="35.847" orientation="longitude" pageId="87" pageNumber="88" precision="1" value="-83.59745">83º35.847′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0C9209C908E9965F" box="[905,967,1512,1538]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="87" pageNumber="88" unit="m" value="42.0">42 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0CC809C90FEA965F" box="[979,1220,1512,1538]" pageId="87" pageNumber="88" value="2009-07-31" valueMax="2009-08-02" valueMin="2009-07-31">31 Jul.–2 Aug. 2009</date>
, Mehrabi 
<emphasis id="1DF1EAFFFFC3FFEA0A2009C80E59965F" box="[1339,1399,1512,1538]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0FE20A2D0A41967B" box="[249,367,1548,1574]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0E9E0A2D09349678" box="[389,538,1547,1574]" pageId="87" pageNumber="88" type="generic">2 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0FA60A0C0A729617" box="[189,348,1581,1610]" degrees="08" direction="north" minutes="28.230" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="8.4705">08º28.230′ N</geoCoordinate>
<geoCoordinate id="4AB1502AFFC3FFEA0E720A0C09219617" box="[361,527,1581,1610]" degrees="83" direction="west" minutes="35.275" orientation="longitude" pageId="87" pageNumber="88" precision="1" value="-83.58791">83º35.275′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0D040A0E094D9617" box="[543,611,1583,1610]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="87" pageNumber="88" unit="m" value="47.0">47 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0D6F0A0E08109614" box="[628,830,1583,1609]" pageId="87" pageNumber="88" value="2009-07-26" valueMax="2009-07-29" valueMin="2009-07-26">26–29 Jul. 2009</date>
, Mehrabi 
<emphasis id="1DF1EAFFFFC3FFEA0CA50A1108D09614" box="[958,1022,1583,1609]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0B510A0E0FEE9614" box="[1098,1216,1583,1609]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0BC20A0E0E5C9614" box="[1241,1394,1583,1609]" pageId="87" pageNumber="88" type="generic">8 specimens</specimenCount>
, Península de Osa, Corcovado National Park, Sirena Biological Station, 
<geoCoordinate id="4AB1502AFFC3FFEA0B3E0A700FED9630" box="[1061,1219,1617,1645]" degrees="08" direction="north" minutes="29.287" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="8.488116">08º29.287′ N</geoCoordinate>
<geoCoordinate id="4AB1502AFFC3FFEA0BD50A700E5C9630" box="[1230,1394,1617,1645]" degrees="83" direction="west" minutes="35.327" orientation="longitude" pageId="87" pageNumber="88" precision="1" value="-83.58878">83º35.327′ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0FA60A560BD596CC" box="[189,251,1655,1681]" metricMagnitude="1" metricUnit="m" metricValue="4.9" pageId="87" pageNumber="88" unit="m" value="49.0">49 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0E1C0A560AE896CC" box="[263,454,1654,1681]" pageId="87" pageNumber="88" value="2009-07-19" valueMax="2009-07-22" valueMin="2009-07-19">19–22 Jul. 2009</date>
, Mehrabi 
<emphasis id="1DF1EAFFFFC3FFEA0D260A56095696CC" box="[573,632,1655,1681]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
leg. (
<collectionCode id="4994AE28FFC3FFEA0DA20A56080196CC" box="[697,815,1655,1681]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0C580A5608E296CD" box="[835,972,1654,1681]" pageId="87" pageNumber="88" type="generic">1 specimen</specimenCount>
, Península de Osa, Osa Biodiversity Centre, 
<geoCoordinate id="4AB1502AFFC3FFEA0E000AB90A8396E8" box="[283,429,1688,1717]" degrees="08" direction="north" minutes="24" orientation="latitude" pageId="87" pageNumber="88" precision="15" seconds="14" value="8.403889">08º24′14″ N</geoCoordinate>
<geoCoordinate id="4AB1502AFFC3FFEA0EAD0AB9096096E8" box="[438,590,1688,1717]" degrees="83" direction="west" minutes="20" orientation="longitude" pageId="87" pageNumber="88" precision="15" seconds="09" value="-83.33584">83º20′09″ W</geoCoordinate>
, 
<quantity id="E87D9B08FFC3FFEA0D400ABB098796E8" box="[603,681,1690,1717]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="87" pageNumber="88" unit="m" value="115.0">115 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0DAF0ABB080096E9" box="[692,814,1690,1716]" pageId="87" pageNumber="88" value="2010-04">Apr. 2010</date>
, Mann and Mehrabi leg. (
<collectionCode id="4994AE28FFC3FFEA0B780ABB0FF796E9" box="[1123,1241,1690,1716]" pageId="87" pageNumber="88">OUMNH</collectionCode>
); 3 ♂♂, 
<specimenCount id="3983FD64FFC3FFEA0A590ABB0E4196E8" box="[1346,1391,1690,1717]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Reserva Biológica Bosque Nuboso Monteverde (“Parque Nacional Monteverde”), 
<date id="5B3B102DFFC3FFEA0B830A9F0E139685" box="[1176,1341,1726,1752]" pageId="87" pageNumber="88" value="1999-08-13">13 Aug. 1999</date>
, J.L. Moreno and T. Mestre leg. (
<collectionCode id="4994AE28FFC3FFEA0D1C0AC3094E96A1" box="[519,608,1762,1788]" pageId="87" pageNumber="88">CEMT</collectionCode>
).
</paragraph>
<paragraph id="2F3A36EDFFC3FFEA0FA60B050996973C" blockId="87.[189,1398,1826,1890]" pageId="87" pageNumber="88">
<collectingCountry id="5792767DFFC3FFEA0FA60B050A4B9763" box="[189,357,1828,1854]" name="Honduras" pageId="87" pageNumber="88">HONDURAS</collectingCountry>
: 
<emphasis id="1DF1EAFFFFC3FFEA0E6F0B020AC59763" bold="true" box="[372,491,1827,1854]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0E6F0B020AC59763" box="[372,491,1827,1854]" country="Honduras" name="Atlantida" pageId="87" pageNumber="88">Atlántida</collectingRegion>
</emphasis>
: 1 ♂ (dissected), 
<specimenCount id="3983FD64FFC3FFEA0DD60B05083B9763" box="[717,789,1828,1854]" pageId="87" pageNumber="88" type="female">2 ♀♀</specimenCount>
, 
<quantity id="E87D9B08FFC3FFEA0C3C0B0208AF9763" box="[807,897,1827,1854]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="87" pageNumber="88" unit="km" value="15.0">15 km</quantity>
W of La Ceiba, 
<date id="5B3B102DFFC3FFEA0B410B020E069760" box="[1114,1320,1827,1854]" pageId="87" pageNumber="88" value="1996-06-15" valueMax="1996-06-19" valueMin="1996-06-15">15–19 Jun. 1996</date>
, flight interception trap, R. Lehman leg. (
<collectionCode id="4994AE28FFC3FFEA0D4A0B660982973C" box="[593,684,1863,1889]" pageId="87" pageNumber="88">TAMU</collectionCode>
).
</paragraph>
<paragraph id="2F3A36EDFFC3FFE40FA60BA8080792CC" blockId="87.[189,1399,1929,2027]" lastBlockId="89.[189,1399,267,658]" lastPageId="89" lastPageNumber="90" pageId="87" pageNumber="88">
<collectingCountry id="5792767DFFC3FFEA0FA60BA80A5C97FE" box="[189,370,1929,1955]" name="Nicaragua" pageId="87" pageNumber="88">NICARAGUA</collectingCountry>
: 
<emphasis id="1DF1EAFFFFC3FFEA0E600BA80AD697FE" bold="true" box="[379,504,1929,1955]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0E600BA80AD697FE" box="[379,504,1929,1955]" country="Nicaragua" name="Chontales" pageId="87" pageNumber="88">Chontales</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFC3FFEA0D1C0BA809A097FE" box="[519,654,1929,1955]" pageId="87" pageNumber="88" type="generic">1 specimen</specimenCount>
, T. Belt leg. (
<collectionCode id="4994AE28FFC3FFEA0C3E0BA808AC97FE" box="[805,898,1929,1955]" pageId="87" pageNumber="88">BMNH</collectionCode>
, 
<collectionCode id="4994AE28FFC3FFEA0C970BA808E397FE" box="[908,973,1929,1955]" pageId="87" pageNumber="88">BCA</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0CEF0BA80F7097FE" bold="true" box="[1012,1118,1929,1955]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0CEF0BA80F7097FE" box="[1012,1118,1929,1955]" country="Nicaragua" name="Jinotega" pageId="87" pageNumber="88">Jinotega</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFC3FFEA0B770BA80F8097F9" box="[1132,1198,1929,1956]" pageId="87" pageNumber="88" type="female">2 ♀♀</specimenCount>
, Reserva Natural Cerro Kilambé, 
<quantity id="E87D9B08FFC3FFEA0E640B8C0AF29795" box="[383,476,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="87" pageNumber="88" unit="m" value="1000.0">1000 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0EF20B8C0947979A" box="[489,617,1965,1991]" pageId="87" pageNumber="88" value="2001-08">Aug. 2001</date>
, M.A. Guatemala leg. (
<collectionCode id="4994AE28FFC3FFEA0C9F0B8C08F9979A" box="[900,983,1965,1991]" pageId="87" pageNumber="88">SEAN</collectionCode>
); 
<specimenCount id="3983FD64FFC3FFEA0CF50B8C0F329795" box="[1006,1052,1965,1992]" pageId="87" pageNumber="88" type="female">1 ♀</specimenCount>
, Santa Maura, 
<quantity id="E87D9B08FFC3FFEA0BCF0B8C0E1F9795" box="[1236,1329,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="87" pageNumber="88" unit="m" value="1215.0">1215 m</quantity>
, 
<date id="5B3B102DFFC3FFEA0A5B0B8F0BD597B6" pageId="87" pageNumber="88" value="2002-03">Mar. 2002</date>
, L. Pivotti leg. (
<collectionCode id="4994AE28FFC3FFEA0EA60BF0093E97B6" box="[445,528,2001,2027]" pageId="87" pageNumber="88">SEAN</collectionCode>
). – 
<emphasis id="1DF1EAFFFFC3FFEA0D270BF009ED97B6" bold="true" box="[572,707,2001,2027]" pageId="87" pageNumber="88">
<collectingRegion id="ED41F80FFFC3FFEA0D270BF009ED97B6" box="[572,707,2001,2027]" country="Nicaragua" name="Matagalpa" pageId="87" pageNumber="88">Matagalpa</collectingRegion>
</emphasis>
: 41 ♂♂, 
<specimenCount id="3983FD64FFC3FFEA0C2D0BF008A797B6" box="[822,905,2001,2027]" pageId="87" pageNumber="88" type="female">17 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFC3FFEA0B0E0BF10FE897B6" box="[1045,1222,2000,2027]" pageId="87" pageNumber="88" value="2003-12-05" valueMax="2003-12-06" valueMin="2003-12-05">5–6 Dec. 2003</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40FE10D2C0A63917A" box="[250,333,269,295]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 3 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40EA20D2C0AD1917A" box="[441,511,269,295]" pageId="89" pageNumber="90" type="female">4 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40D960D2C0810917A" box="[653,830,269,295]" pageId="89" pageNumber="90" value="2004-02-08" valueMax="2004-02-09" valueMin="2004-02-08">8–9 Feb. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40B2D0D2C0FA7917A" box="[1078,1161,269,295]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 5 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40BEE0D2C0E14917A" box="[1269,1338,269,295]" pageId="89" pageNumber="90" type="female">2 ♀♀</specimenCount>
, 
<date id="5B3B102DFFCDFFE40A510D2C0A749116" pageId="89" pageNumber="90" value="2004-02-13" valueMax="2004-02-14" valueMin="2004-02-13">13– 14 Feb. 2004</date>
, B. Hernández leg. 5 (
<collectionCode id="4994AE28FFCDFFE40D450D10099F9116" box="[606,689,305,331]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 18 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40C3F0D1008589111" box="[804,886,305,332]" pageId="89" pageNumber="90" type="female">21 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40CE40D100FE49116" box="[1023,1226,305,332]" pageId="89" pageNumber="90" value="2004-03-10" valueMax="2004-03-11" valueMin="2004-03-10">10–11 Mar. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40FE30D740A659132" box="[248,331,341,367]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 3 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40EA90D740AF1912D" box="[434,479,341,368]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40D700D7408179132" box="[619,825,341,368]" pageId="89" pageNumber="90" value="2004-03-14" valueMax="2004-03-15" valueMin="2004-03-14">14–15 Mar. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40B310D740F539132" box="[1066,1149,341,367]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40B890D740F91912D" box="[1170,1215,341,368]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40A510D740A7891C9" pageId="89" pageNumber="90" value="2004-04-21" valueMax="2004-04-22" valueMin="2004-04-21">21– 22 Apr. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40D240D5B09BC91C9" box="[575,658,378,404]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 11 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40C1B0D5B087E91C9" box="[768,848,378,404]" pageId="89" pageNumber="90" type="female">17 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40CCC0D5B0FB291C9" box="[983,1180,378,404]" pageId="89" pageNumber="90" value="2004-04-23" valueMax="2004-04-24" valueMin="2004-04-23">23–24 Apr. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40FDE0DBF0A3691E5" box="[197,280,414,440]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 27 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40E9E0DBF0AFB91E5" box="[389,469,414,440]" pageId="89" pageNumber="90" type="female">23 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40D410DBF083091E5" box="[602,798,414,440]" pageId="89" pageNumber="90" value="2004-04-26" valueMax="2004-04-27" valueMin="2004-04-26">26–27Apr. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40B1F0DBF0F7991E5" box="[1028,1111,414,440]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 9 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40BAE0DBF0FD891E5" box="[1205,1270,414,440]" pageId="89" pageNumber="90" type="female">2 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40FA60DE30A459181" box="[189,363,450,477]" pageId="89" pageNumber="90" value="2004-06-03" valueMax="2004-06-04" valueMin="2004-06-03">3–4 Jun. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40D7E0DE309969181" box="[613,696,450,476]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 9 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40C3F0DE308449180" box="[804,874,450,477]" pageId="89" pageNumber="90" type="female">5 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40CE20DE30FEA9181" box="[1017,1220,450,477]" pageId="89" pageNumber="90" value="2004-06-19" valueMax="2004-06-20" valueMin="2004-06-19">19–20 Jun. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40FE20DC70A62925D" box="[249,332,486,512]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 7 ♂♂, Matiguas, 
<date id="5B3B102DFFCDFFE40D280DC709D3925D" box="[563,765,486,513]" pageId="89" pageNumber="90" value="2004-06-23" valueMax="2004-06-24" valueMin="2004-06-23">23–24 Jun. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40CE90DC70F6B925D" box="[1010,1093,486,512]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 2 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40BB50DC70FDD925C" box="[1198,1267,486,513]" pageId="89" pageNumber="90" type="female">4 ♀♀</specimenCount>
, Matiguas, 
<date id="5B3B102DFFCDFFE40FA60E2A0AAC9278" box="[189,386,523,549]" pageId="89" pageNumber="90" value="2004-06-29" valueMax="2004-06-30" valueMin="2004-06-29">29–30 Jun. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40D750E2A09EF9278" box="[622,705,523,549]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 1 ♂, Matiguas, 
<date id="5B3B102DFFCDFFE40C920E2A0F049278" box="[905,1066,522,549]" pageId="89" pageNumber="90" value="2004-07-02" valueMax="2004-07-03" valueMin="2004-07-02">2–3 Jul. 2004</date>
, B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40A0D0E2A0E479278" box="[1302,1385,523,549]" pageId="89" pageNumber="90">SEAN</collectionCode>
); 2 ♂♂, Reserva Natural Biósfera de Bosawas, Macizo de Peñas Blancas, 
<date id="5B3B102DFFCDFFE40B1D0E0E0FE29214" box="[1030,1228,559,585]" pageId="89" pageNumber="90" value="2017-04-29" valueMax="2017-04-30" valueMin="2017-04-29">29–30 Apr. 2017</date>
, B. Hernández leg (
<collectionCode id="4994AE28FFCDFFE40FE90E720A649230" box="[242,330,595,621]" pageId="89" pageNumber="90">CEMT</collectionCode>
). – 
<emphasis id="1DF1EAFFFFCDFFE40E6D0E73090C9230" bold="true" box="[374,546,594,621]" pageId="89" pageNumber="90">
<collectingRegion id="ED41F80FFFCDFFE40E6D0E73090C9230" box="[374,546,594,621]" country="Nicaragua" name="Rio San Juan" pageId="89" pageNumber="90">Río San Juan</collectingRegion>
</emphasis>
: 1 ♂, Reserva Biológica Indio Maíz, 
<geoCoordinate id="4AB1502AFFCDFFE40CFD0E700F549233" box="[998,1146,593,622]" degrees="10" direction="north" minutes="58" orientation="latitude" pageId="89" pageNumber="90" precision="15" seconds="24" value="10.973333">10º58′24″ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFCDFFE40B930E700E0E9233" box="[1160,1312,593,622]" degrees="84" direction="west" minutes="04" orientation="longitude" pageId="89" pageNumber="90" precision="15" seconds="52" value="-84.08111">84º04′52″ W</geoCoordinate>
, 
<date id="5B3B102DFFCDFFE40A360E720A1C92CC" pageId="89" pageNumber="90" value="2002-06-14" valueMax="2002-06-15" valueMin="2002-06-14">14–15 Jun. 2002</date>
, P. Schmit and B. Hernández leg. (
<collectionCode id="4994AE28FFCDFFE40DD20E56083292CC" box="[713,796,631,657]" pageId="89" pageNumber="90">SEAN</collectionCode>
).
</paragraph>
<caption id="7BFA6665FFCCFFE50FA60BA80E5897B4" ID-DOI="http://doi.org/10.5281/zenodo.3824837" ID-Zenodo-Dep="3824837" httpUri="https://zenodo.org/record/3824837/files/figure.png" pageId="88" pageNumber="89" startId="88.[189,232,1929,1955]" targetBox="[192,1399,370,1881]" targetPageId="88">
<paragraph id="2F3A36EDFFCCFFE50FA60BA80E5897B4" blockId="88.[189,1399,1929,2026]" pageId="88" pageNumber="89">
<emphasis id="1DF1EAFFFFCCFFE50FA60BA80A3797FE" bold="true" box="[189,281,1929,1955]" pageId="88" pageNumber="89">Fig. 29.</emphasis>
Type material of the 
<taxonomicName id="E8854D6EFFCCFFE50D0A0BA8099997FE" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[529,695,1929,1955]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCCFFE50D0A0BA8099997FE" box="[529,695,1929,1955]" italics="true" pageId="88" pageNumber="89">aequinoctialis</emphasis>
</taxonomicName>
subgroup. 
<emphasis id="1DF1EAFFFFCCFFE50C230BA8084197FE" bold="true" box="[824,879,1929,1955]" pageId="88" pageNumber="89">A–B</emphasis>
. 
<taxonomicName id="E8854D6EFFCCFFE50C660BA80E5897F9" authority="(Harold, 1868)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[893,1398,1929,1956]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="aequinoctialis" status="comb. nov.">
<emphasis id="1DF1EAFFFFCCFFE50C660BA80FEC97FE" box="[893,1218,1929,1955]" italics="true" pageId="88" pageNumber="89">Sylvicanthon aequinoctialis</emphasis>
(Harold, 1868)
</taxonomicName>
<taxonomicNameLabel id="06C25784FFCCFFE50FA60B8D0A6A979A" box="[189,324,1964,1991]" pageId="88" pageNumber="89" rank="species">comb. nov.</taxonomicNameLabel>
, type series. 
<emphasis id="1DF1EAFFFFCCFFE50EF80B8D0AD6979B" bold="true" box="[483,504,1964,1990]" pageId="88" pageNumber="89">A</emphasis>
. Lectotype and its labels. 
<emphasis id="1DF1EAFFFFCCFFE50C230B8D0862979B" bold="true" box="[824,844,1964,1990]" pageId="88" pageNumber="89">B</emphasis>
. Paralectotype and its labels. 
<emphasis id="1DF1EAFFFFCCFFE50BAC0B8D0FC0979B" bold="true" box="[1207,1262,1964,1990]" pageId="88" pageNumber="89">C–E</emphasis>
. 
<taxonomicName id="E8854D6EFFCCFFE50A1B0B8C0AAC97B4" authority="(Martinez, 1949)" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFCCFFE50A1B0B8C0E59979B" box="[1280,1399,1965,1991]" italics="true" pageId="88" pageNumber="89">S. proseni</emphasis>
(Martínez, 1949)
</taxonomicName>
stat. et comb. nov., type series. 
<emphasis id="1DF1EAFFFFCCFFE50DF40BEE082A97B4" bold="true" box="[751,772,1999,2025]" pageId="88" pageNumber="89">C</emphasis>
. Holotype and its labels. 
<emphasis id="1DF1EAFFFFCCFFE50B330BF10F1397B7" bold="true" box="[1064,1085,2000,2026]" pageId="88" pageNumber="89">D</emphasis>
. Paratype 1. 
<emphasis id="1DF1EAFFFFCCFFE50BC80BF10FC997B7" bold="true" box="[1235,1255,2000,2026]" pageId="88" pageNumber="89">E</emphasis>
. Paratype 2.
</paragraph>
</caption>
<paragraph id="2F3A36EDFFCDFFE40FA60E9B093E96B2" blockId="89.[189,1399,696,1776]" pageId="89" pageNumber="90">
<collectingCountry id="5792767DFFCDFFE40FA60E9B0A109289" box="[189,318,698,724]" name="Panama" pageId="89" pageNumber="90">PANAMA</collectingCountry>
: 
<emphasis id="1DF1EAFFFFCDFFE40E500E980A999289" bold="true" box="[331,439,697,724]" pageId="89" pageNumber="90">
<collectingRegion id="ED41F80FFFCDFFE40E500E980A999289" box="[331,439,697,724]" country="Panama" name="Chiriqui" pageId="89" pageNumber="90">Chiriquí</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFCDFFE40ED20E9B097A9289" box="[457,596,698,724]" pageId="89" pageNumber="90" type="generic">1 specimen</specimenCount>
, David, San José de David, Champion leg. (
<collectionCode id="4994AE28FFCDFFE40B770E9B0FE89289" box="[1132,1222,698,724]" pageId="89" pageNumber="90">BMNH</collectionCode>
– 
<collectionCode id="4994AE28FFCDFFE40BF30E9B0E079289" box="[1256,1321,698,724]" pageId="89" pageNumber="90">BCA</collectionCode>
); 1 ♂, Gualaca, Hornito, Finca La Suiza, 
<date id="5B3B102DFFCDFFE40D430EFF083B92A5" box="[600,789,734,761]" pageId="89" pageNumber="90" value="1999-05-09" valueMax="1999-05-15" valueMin="1999-05-09">9–15 May 1999</date>
, pitfall with human faeces, Wappes and Morris leg. (
<collectionCode id="4994AE28FFCDFFE40FDE0F230A0E9341" box="[197,288,770,796]" pageId="89" pageNumber="90">TAMU</collectionCode>
). – 
<emphasis id="1DF1EAFFFFCDFFE40E5D0F200ABF9341" bold="true" box="[326,401,769,796]" pageId="89" pageNumber="90">
<collectingRegion id="ED41F80FFFCDFFE40E5D0F200ABF9341" box="[326,401,769,796]" country="Panama" name="Colon" pageId="89" pageNumber="90">Colón</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFCDFFE40E840F230AE49340" box="[415,458,770,797]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Chagres, Fort San Lorenzo (“San Lorenzo Forest” sic), 
<geoCoordinate id="4AB1502AFFCDFFE40B420F210FEF9340" box="[1113,1217,768,797]" degrees="09" direction="north" minutes="17" orientation="latitude" pageId="89" pageNumber="90" precision="925" value="9.283334">09º17′ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFCDFFE40BD00F210E189340" box="[1227,1334,768,797]" degrees="79" direction="west" minutes="58" orientation="longitude" pageId="89" pageNumber="90" precision="925" value="-79.96667">79º58′ W</geoCoordinate>
, 
<date id="5B3B102DFFCDFFE40A5A0F220BD5931C" pageId="89" pageNumber="90" value="2004-05">May 2004</date>
, flight interception trap, A. Tishechkin leg. (
<collectionCode id="4994AE28FFCDFFE40DEF0F060862931C" box="[756,844,807,833]" pageId="89" pageNumber="90">CEMT</collectionCode>
); 1 ♂, Colón, 
<quantity id="E87D9B08FFCDFFE40CF10F060F07931D" box="[1002,1065,806,833]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="90" unit="km" value="2.0">2 km</quantity>
S of Sabanitas, 
<geoCoordinate id="4AB1502AFFCDFFE40BFA0F040E5C931C" box="[1249,1394,805,833]" degrees="09" direction="north" minutes="19" orientation="latitude" pageId="89" pageNumber="90" precision="15" seconds="19" value="9.321944">09º19′19″ N</geoCoordinate>
, 
<geoCoordinate id="4AB1502AFFCDFFE40FA60F680A789338" box="[189,342,841,869]" degrees="79" direction="west" minutes="47" orientation="longitude" pageId="89" pageNumber="90" precision="15" seconds="54" value="-79.79833">79º47′54″ W</geoCoordinate>
, 
<date id="5B3B102DFFCDFFE40E7E0F6A091E9338" box="[357,560,843,870]" pageId="89" pageNumber="90" value="1999-06-25" valueMax="1999-06-26" valueMin="1999-06-25">25–26 Jun. 1999</date>
, UV light, A. Gillogly leg. (
<collectionCode id="4994AE28FFCDFFE40C910F6A08CB9338" box="[906,997,843,869]" pageId="89" pageNumber="90">TAMU</collectionCode>
). – 
<emphasis id="1DF1EAFFFFCDFFE40B0F0F6B0F559338" bold="true" box="[1044,1147,842,869]" pageId="89" pageNumber="90">
<collectingRegion id="ED41F80FFFCDFFE40B0F0F6B0F559338" box="[1044,1147,842,869]" country="Panama" name="Panama" pageId="89" pageNumber="90">Panamá</collectingRegion>
</emphasis>
: 
<specimenCount id="3983FD64FFCDFFE40B960F6A0FFD9338" box="[1165,1235,843,869]" pageId="89" pageNumber="90" type="female">3 ♀♀</specimenCount>
, Canal Zone, Fort Kobbe, 
<date id="5B3B102DFFCDFFE40E4F0F4E0AC193D4" box="[340,495,879,906]" pageId="89" pageNumber="90" value="1976-06-14">14 Jun. 1976</date>
, E.G. Riley leg. (
<collectionCode id="4994AE28FFCDFFE40DA50F4E083793D4" box="[702,793,879,905]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 9 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40C9A0F4E08EB93D7" box="[897,965,879,906]" pageId="89" pageNumber="90" type="female">4 ♀♀</specimenCount>
, Canal Zone, Fort Kobbe, 
<date id="5B3B102DFFCDFFE40A180F4E0BD493F3" pageId="89" pageNumber="90" value="1985-06-04" valueMax="1985-06-21" valueMin="1985-06-04">4–21 Jun. 1985</date>
, pitfall with human faeces, E.G. Riley leg. (
<collectionCode id="4994AE28FFCDFFE40DE40FB2087493F0" box="[767,858,915,941]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40C750FB208D893F0" box="[878,1014,915,941]" pageId="89" pageNumber="90" type="generic">1 specimen</specimenCount>
, Canal Zone, Lago Gatún, Barro Colorado Island, K.W. Cooper leg. (
<collectionCode id="4994AE28FFCDFFE40D6A0F9909FF938F" box="[625,721,952,978]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="89" pageNumber="90">MNHN</collectionCode>
); 1 ♂, Canal Zone, Lago Gatún, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40FA60FFD0A7393AA" box="[189,349,988,1015]" pageId="89" pageNumber="90" value="1955-02-22">22 Feb. 1955</date>
, R. Freund leg. (
<collectionCode id="4994AE28FFCDFFE40D320FFD095B93AB" box="[553,629,988,1014]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="89" pageNumber="90">CNCI</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40D900FFD099793AB" box="[651,697,988,1014]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Canal Zone, Lago Gatún, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40A060FFD0BD49447" pageId="89" pageNumber="90" value="1960-02-13">13 Feb. 1960</date>
, S. Breeland leg. (
<collectionCode id="4994AE28FFCDFFE40EFA082109039447" box="[481,557,1024,1050]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="89" pageNumber="90">CNCI</collectionCode>
); 4 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40D84082109C89446" box="[671,742,1024,1051]" pageId="89" pageNumber="90" type="female">3 ♀♀</specimenCount>
, Canal Zone, Lago Gatún, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40A4208210A039463" pageId="89" pageNumber="90" value="1963-07-18">18 Jul. 1963</date>
, L. J. Bottimer leg. (
<collectionCode id="4994AE28FFCDFFE40D330805095A9463" box="[552,628,1060,1086]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="89" pageNumber="90">CNCI</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40D900804080E9463" box="[651,800,1060,1087]" pageId="89" pageNumber="90" type="generic">7 specimens</specimenCount>
, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40FA608680A16943E" box="[189,312,1096,1123]" pageId="89" pageNumber="90" value="1975-03">Mar. 1975</date>
, O.P. Young leg. (
<collectionCode id="4994AE28FFCDFFE40D1608680942943E" box="[525,620,1097,1123]" pageId="89" pageNumber="90">BMNH</collectionCode>
); 1 ♂, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40A1F08680E59943E" box="[1284,1399,1097,1123]" pageId="89" pageNumber="90" value="1978-06">Jun. 1978</date>
(
<collectionCode id="4994AE28FFCDFFE40FDE084C0A3394DA" box="[197,285,1133,1159]" pageId="89" pageNumber="90">CEMT</collectionCode>
); 2 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40E64084C0A8494DA" box="[383,426,1133,1159]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40CE8084C0F5B94DA" box="[1011,1141,1133,1159]" pageId="89" pageNumber="90" value="1978-07-09">9 Jul. 1978</date>
, dung trap, A. Forsyth leg. (
<collectionCode id="4994AE28FFCDFFE40FEC08B00A7C94F6" box="[247,338,1169,1195]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 1 ♂, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40CFB08B00F5C94F6" box="[992,1138,1169,1195]" pageId="89" pageNumber="90" value="1978-07-19">19 Jul. 1978</date>
, dung trap, A. Forsyth leg. (
<collectionCode id="4994AE28FFCDFFE40FED08940A7F9492" box="[246,337,1205,1231]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 1 ♂, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40CCC08940F5B9492" box="[983,1141,1205,1231]" pageId="89" pageNumber="90" value="1978-08-16">16 Aug. 1978</date>
, dung trap, A. Forsyth leg. (
<collectionCode id="4994AE28FFCDFFE40FE308FB0A7D94A9" box="[248,339,1242,1268]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40E7C08FB0AD594AE" box="[359,507,1241,1268]" pageId="89" pageNumber="90" type="generic">4 specimens</specimenCount>
, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40B5508FB0E3794A9" box="[1102,1305,1242,1268]" pageId="89" pageNumber="90" value="1981-05-12" valueMax="1981-05-14" valueMin="1981-05-12">12–14 May 1981</date>
, B. Gill leg. (
<collectionCode id="4994AE28FFCDFFE40FE208DF0A769545" box="[249,344,1278,1304]" pageId="89" pageNumber="90">BMNH</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40E7708DF0ADB9545" box="[364,501,1278,1304]" pageId="89" pageNumber="90" type="generic">1 specimen</specimenCount>
, Canal Zone, Gatún Lake, Barro Colorado Island, 
<date id="5B3B102DFFCDFFE40B5708DF0E369545" box="[1100,1304,1278,1304]" pageId="89" pageNumber="90" value="1981-05-20" valueMax="1981-05-28" valueMin="1981-05-20">20–28 May 1981</date>
, B. Gill leg. (
<collectionCode id="4994AE28FFCDFFE40FE009030A749561" box="[251,346,1314,1340]" pageId="89" pageNumber="90">BMNH</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FFCDFFE40EA809030ACC9560" box="[435,482,1314,1341]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Canal Zone, Río Changena, 
<date id="5B3B102DFFCDFFE40C55090308DF9561" box="[846,1009,1314,1340]" pageId="89" pageNumber="90" value="1961-09-21">21 Sep. 1961</date>
, C.E. Yunker leg. (
<collectionCode id="4994AE28FFCDFFE40BC709030E069561" box="[1244,1320,1314,1340]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="89" pageNumber="90">CNCI</collectionCode>
); 1 ♂, Canal Zone, Skunk Hollow, 
<date id="5B3B102DFFCDFFE40D13096709B4953C" box="[520,666,1350,1377]" pageId="89" pageNumber="90" value="1975-07-13">13 Jul. 1975</date>
, B.C. Ratcliffe leg. (
<collectionCode id="4994AE28FFCDFFE40C92096708CA953D" box="[905,996,1350,1376]" pageId="89" pageNumber="90">ZMHB</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40CE309670FA5953D" box="[1016,1163,1350,1377]" pageId="89" pageNumber="90" type="generic">5 specimens</specimenCount>
, Canal Zone, Skunk Hollow, 
<quantity id="E87D9B08FFCDFFE40E3F094B0A4D95D9" box="[292,355,1386,1412]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="89" pageNumber="90" unit="mi" value="6.0">6 mi.</quantity>
NW of Gatún Lake, 
<date id="5B3B102DFFCDFFE40D7D094A081B95D8" box="[614,821,1387,1413]" pageId="89" pageNumber="90" value="1980-05-17" valueMax="1980-05-31" valueMin="1980-05-17">17–31 May 1980</date>
, B.C. Ratcliffe leg. (
<collectionCode id="4994AE28FFCDFFE40B2B094A0FA195D8" box="[1072,1167,1387,1413]" pageId="89" pageNumber="90">BMNH</collectionCode>
); 4 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40BE0094A0E0795D8" box="[1275,1321,1387,1413]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, same collecting data as for preceding (
<collectionCode id="4994AE28FFCDFFE40D5E09AE098E95F4" box="[581,672,1423,1449]" pageId="89" pageNumber="90">ZMHB</collectionCode>
); 2 ♂♂, Cerro Azul, 
<date id="5B3B102DFFCDFFE40C8609AE0F4595F4" box="[925,1131,1423,1449]" pageId="89" pageNumber="90" value="1996-05-12" valueMax="1996-05-13" valueMin="1996-05-12">12–13 May 1996</date>
, Wappes, Huether and Morris leg. (
<collectionCode id="4994AE28FFCDFFE40E5409920A849590" box="[335,426,1459,1485]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40EA40992092C9593" box="[447,514,1459,1486]" pageId="89" pageNumber="90" type="female">2 ♀♀</specimenCount>
, Cerro Azul, 
<date id="5B3B102DFFCDFFE40DBA099208439590" box="[673,877,1459,1486]" pageId="89" pageNumber="90" value="1996-05-21" valueMax="1996-05-24" valueMin="1996-05-21">21–24 May 1996</date>
, Wappes, Huether and Morris leg. (
<collectionCode id="4994AE28FFCDFFE40A1409920E449590" box="[1295,1386,1459,1485]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40FA609F60BC595AF" box="[189,235,1495,1522]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, El Llano-Carti Road, Km 7.5, 
<quantity id="E87D9B08FFCDFFE40D7709F6099595AF" box="[620,699,1495,1522]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="89" pageNumber="90" unit="m" value="350.0">350 m</quantity>
, 
<date id="5B3B102DFFCDFFE40DD109F6085895AF" box="[714,886,1495,1522]" pageId="89" pageNumber="90" value="1995-06-04" valueMax="1995-06-06" valueMin="1995-06-04">4–6 Jun. 1995</date>
, flight interception trap, A.R. Gillogly leg. (
<collectionCode id="4994AE28FFCDFFE40FDE09DD0A0E964B" box="[197,288,1532,1558]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FFCDFFE40E6D09DD0A8B964B" box="[374,421,1532,1558]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, El Llano-Carti Road, Km 8–13, 
<date id="5B3B102DFFCDFFE40C2509DD0F209648" box="[830,1038,1531,1558]" pageId="89" pageNumber="90" value="1996-05-21" valueMax="1996-05-24" valueMin="1996-05-21">21–24 May 1996</date>
, Wappes, Huether and Morris leg. (
<collectionCode id="4994AE28FFCDFFE40FE30A010A7D9667" box="[248,339,1568,1594]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40E7D0A010ABC9667" box="[358,402,1568,1594]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Nusagandi, Ina Igar (Trail), 
<date id="5B3B102DFFCDFFE40DF00A0108989667" box="[747,950,1568,1594]" pageId="89" pageNumber="90" value="1993-05-18" valueMax="1993-05-21" valueMin="1993-05-18">18–21 May 1993</date>
, pitfall with human faeces, E.G. Riley leg. (
<collectionCode id="4994AE28FFCDFFE40FE30A650A7E9603" box="[248,336,1604,1630]" pageId="89" pageNumber="90">CEMT</collectionCode>
); 19 ♂♂ (1 dissected), 
<specimenCount id="3983FD64FFCDFFE40D790A6509889602" box="[610,678,1604,1631]" pageId="89" pageNumber="90" type="female">9 ♀♀</specimenCount>
, same collecting data as for preceding (
<collectionCode id="4994AE28FFCDFFE40B610A650FFB9603" box="[1146,1237,1604,1630]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40BF20A650E389602" box="[1257,1302,1604,1631]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Parque nacional Soberanía, 
<date id="5B3B102DFFCDFFE40EB60A49095796DF" box="[429,633,1640,1667]" pageId="89" pageNumber="90" value="1996-05-23" valueMax="1996-05-27" valueMin="1996-05-23">23–27 May 1996</date>
, Wappes, Huether and Morris leg. (
<collectionCode id="4994AE28FFCDFFE40B060A490F5696DF" box="[1053,1144,1640,1666]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 
<specimenCount id="3983FD64FFCDFFE40B960A490F9496DE" box="[1165,1210,1640,1667]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Pipeline Road, Km 01–12, 
<date id="5B3B102DFFCDFFE40E500AAD093D96FA" box="[331,531,1676,1703]" pageId="89" pageNumber="90" value="1997-06-26" valueMax="1997-06-30" valueMin="1997-06-26">26–30 Jun. 1997</date>
, Wappes and Morris leg. (
<collectionCode id="4994AE28FFCDFFE40C550AAC088896FA" box="[846,934,1677,1703]" pageId="89" pageNumber="90">CEMT</collectionCode>
); 3 ♂♂, 
<specimenCount id="3983FD64FFCDFFE40B140AAC0F7A96FA" box="[1039,1108,1677,1703]" pageId="89" pageNumber="90" type="female">3 ♀♀</specimenCount>
, same collecting data as for preceding (
<collectionCode id="4994AE28FFCDFFE40E730A900AED9696" box="[360,451,1713,1739]" pageId="89" pageNumber="90">TAMU</collectionCode>
); 1 ♂, 
<specimenCount id="3983FD64FFCDFFE40D0B0A9009159696" box="[528,571,1713,1739]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
, Pipline Road, Km 6.1, “near Gambba”, 
<quantity id="E87D9B08FFCDFFE40B020A900F789696" box="[1049,1110,1713,1739]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="89" pageNumber="90" unit="m" value="40.0">40 m</quantity>
, 
<date id="5B3B102DFFCDFFE40B790A900E399691" box="[1122,1303,1713,1740]" pageId="89" pageNumber="90" value="1995-06-07" valueMax="1995-06-21" valueMin="1995-06-07">7–21 Jun. 1995</date>
, J. Ashe and R. Brooks leg. (
<collectionCode id="4994AE28FFCDFFE40EB30AF4092A96B2" box="[424,516,1749,1775]" country="Brazil" httpUri="http://grbio.org/cool/wyk1-wacz" name="Porto Alegre, Museu de Ciencias Naturais da Fundacao Zoo-Botanica do Rio Grande do Sul" pageId="89" pageNumber="90">MCNZ</collectionCode>
).
</paragraph>
<paragraph id="2F3A36EDFFCDFFE40FA60B390E0F976F" blockId="89.[189,1313,1814,1842]" box="[189,1313,1814,1842]" pageId="89" pageNumber="90">
No data: 1 ♂, 
<specimenCount id="3983FD64FFCDFFE40E780B390AA1976F" box="[355,399,1816,1842]" pageId="89" pageNumber="90" type="female">1 ♀</specimenCount>
(
<collectionCode id="4994AE28FFCDFFE40E840B390AC8976F" box="[415,486,1816,1842]" pageId="89" pageNumber="90">ISNB</collectionCode>
, Gillet collection), 
<specimenCount id="3983FD64FFCDFFE40DD30B390824976F" box="[712,778,1816,1842]" pageId="89" pageNumber="90" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="4994AE28FFCDFFE40C010B39084F976F" box="[794,865,1816,1842]" pageId="89" pageNumber="90">ISNB</collectionCode>
, J. Thomson collection, pseudotypes).
</paragraph>
</subSubSection>
<subSubSection id="679F6566FFCDFFE10FA60B7B0F349131" lastPageId="92" lastPageNumber="93" pageId="89" pageNumber="90" type="description">
<paragraph id="2F3A36EDFFCDFFE40FA60B7B0A459729" blockId="89.[189,363,1882,1908]" box="[189,363,1882,1908]" pageId="89" pageNumber="90">
<heading id="74728181FFCDFFE40FA60B7B0A459729" bold="true" box="[189,363,1882,1908]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="1DF1EAFFFFCDFFE40FA60B7B0A459729" bold="true" box="[189,363,1882,1908]" pageId="89" pageNumber="90">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFCDFFE40FA60BAB0F6997B1" blockId="89.[189,1399,1930,2029]" pageId="89" pageNumber="90">COLOURATION. Entire body surface bright and glossy, including the surface of pygidium. Head, pronotum, elytra, and pygidium usually purple, blue, or, occasionally, green. Metaventrite dark green or dark blue. Meso- and metafemora dark brown and, in general, with greenish reflections.</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA60D2D085691E7" blockId="90.[189,1399,267,442]" pageId="90" pageNumber="91">HEAD. Tegument entirely covered by alveolar microsculpture and without any trace of micropunctation or, occasionally, with very weak, almost imperceptible micropunctures among microsculpture. Clypeus with two apical teeth obtuse and contiguous at base; with a single transverse row of very short setae covering the base of both teeth. Genae with evident tooth immediately behind clypeal-genal juncture. Posterior edge of head with complete margin between eyes.</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA60DC20AB79314" blockId="90.[189,1399,482,842]" pageId="90" pageNumber="91">
THORAX. Pronotum very shiny and with tegument at centre ranging from completely smooth, with no trace of microsculpture or micropunctation, to with very weak and effaced alveolar microsculpture and micropunctation; on the sides, always with fine alveolar microsculpture. Posterior edge, in general, without transverse line at centre; in some specimens, with traces of fine transverse line at centre. Hypomeral cavity with long setae on posterior region and as long or shorter setae at centre and on external edge; external edge with a turbercle which can be clearly present or almost imperceptible. Metaventrite entirely glabrous (or, occasionally, with few setae at lateral region close to metacoxae, 
<figureCitation id="B7BE2A68FFCEFFE70BD70EE10E029287" box="[1228,1324,704,730]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,372,1844]" captionTargetId="figure@21.[189,1398,372,1849]" captionTargetPageId="21" captionText="Fig. 7. Pterothorax, ventrolateral views. A. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov. B. S. obscurus (Schmidt, 1920). C. Canthon xanthopus Blanchard, 1846. Abbreviations: MsC = mesocoxa; MMsC = external margin of mesocoxa; CMtE = centre of metaventrite; LMtE = lateral region of metaventrite; MtC = metacoxa." figureDoi="http://doi.org/10.5281/zenodo.3824793" httpUri="https://zenodo.org/record/3824793/files/figure.png" pageId="90" pageNumber="91">Fig. 7A</figureCitation>
); with uniform alveolar microsculpture throughout ventral region of metaventrite (i.e., between metacoxae) and with very tenuous micropunctation, which, in general, is only visible under high magnification (higher than 35 ×).
</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA60F52098A9484" blockId="90.[189,1399,882,1241]" pageId="90" pageNumber="91">
LEGS. Ventral surface of all femora bright and with silky appearance. Profemora with tegument completely covered by alveolar microsculpture. Protibiae broad and without internal expansion; at their apical half, external edge with three medium-sized, broad and acute or obstuse teeth – the two apical much larger than basal ones (
<figureCitation id="B7BE2A68FFCEFFE70D610FC009C793A6" box="[634,745,993,1019]" captionStart="Fig" captionStartId="25.[189,232,1787,1813]" captionTargetBox="[196,1391,645,1731]" captionTargetId="figure@25.[189,1399,645,1755]" captionTargetPageId="25" captionText="Fig. 11. Morphological diversity in the protibiae of Sylvicanthon Halffter &amp; Martínez, 1977. A. S. enkerlini (Martínez et al., 1964) comb. nov. B. S. aequinoctialis (Harold, 1869) comb. nov. C. S. securus (Schmidt, 1920) comb. nov. D. S. obscurus (Schmidt, 1920). E. S. furvus (Schmidt, 1920). F. S. bridarollii (Martínez, 1949) (southern populations). G. S. bridarollii (northern populations). H. S. attenboroughi sp. nov. I. S. edmondsi sp. nov. J. S. candezei (Harold, 1869). Note the variation related to the presence and degree of development of an expansion on the internal edge of protibiae and to the number, size and position of the external teeth." figureDoi="http://doi.org/10.5281/zenodo.3824801" httpUri="https://zenodo.org/record/3824801/files/figure.png" pageId="90" pageNumber="91">Fig. 11B</figureCitation>
). Mesofemora margined anteriorly only at basal half; unmargined portion of anterior edge with row of very short setae; tegument with rivose microsculpture at external (anterior) half and with alveolar microsculpture at internal (posterior) half. Metafemora with both anterior and posterior margins (
<figureCitation id="B7BE2A68FFCEFFE70D7E087109FF9437" box="[613,721,1104,1130]" captionStart="Fig" captionStartId="93.[189,232,1893,1919]" captionTargetBox="[189,1399,1019,1850]" captionTargetId="figure@93.[189,1399,1019,1850]" captionTargetPageId="93" captionText="Fig. 31. Metafemora of the two species of the aequinoctialis subgroup. A. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov. B. S. proseni (Martínez, 1949) stat. et comb. nov. Arrow indicates the basal end of the posterior margin of the ventral face of metafemora (note that it is longer in S. aequinoctialis comb. nov. than in S. proseni stat. et comb. nov. and may exceed the trochanter in the first species)." figureDoi="http://doi.org/10.5281/zenodo.3824843" httpUri="https://zenodo.org/record/3824843/files/figure.png" pageId="90" pageNumber="91">Fig. 31A</figureCitation>
); posterior margin extending from the apex of femur until, at most, little beyond the trochanter; tegument as on mesofemora, with no trace of micropunctation or coarse punctation at base. Metatarsomeres II and V subequal in length and longer than the others; metatarsomere IV shorter than the others.
</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA60923084295ED" blockId="90.[189,1399,1282,1456]" pageId="90" pageNumber="91">ELYTRA. With only eight very subtly-marked, fine and superficial striae, all of which without carinulae; first seven striae equally marked (only seventh stria, occasionally, more effaced and discontinuous); eighth stria present only on humerus and subtly carinate (humeral carina). Tegument bright with alveolar microsculpture and micropunctation very subtle and effaced, only visible under magnification of at least 30 ×; microsculpture more evident only on the outer sides.</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA609F808709660" blockId="90.[189,1399,1497,1597]" pageId="90" pageNumber="91">ABDOMEN. Ventrite V and, especially, ventrite VI smooth at centre and with weaker rivose microsculpture on the sides. Lateral foveae absent in both sexes. Pygidium with shiny tegument and very diffuse effaced rivose microsculpture; of equivalent length in both sexes.</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA60A470A7096B2" blockId="90.[189,1399,1638,1775]" pageId="90" pageNumber="91">AEDEAGUS. Parameres long, slightly shorter than phallobase, and clearly asymmetrical: left paramere laterally with apical depression more profound and wide than depression on left paramere (difference more easily seen in dorsal view). In lateral view, parameres with central angularity and without ventral keel or notch.</paragraph>
<paragraph id="2F3A36EDFFCEFFE70FA60B390AD397B6" blockId="90.[189,1399,1816,2027]" pageId="90" pageNumber="91">
SEXUAL DIMORPHISM. 
<emphasis id="1DF1EAFFFFCEFFE70EAA0B390AD2976F" bold="true" box="[433,508,1816,1842]" pageId="90" pageNumber="91">Males</emphasis>
: Protibial spur wide and foliaceous, external edge projected into an acute long spine, while internal edge has no prolongation or has a very short spine; area between both spines straight or slightly excavated (
<figureCitation id="B7BE2A68FFCEFFE70D360B4309B49720" box="[557,666,1890,1917]" captionStart="Fig" captionStartId="30.[189,232,1790,1816]" captionTargetBox="[196,1383,292,1753]" captionTargetId="figure@30.[196,1383,289,1754]" captionTargetPageId="30" captionText="Fig. 15. Morphological variation and sexual dimorphism in the protibial spur of Sylvicanthon. A. S. enkerlini (Martínez et al., 1964) comb. nov., ♂. B. S. foveiventris (Schmidt, 1920), ♂. C. S. candezei (Harold, 1869), ♂. D. S. genieri sp. nov., ♂. E. S. aequinoctialis (Harold, 1868), ♂. F. S. aequinoctialis comb. nov., ♀. G. S. proseni (Martínez, 1949) stat. et comb. nov., ♂. H. S. proseni stat. et comb. nov., ♀. I. S. bridarollii (Martínez, 1949), ♂. J. S. seag sp. nov., ♂. K. S. edmondsi sp. nov.,♂. L. S. attenboroughi sp. nov., ♂. M. S. obscurus (Schmidt, 1920), ♂. N. S. furvus (Schmidt, 1920), ♂. O. S. securus (Schmidt, 1920) comb. nov., ♂." figureDoi="http://doi.org/10.5281/zenodo.3824809" httpUri="https://zenodo.org/record/3824809/files/figure.png" pageId="90" pageNumber="91">Fig. 15E</figureCitation>
). Ventrite VI strongly narrowed at centre; ventrite V usually without medial flange or only slightly indicated over anterior edge of ventrite VI. 
<emphasis id="1DF1EAFFFFCEFFE70B820BA60E2E97FC" bold="true" box="[1177,1280,1927,1953]" pageId="90" pageNumber="91">Females</emphasis>
: Protibial spur spiniform (
<figureCitation id="B7BE2A68FFCEFFE70E600B8D0ACD979A" box="[379,483,1964,1991]" captionStart="Fig" captionStartId="30.[189,232,1790,1816]" captionTargetBox="[196,1383,292,1753]" captionTargetId="figure@30.[196,1383,289,1754]" captionTargetPageId="30" captionText="Fig. 15. Morphological variation and sexual dimorphism in the protibial spur of Sylvicanthon. A. S. enkerlini (Martínez et al., 1964) comb. nov., ♂. B. S. foveiventris (Schmidt, 1920), ♂. C. S. candezei (Harold, 1869), ♂. D. S. genieri sp. nov., ♂. E. S. aequinoctialis (Harold, 1868), ♂. F. S. aequinoctialis comb. nov., ♀. G. S. proseni (Martínez, 1949) stat. et comb. nov., ♂. H. S. proseni stat. et comb. nov., ♀. I. S. bridarollii (Martínez, 1949), ♂. J. S. seag sp. nov., ♂. K. S. edmondsi sp. nov.,♂. L. S. attenboroughi sp. nov., ♂. M. S. obscurus (Schmidt, 1920), ♂. N. S. furvus (Schmidt, 1920), ♂. O. S. securus (Schmidt, 1920) comb. nov., ♂." figureDoi="http://doi.org/10.5281/zenodo.3824809" httpUri="https://zenodo.org/record/3824809/files/figure.png" pageId="90" pageNumber="91">Fig. 15F</figureCitation>
). Ventrite VI broad at centre and with anterior edge slightly covered by weak medial flange of ventrite V.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60D2A0A5B9178" blockId="91.[189,373,267,293]" box="[189,373,267,293]" pageId="91" pageNumber="92">
<heading id="74728181FFCFFFE60FA60D2A0A5B9178" bold="true" box="[189,373,267,293]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="1DF1EAFFFFCFFFE60FA60D2A0A5B9178" bold="true" box="[189,373,267,293]" pageId="91" pageNumber="92">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60D1B08A991C1" blockId="91.[189,1399,314,413]" pageId="91" pageNumber="92">
Males (N = 20). 
<emphasis id="1DF1EAFFFFCFFFE60E930D1B0A979109" bold="true" box="[392,441,314,340]" pageId="91" pageNumber="92">TL:</emphasis>
AV: 10.5 ± 0.93; 
<collectingCountry id="5792767DFFCFFFE60D880D1A09E99108" box="[659,711,315,341]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 12.3; MN: 9.0. 
<emphasis id="1DF1EAFFFFCFFFE60C950D1B08E99109" bold="true" box="[910,967,314,340]" pageId="91" pageNumber="92">EW:</emphasis>
AV: 7.1 ± 0.66; 
<collectingCountry id="5792767DFFCFFFE60B880D1A0FE99108" box="[1171,1223,315,341]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 7.8; MN: 5.7. 
<emphasis id="1DF1EAFFFFCFFFE60FA60D7F0BC39125" bold="true" box="[189,237,350,376]" pageId="91" pageNumber="92">PL:</emphasis>
AV: 3.4 ± 0.3; 
<collectingCountry id="5792767DFFCFFFE60EBE0D7E0AF79124" box="[421,473,351,377]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 3.9; MN: 2.7. 
<emphasis id="1DF1EAFFFFCFFFE60D940D7F09EB9125" bold="true" box="[655,709,350,376]" pageId="91" pageNumber="92">PW:</emphasis>
AV: 6.3 ± 0.83; 
<collectingCountry id="5792767DFFCFFFE60C950D7E08EC9124" box="[910,962,351,377]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 7.4; MN: 5.3. 
<emphasis id="1DF1EAFFFFCFFFE60B620D7F0F989125" bold="true" box="[1145,1206,350,376]" pageId="91" pageNumber="92">PgL:</emphasis>
AV: 1.9 ± 0.16; 
<collectingCountry id="5792767DFFCFFFE60FA60DA20BDF91C0" box="[189,241,387,413]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 2.1; MN: 1.6. 
<emphasis id="1DF1EAFFFFCFFFE60EB90DA30AC691C1" bold="true" box="[418,488,386,412]" pageId="91" pageNumber="92">PgW:</emphasis>
AV: 3.4 ± 0.29; 
<collectingCountry id="5792767DFFCFFFE60DB10DA209F091C0" box="[682,734,387,413]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 3.7; MN: 2.8.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60DE508A8927A" blockId="91.[189,1399,452,551]" pageId="91" pageNumber="92">
Females (N = 21). 
<emphasis id="1DF1EAFFFFCFFFE60EBA0DE50AFC9183" bold="true" box="[417,466,452,478]" pageId="91" pageNumber="92">CT:</emphasis>
AV: 10.4 ± 0.88; 
<collectingCountry id="5792767DFFCFFFE60DB00DE409F19182" box="[683,735,453,479]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 11.8; MN: 8.7. 
<emphasis id="1DF1EAFFFFCFFFE60CBE0DE408F09182" bold="true" box="[933,990,453,479]" pageId="91" pageNumber="92">EW:</emphasis>
AV: 7.0 ± 0.51; 
<collectingCountry id="5792767DFFCFFFE60BB20DE40FF39182" box="[1193,1245,453,479]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 7.6; MN: 6. 
<emphasis id="1DF1EAFFFFCFFFE60FA60DC80BC3925E" bold="true" box="[189,237,489,515]" pageId="91" pageNumber="92">PL:</emphasis>
AV: 3.4 ± 0.29; 
<collectingCountry id="5792767DFFCFFFE60EAA0DC80ACB925E" box="[433,485,489,515]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 3.9; MN: 2.9. 
<emphasis id="1DF1EAFFFFCFFFE60D820DC809FE925E" bold="true" box="[665,720,489,515]" pageId="91" pageNumber="92">PW:</emphasis>
AV: 6.3 ± 0.47; 
<collectingCountry id="5792767DFFCFFFE60C8E0DC808E7925E" box="[917,969,489,515]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 5.3; MN: 6.8. 
<emphasis id="1DF1EAFFFFCFFFE60B660DC80F95925E" bold="true" box="[1149,1211,489,515]" pageId="91" pageNumber="92">PgL:</emphasis>
AV: 1.7 ± 0.14; 
<collectingCountry id="5792767DFFCFFFE60FA60E2C0BDF927A" box="[189,241,525,551]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 2.0; MN: 1.5. 
<emphasis id="1DF1EAFFFFCFFFE60EB90E2C0AC6927A" bold="true" box="[418,488,525,551]" pageId="91" pageNumber="92">PgW:</emphasis>
AV: 3.3 ± 0.27; 
<collectingCountry id="5792767DFFCFFFE60DB30E2C09F2927A" box="[680,732,525,551]" name="Mexico" pageId="91" pageNumber="92">MX</collectingCountry>
: 3.7; MN: 2.8.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60E6E092C9234" blockId="91.[189,514,591,617]" box="[189,514,591,617]" pageId="91" pageNumber="92">
<heading id="74728181FFCFFFE60FA60E6E092C9234" bold="true" box="[189,514,591,617]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="1DF1EAFFFFCFFFE60FA60E6E092C9234" bold="true" box="[189,514,591,617]" pageId="91" pageNumber="92">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60E5F0F5992C5" blockId="91.[189,1143,638,665]" box="[189,1143,638,665]" pageId="91" pageNumber="92">
Widespread throughout the tropical forests from 
<collectingCountry id="5792767DFFCFFFE60DE20E5F084592C5" box="[761,875,638,664]" name="Honduras" pageId="91" pageNumber="92">Honduras</collectingCountry>
to northern 
<collectingCountry id="5792767DFFCFFFE60CE70E5F0F5A92C5" box="[1020,1140,638,664]" name="Colombia" pageId="91" pageNumber="92">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60EE10A699287" blockId="91.[189,327,704,730]" box="[189,327,704,730]" pageId="91" pageNumber="92">
<heading id="74728181FFCFFFE60FA60EE10A699287" bold="true" box="[189,327,704,730]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="1DF1EAFFFFCFFFE60FA60EE10A699287" bold="true" box="[189,327,704,730]" pageId="91" pageNumber="92">Ecoregions</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60ED10F12932B" blockId="91.[189,1398,752,886]" pageId="91" pageNumber="92">
Central American Atlantic Moist Forests, Central American Montane Forests, Isthmian-Atlantic Moist Forests, Talamancan Montane Forests, Costa Rican Seasonal Moist Forests, Isthmian-Pacific Moist Forests, Chocó-Darién Moist Forests, Guajira-Barranquilla Xeric Scrub, Magdalena-Urabá Moist Forests, 
<collectingRegion id="ED41F80FFFCFFFE60E3B0F7D0A8C932B" box="[288,418,860,886]" country="Colombia" name="Magdalena" pageId="91" pageNumber="92">Magdalena</collectingRegion>
Valley Montane Forests, 
<collectingRegion id="ED41F80FFFCFFFE60DD40F7D087F932B" box="[719,849,860,886]" country="Colombia" name="Magdalena" pageId="91" pageNumber="92">Magdalena</collectingRegion>
Valley Dry Forests.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60FBF0AC993E5" blockId="91.[189,487,926,952]" box="[189,487,926,952]" pageId="91" pageNumber="92">
<emphasis id="1DF1EAFFFFCFFFE60FA60FBF0A5793E5" bold="true" box="[189,377,926,952]" pageId="91" pageNumber="92">Collecting sites</emphasis>
(
<figureCitation id="B7BE2A68FFCFFFE60E920FBF0AF193E5" box="[393,479,926,952]" captionStart="Fig" captionStartId="92.[189,232,2000,2026]" captionTargetBox="[189,1399,1037,1965]" captionTargetId="figure@92.[189,1399,1037,1965]" captionTargetPageId="92" captionText="Fig. 30. Distribution of the two species of the aequinoctialis subgroup." figureDoi="http://doi.org/10.5281/zenodo.3824841" httpUri="https://zenodo.org/record/3824841/files/figure.png" pageId="91" pageNumber="92">Fig. 30</figureCitation>
)
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60FEF094693BA" blockId="91.[189,616,973,1000]" box="[189,616,973,1000]" pageId="91" pageNumber="92">
<collectingCountry id="5792767DFFCFFFE60FA60FEF0A4A93B5" box="[189,356,974,1000]" name="Honduras" pageId="91" pageNumber="92">HONDURAS</collectingCountry>
. 
<emphasis id="1DF1EAFFFFCFFFE60E740FEC0AC893B5" bold="true" box="[367,486,973,1000]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60E740FEC0AC893B5" box="[367,486,973,1000]" country="Honduras" name="Atlantida" pageId="91" pageNumber="92">Atlántida</collectingRegion>
</emphasis>
: La Ceiba.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA608310A68942F" blockId="91.[189,1398,1040,1138]" pageId="91" pageNumber="92">
<collectingCountry id="5792767DFFCFFFE60FA608310A5F9477" box="[189,369,1040,1066]" name="Nicaragua" pageId="91" pageNumber="92">NICARAGUA</collectingCountry>
. 
<emphasis id="1DF1EAFFFFCFFFE60E6208310ACD9477" bold="true" box="[377,483,1040,1066]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60E6208310ACD9477" box="[377,483,1040,1066]" country="Nicaragua" name="Jinotega" pageId="91" pageNumber="92">Jinotega</collectingRegion>
</emphasis>
: Reserva Natural Cerro Kilambé, Santa Maura. 
<emphasis id="1DF1EAFFFFCFFFE60B1108310FBF9477" bold="true" box="[1034,1169,1040,1066]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60B1108310FBF9477" box="[1034,1169,1040,1066]" country="Nicaragua" name="Matagalpa" pageId="91" pageNumber="92">Matagalpa</collectingRegion>
</emphasis>
: Matiguas, Reserva Natural Biósfera de Bosawas (Macizo de Peñas Blancas). 
<emphasis id="1DF1EAFFFFCFFFE60C7B081508F39413" bold="true" box="[864,989,1076,1102]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60C7B081508F39413" box="[864,989,1076,1102]" country="Nicaragua" name="Chontales" pageId="91" pageNumber="92">Chontales</collectingRegion>
</emphasis>
. 
<emphasis id="1DF1EAFFFFCFFFE60CF008120FBF9413" bold="true" box="[1003,1169,1075,1102]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60CF008120FBF9413" box="[1003,1169,1075,1102]" country="Nicaragua" name="Rio San Juan" pageId="91" pageNumber="92">Río San Juan</collectingRegion>
</emphasis>
: Reserva Biológica Indio Maíz.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA608BB0E5F95D0" blockId="91.[189,1399,1178,1421]" pageId="91" pageNumber="92">
<collectingCountry id="5792767DFFCFFFE60FA608BB0A4094E9" box="[189,366,1178,1204]" name="Costa Rica" pageId="91" pageNumber="92">COSTA RICA</collectingCountry>
. 
<emphasis id="1DF1EAFFFFCFFFE60E6708BB092194E9" bold="true" box="[380,527,1178,1204]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60E6708BB092194E9" box="[380,527,1178,1204]" country="Costa Rica" name="Guanacaste" pageId="91" pageNumber="92">Guanacaste</collectingRegion>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60D3908BA09CB94E9" box="[546,741,1178,1205]" italics="true" pageId="91" pageNumber="92">Cerro El Hacha</emphasis>
, Volcán Rincón de la Vieja. 
<emphasis id="1DF1EAFFFFCFFFE60B5208BB08F49484" italics="true" pageId="91" pageNumber="92">
<emphasis id="1DF1EAFFFFCFFFE60B5208BB0F9E94E8" bold="true" box="[1097,1200,1178,1205]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60B5208BB0F9E94E8" box="[1097,1200,1178,1205]" country="Costa Rica" name="Alajuela" pageId="91" pageNumber="92">Alajuela</collectingRegion>
</emphasis>
: Playuelas, San Ramón de dos Ríos, Volcán Tenorio (Colonia Río Celeste). 
<emphasis id="1DF1EAFFFFCFFFE60C63089F08F49484" bold="true" box="[888,986,1214,1241]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60C63089F08F49484" box="[888,986,1214,1241]" country="Costa Rica" name="Heredia" pageId="91" pageNumber="92">Heredia</collectingRegion>
</emphasis>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60CF1089E0A2094A1" italics="true" pageId="91" pageNumber="92">Sarapiquí (Estación Biológica La Selva).</emphasis>
<emphasis id="1DF1EAFFFFCFFFE60E0E08C30A8A94A1" bold="true" box="[277,420,1250,1276]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60E0E08C30A8A94A1" box="[277,420,1250,1276]" country="Costa Rica" name="Puntarenas" pageId="91" pageNumber="92">Puntarenas</collectingRegion>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60EA808C3098994A0" box="[435,679,1250,1277]" italics="true" pageId="91" pageNumber="92">Golfitos (Las Torres)</emphasis>
, Osa (
<emphasis id="1DF1EAFFFFCFFFE60DEE08C30FB594A0" box="[757,1179,1250,1277]" italics="true" pageId="91" pageNumber="92">Albergue Ecoturístico Cerro de Oro</emphasis>
), Parque Nacional 
<taxonomicName id="E8854D6EFFCFFFE60FA6092709B7957C" authority="(Estacion Biologica Sirena)" baseAuthorityName="Estacion Biologica Sirena" box="[189,665,1286,1313]" class="Insecta" family="Cerambycidae" genus="Corcovado" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">Corcovado (Estación Biológica Sirena)</taxonomicName>
, Reserva Biológica Bosque Nuboso Monteverde. 
<emphasis id="1DF1EAFFFFCFFFE60BE609270E41957D" bold="true" box="[1277,1391,1286,1312]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60BE609270E41957D" box="[1277,1391,1286,1312]" country="Costa Rica" name="San Jose" pageId="91" pageNumber="92">San José</collectingRegion>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60FA6090A09BB9518" box="[189,661,1322,1349]" italics="true" pageId="91" pageNumber="92">Santa Ana (Ciudad Colon), Turrúcares.</emphasis>
<emphasis id="1DF1EAFFFFCFFFE60D85090809DF9519" bold="true" box="[670,753,1321,1348]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60D85090809DF9519" box="[670,753,1321,1348]" country="Costa Rica" name="Limon" pageId="91" pageNumber="92">Limón</collectingRegion>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60C19090A0E4495D1" italics="true" pageId="91" pageNumber="92">Guácimo (Parque Nacional Tortuguero), Limón (Río Blanco), Pococí (Cedrales de la Rita, Cerro Cocorí, Refugio Nacional de Fauna Silvestre Barra del Colorado), Refugio de Vida Silvestre Gandoca-Manzanillo (Manzanillo), Talamanca (Bratsi: Amubri)</emphasis>
.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA6099408CD964B" blockId="91.[189,1398,1460,1559]" pageId="91" pageNumber="92">
<collectingCountry id="5792767DFFCFFFE60FA609940A139592" box="[189,317,1461,1487]" name="Panama" pageId="91" pageNumber="92">PANAMA</collectingCountry>
. 
<emphasis id="1DF1EAFFFFCFFFE60E5309950A9A9592" bold="true" box="[328,436,1460,1487]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60E5309950A9A9592" box="[328,436,1460,1487]" country="Panama" name="Chiriqui" pageId="91" pageNumber="92">Chiriquí</collectingRegion>
</emphasis>
: 
<taxonomicName id="E8854D6EFFCFFFE60ED9099409F79593" authority=", David" authorityName="David" box="[450,729,1460,1487]" class="Reptilia" family="Crocodylidae" genus="Baru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="91" pageNumber="92" phylum="Chordata" rank="subGenus" subGenus="Progreso">
<emphasis id="1DF1EAFFFFCFFFE60ED9099409AD9592" box="[450,643,1461,1487]" italics="true" pageId="91" pageNumber="92">Barú (Progreso)</emphasis>
, David
</taxonomicName>
(San José de David), Gualaca (Hornito), 
<emphasis id="1DF1EAFFFFCFFFE60BA709950E299592" bold="true" box="[1212,1287,1460,1487]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60BA709950E299592" box="[1212,1287,1460,1487]" country="Panama" name="Colon" pageId="91" pageNumber="92">Colón</collectingRegion>
</emphasis>
: Chagres (Fort San Lorenzo), Colón. 
<emphasis id="1DF1EAFFFFCFFFE60D1609F9095A95AE" bold="true" box="[525,628,1496,1523]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60D1609F9095A95AE" box="[525,628,1496,1523]" country="Panama" name="Panama" pageId="91" pageNumber="92">Panamá</collectingRegion>
</emphasis>
: Panama Canal (Fort Kobbe; Lago Gatún: Isla Barro Colorado; Skunk Hollow), Cerro Azul, Nusagandi, Parque Nacional Soberanía.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60A1E0F7D96B4" blockId="91.[189,1398,1598,1770]" pageId="91" pageNumber="92">
<collectingCountry id="5792767DFFCFFFE60FA60A1E0A709604" box="[189,350,1599,1625]" name="Colombia" pageId="91" pageNumber="92">COLOMBIA</collectingCountry>
: 
<emphasis id="1DF1EAFFFFCFFFE60E720A1F0AF69604" bold="true" box="[361,472,1598,1625]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60E720A1F0AF69604" box="[361,472,1598,1625]" country="Colombia" name="Atlantico" pageId="91" pageNumber="92">Atlántico</collectingRegion>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60EF30A6108CE9604" box="[488,992,1599,1625]" italics="true" pageId="91" pageNumber="92">Juan de Acosta (Reserva de Tierra Arena).</emphasis>
<emphasis id="1DF1EAFFFFCFFFE60CFC0A1E0F1E9604" bold="true" box="[999,1072,1599,1625]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60CFC0A1E0F1E9604" box="[999,1072,1599,1625]" country="Colombia" name="Cesar" pageId="91" pageNumber="92">Cesar</collectingRegion>
</emphasis>
: Chimichagua. 
<emphasis id="1DF1EAFFFFCFFFE60BF70A1E0E409604" bold="true" box="[1260,1390,1599,1625]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60BF70A1E0E409604" box="[1260,1390,1599,1625]" country="Colombia" name="Santander" pageId="91" pageNumber="92">Santander</collectingRegion>
</emphasis>
: Serranía de las Quinchas (Reserva El Paujil). 
<emphasis id="1DF1EAFFFFCFFFE60DDB0A4208109620" bold="true" box="[704,830,1635,1661]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60DDB0A4208109620" box="[704,830,1635,1661]" country="Colombia" name="Antioquia" pageId="91" pageNumber="92">Antioquia</collectingRegion>
</emphasis>
: San Luis (Sonsón). 
<emphasis id="1DF1EAFFFFCFFFE60B320A430F579620" bold="true" box="[1065,1145,1634,1661]" pageId="91" pageNumber="92">Chocó</emphasis>
:Acandí (Capurganá), Lloró, Parque Nacional Natural Ensenada de Utría, Quibdó (Tutunendó, Estación Ambiental Pandó, Pacurita), Unión Panamericana (Salero). 
<emphasis id="1DF1EAFFFFCFFFE60DAB0A8B082D9698" bold="true" box="[688,771,1706,1733]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60DAB0A8B082D9698" box="[688,771,1706,1733]" country="Colombia" name="Caldas" pageId="91" pageNumber="92">Caldas</collectingRegion>
</emphasis>
: 
<emphasis id="1DF1EAFFFFCFFFE60C0D0A8D0FD29698" box="[790,1276,1707,1733]" italics="true" pageId="91" pageNumber="92">Norcasia (Reserva Natural Río Manso).</emphasis>
<collectingRegion id="ED41F80FFFCFFFE60A1C0A8A0A2396B4" country="Colombia" name="Valle del Cauca" pageId="91" pageNumber="92">
<emphasis id="1DF1EAFFFFCFFFE60A1C0A8A0A2396B4" bold="true" pageId="91" pageNumber="92">
Valle del 
<taxonomicName id="E8854D6EFFCFFFE60FA60AEE0A2396B4" authority=": Estacion Agro-Forestal Bajo Calima, Dagua." authorityName="Estacion Agro-Forestal Bajo Calima, Dagua." box="[189,269,1743,1769]" class="Insecta" family="Cerambycidae" genus="Cauca" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">Cauca</taxonomicName>
</emphasis>
</collectingRegion>
: Estación Agro-Forestal Bajo Calima, Dagua. 
<emphasis id="1DF1EAFFFFCFFFE60C2B0AEE08C996B7" bold="true" box="[816,999,1743,1770]" italics="true" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60C2B0AEE08C996B7" box="[816,999,1743,1770]" country="Colombia" name="Cundinamarca" pageId="91" pageNumber="92">Cundinamarca</collectingRegion>
</emphasis>
. 
<emphasis id="1DF1EAFFFFCFFFE60CED0AEE0F6396B4" bold="true" box="[1014,1101,1743,1769]" pageId="91" pageNumber="92">
<collectingRegion id="ED41F80FFFCFFFE60CED0AEE0F6396B4" box="[1014,1101,1743,1769]" country="Colombia" name="Tolima" pageId="91" pageNumber="92">Tolima</collectingRegion>
</emphasis>
.
</paragraph>
<paragraph id="2F3A36EDFFCFFFE60FA60B3008379776" blockId="91.[189,793,1809,1835]" box="[189,793,1809,1835]" pageId="91" pageNumber="92">
<heading id="74728181FFCFFFE60FA60B3008379776" bold="true" box="[189,793,1809,1835]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="1DF1EAFFFFCFFFE60FA60B3008379776" bold="true" box="[189,793,1809,1835]" pageId="91" pageNumber="92">Intraspecific variation and taxonomic discussion</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFCFFFE10FA60B610F349131" blockId="91.[189,1399,1856,2027]" lastBlockId="92.[189,1398,267,364]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
Despite its extensive geographical distribution and the large number of specimens studied for this work, limited morphological variation was observed in 
<taxonomicName id="E8854D6EFFCFFFE60DEC0B4408949722" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[759,954,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCFFFE60DEC0B4408949722" box="[759,954,1893,1919]" italics="true" pageId="91" pageNumber="92">S. aequinoctialis</emphasis>
</taxonomicName>
apart from colouration and size 
<superScript id="D8F09BA5FFCFFFE60A300B420E12972C" attach="left" box="[1323,1340,1891,1905]" fontSize="6" pageId="91" pageNumber="92">19</superScript>
. The most noticeable variation is related to the sculpture at the centre of the pronotum, which can be almost completely smooth, without any trace of alveolar microsculpture or micropunctation, in some specimens, or have microsculpture and micropunctation present, but very fine and effaced, in others. It is interesting to note that in its sister species, 
<taxonomicName id="E8854D6EFFC8FFE10D2F0D2D09879178" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[564,681,268,294]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC8FFE10D2F0D2D09879178" box="[564,681,268,294]" italics="true" pageId="92" pageNumber="93">S. proseni</emphasis>
</taxonomicName>
, the degree to which the pronotal micropunctation is marked varies intrapopulationally at a much stronger intensity. A detailed comparison between 
<taxonomicName id="E8854D6EFFC8FFE10BA80D0E0E589114" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1203,1398,303,329]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10BA80D0E0E589114" box="[1203,1398,303,329]" italics="true" pageId="92" pageNumber="93">S. aequinoctialis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E8854D6EFFC8FFE10FF40D730A4D9131" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[239,355,338,364]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC8FFE10FF40D730A4D9131" box="[239,355,338,364]" italics="true" pageId="92" pageNumber="93">S. proseni</emphasis>
</taxonomicName>
is present in the taxonomic discussion of the latter species.
</paragraph>
</subSubSection>
<subSubSection id="679F6566FFC8FFE20FA60DB208969114" lastPageId="95" lastPageNumber="96" pageId="92" pageNumber="93" type="discussion">
<paragraph id="2F3A36EDFFC8FFE10FA60DB20A6A91F0" blockId="92.[189,324,403,429]" box="[189,324,403,429]" pageId="92" pageNumber="93">
<heading id="74728181FFC8FFE10FA60DB20A6A91F0" bold="true" box="[189,324,403,429]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="1DF1EAFFFFC8FFE10FA60DB20A6A91F0" bold="true" box="[189,324,403,429]" pageId="92" pageNumber="93">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFC8FFE10FA60DE3092092F2" blockId="92.[189,1399,450,687]" pageId="92" pageNumber="93">
The earliest references found in the literature on the species we now know as 
<taxonomicName id="E8854D6EFFC8FFE10B410DE30E309181" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1114,1310,450,476]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10B410DE30E309181" box="[1114,1310,450,476]" italics="true" pageId="92" pageNumber="93">S. aequinoctialis</emphasis>
</taxonomicName>
are the records by 
<bibRefCitation id="4B144B1CFFC8FFE10E260DC40AF291A2" author="Dejean P. F. M. A." box="[317,476,485,511]" pageId="92" pageNumber="93" pagination="1833 - 1836" refId="ref133982" refString="Dejean P. F. M. A. 1833. [Livraison 2.]. In: Dejean P. F. M. A. 1833 - 1836. Catalogue des Coleopteres de la collection de M. le Comte Dejean: 97 - 176. Mequignon-Marvis Pere et Fils, Paris. [Publication date according to Madge 1988.] Available from http: // biodiversitylibrary. org / page / 9322478 [accessed 30 May 2018]." type="journal article" year="1833">Dejean (1833</bibRefCitation>
, 1836) of the presence of “ 
<taxonomicName id="E8854D6EFFC8FFE10C090DC40F1F91A2" box="[786,1073,485,511]" class="Insecta" family="Scarabaeidae" genus="Coprobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10C090DC40F1F91A2" box="[786,1073,485,511]" italics="true" pageId="92" pageNumber="93">Coprobius aequinoctialis</emphasis>
</taxonomicName>
”, from “Carthagena”, in his beetle collection. As for other nominal species firstly established in the catalogues of Dejean, this name is unavailable as it was not published along with a description or indication and, therefore, it is out of the availability criteria adopted by the Code (Articles 11 and 12). Only thirty years later, 
<bibRefCitation id="4B144B1CFFC8FFE10B850E6F0E7A9234" author="Harold E." box="[1182,1364,590,617]" pageId="92" pageNumber="93" pagination="1 - 144" refId="ref136501" refString="Harold E. 1868 a. Monographie der Gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120104" type="journal article" year="1868">Harold (1868a)</bibRefCitation>
, in his revision of 
<taxonomicName id="E8854D6EFFC8FFE10E760E530AFC92D1" baseAuthorityName="Halffter &amp; Martinez" baseAuthorityYear="1977" box="[365,466,626,652]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFFC8FFE10E760E530AFC92D1" box="[365,466,626,652]" italics="true" pageId="92" pageNumber="93">Canthon</emphasis>
</taxonomicName>
, at last described the species, now with the name 
<taxonomicName id="E8854D6EFFC8FFE10B070E530E0092D1" authorityName="Harold" authorityYear="1868" box="[1052,1326,626,652]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10B070E530E0092D1" box="[1052,1326,626,652]" italics="true" pageId="92" pageNumber="93">Canthon aequinoctialis</emphasis>
</taxonomicName>
, from “Columbien, Neu-Granada”.
</paragraph>
<paragraph id="2F3A36EDFFC8FFE00FA60EF70E0591A5" blockId="92.[189,1399,726,999]" lastBlockId="93.[189,1399,267,505]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
There is no doubt that the two specimens here recognized as part of the type series of 
<taxonomicName id="E8854D6EFFC8FFE10BA90EF60E5892AD" authorityName="Harold" authorityYear="1868" box="[1202,1398,726,753]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10BA90EF60E5892AD" box="[1202,1398,726,753]" italics="true" pageId="92" pageNumber="93">S. aequinoctialis</emphasis>
</taxonomicName>
were indeed described by 
<bibRefCitation id="4B144B1CFFC8FFE10EEB0ED809899349" author="Harold E." box="[496,679,761,788]" pageId="92" pageNumber="93" pagination="1 - 144" refId="ref136501" refString="Harold E. 1868 a. Monographie der Gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120104" type="journal article" year="1868">Harold (1868a)</bibRefCitation>
. The now-lectotype, besides having originated from Harold’s personal collection (nowadays deposited at the MNHN), bears a label with the information “ 
<taxonomicName id="E8854D6EFFC8FFE10BC30F3C0E58936A" authorityName="Harold" authorityYear="1868" box="[1240,1398,797,823]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10BC30F3C0E58936A" box="[1240,1398,797,823]" italics="true" pageId="92" pageNumber="93">aequinoctialis</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFFC8FFE10FD60F600A3C9307" box="[205,274,833,858]" italics="true" pageId="92" pageNumber="93">t. Har</emphasis>
” in Harold’s own handwriting (
<figureCitation id="B7BE2A68FFC8FFE10D910F6109D69307" box="[650,760,832,858]" captionStart="Fig" captionStartId="88.[189,232,1929,1955]" captionTargetBox="[192,1399,370,1881]" captionTargetId="figure@88.[189,1399,364,1885]" captionTargetPageId="88" captionText="Fig. 29. Type material of the aequinoctialis subgroup. A–B. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov., type series. A. Lectotype and its labels. B. Paralectotype and its labels. C–E. S. proseni (Martínez, 1949) stat. et comb. nov., type series. C. Holotype and its labels. D. Paratype 1. E. Paratype 2." figureDoi="http://doi.org/10.5281/zenodo.3824837" httpUri="https://zenodo.org/record/3824837/files/figure.png" pageId="92" pageNumber="93">Fig. 29A</figureCitation>
). The 
<typeStatus id="F03E884FFFC8FFE10C5B0F6108CF9307" box="[832,993,832,858]" pageId="92" pageNumber="93" type="paralectotype">paralectotype</typeStatus>
, in turn, housed at the ZMHB, has an old drawer label written “ 
<taxonomicName id="E8854D6EFFC8FFE10D130F4209809320" authorityName="Harold" authorityYear="1868" box="[520,686,867,893]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10D130F4209809320" box="[520,686,867,893]" italics="true" pageId="92" pageNumber="93">aequinoctialis</emphasis>
</taxonomicName>
/ 
<emphasis id="1DF1EAFFFFC8FFE10DD90F42080A9320" box="[706,804,867,893]" italics="true" pageId="92" pageNumber="93">Harold*</emphasis>
/ 
<emphasis id="1DF1EAFFFFC8FFE10C220F4508F09320" box="[825,990,867,893]" italics="true" pageId="92" pageNumber="93">
Nov. 
<collectingCountry id="5792767DFFC8FFE10C6F0F4208F09320" box="[884,990,867,893]" name="Grenada" pageId="92" pageNumber="93">Granada</collectingCountry>
</emphasis>
” (
<figureCitation id="B7BE2A68FFC8FFE10CE20F420F4A9320" box="[1017,1124,867,893]" captionStart="Fig" captionStartId="88.[189,232,1929,1955]" captionTargetBox="[192,1399,370,1881]" captionTargetId="figure@88.[189,1399,364,1885]" captionTargetPageId="88" captionText="Fig. 29. Type material of the aequinoctialis subgroup. A–B. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov., type series. A. Lectotype and its labels. B. Paralectotype and its labels. C–E. S. proseni (Martínez, 1949) stat. et comb. nov., type series. C. Holotype and its labels. D. Paratype 1. E. Paratype 2." figureDoi="http://doi.org/10.5281/zenodo.3824837" httpUri="https://zenodo.org/record/3824837/files/figure.png" pageId="92" pageNumber="93">Fig. 29B</figureCitation>
). According to Joachim Willers (personal communication to MC, 2015), curator at ZMHB, the asterisk following the author’s name indicates that the collection has at least one type specimen of that nominal species which, in this case, should be the female here recognized as 
<typeStatus id="F03E884FFFC8FFE10DC60FEC085593BA" box="[733,891,973,999]" pageId="92" pageNumber="93" type="paralectotype">paralectotype</typeStatus>
(and it is worth noting that 
<bibRefCitation id="4B144B1CFFC8FFE00BDF0FEC0BCA9178" author="Harold E." lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" pagination="1 - 144" refId="ref136501" refString="Harold E. 1868 a. Monographie der Gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120104" type="journal article" year="1868">Harold (1868a: 10)</bibRefCitation>
indeed made it clear he had examined specimens from the ZMHB for his revision of 
<taxonomicName id="E8854D6EFFC9FFE00BD70D2A0E1F9178" baseAuthorityName="Halffter &amp; Martinez" baseAuthorityYear="1977" box="[1228,1329,267,293]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFFC9FFE00BD70D2A0E1F9178" box="[1228,1329,267,293]" italics="true" pageId="93" pageNumber="94">Canthon</emphasis>
</taxonomicName>
). The choice which of the two known 
<typeStatus id="F03E884FFFC9FFE00D2F0D0E09B59114" box="[564,667,303,329]" pageId="93" pageNumber="94" type="syntype">syntypes</typeStatus>
should be designated as 
<typeStatus id="F03E884FFFC9FFE00CA50D0F0F049115" box="[958,1066,302,328]" pageId="93" pageNumber="94" type="lectotype">lectotype</typeStatus>
was not a simple one: while the female is in perfect condition, the male is significantly damaged around the pin, although no known diagnostic characters of the species have been lost. On the other hand, the male belongs to the Edgar von Harold collection, has a label handwritten by the nominal species’ author and, the most important fact, it is of the same sex as all the other name-bearing types in 
<taxonomicName id="E8854D6EFFC9FFE00C720D9A0F2F9188" authorityName="Halffter &amp; Martinez" authorityYear="1977" box="[873,1025,443,469]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFFC9FFE00C720D9A0F2F9188" box="[873,1025,443,469]" italics="true" pageId="93" pageNumber="94">Sylvicanthon</emphasis>
</taxonomicName>
(i.e., 
<typeStatus id="F03E884FFFC9FFE00B5F0D9A0F989188" box="[1092,1206,443,469]" pageId="93" pageNumber="94" type="holotype">holotypes</typeStatus>
and 
<typeStatus id="F03E884FFFC9FFE00BF50D9A0E44918B" box="[1262,1386,443,470]" pageId="93" pageNumber="94" type="lectotype">lectotypes</typeStatus>
). Therefore, the male 
<typeStatus id="F03E884FFFC9FFE00EB00DFE092891A4" box="[427,518,479,505]" pageId="93" pageNumber="94" type="syntype">syntype</typeStatus>
was chosen instead of the female to be here designated as 
<typeStatus id="F03E884FFFC9FFE00BAC0DFF0E0891A5" box="[1207,1318,478,504]" pageId="93" pageNumber="94" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<caption id="7BFA6665FFC8FFE10FA60BF108D397B7" ID-DOI="http://doi.org/10.5281/zenodo.3824841" ID-Zenodo-Dep="3824841" box="[189,1021,2000,2027]" httpUri="https://zenodo.org/record/3824841/files/figure.png" pageId="92" pageNumber="93" startId="92.[189,232,2000,2026]" targetBox="[189,1399,1037,1965]" targetPageId="92">
<paragraph id="2F3A36EDFFC8FFE10FA60BF108D397B7" blockId="92.[189,1021,2000,2027]" box="[189,1021,2000,2027]" pageId="92" pageNumber="93">
<emphasis id="1DF1EAFFFFC8FFE10FA60BF10A3797B7" bold="true" box="[189,281,2000,2026]" pageId="92" pageNumber="93">Fig. 30.</emphasis>
Distribution of the two species of the 
<taxonomicName id="E8854D6EFFC8FFE10DC00BF108AF97B7" authorityName="Harold" authorityYear="1868" box="[731,897,2000,2026]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC8FFE10DC00BF108AF97B7" box="[731,897,2000,2026]" italics="true" pageId="92" pageNumber="93">aequinoctialis</emphasis>
</taxonomicName>
subgroup.
</paragraph>
</caption>
<paragraph id="2F3A36EDFFC9FFE30FA60E01099A918A" blockId="93.[189,1399,544,958]" lastBlockId="94.[189,1399,267,472]" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94">
As 
<taxonomicName id="E8854D6EFFC9FFE00FFE0E010A869267" authorityName="Harold" authorityYear="1868" box="[229,424,544,570]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC9FFE00FFE0E010A869267" box="[229,424,544,570]" italics="true" pageId="93" pageNumber="94">S. aequinoctialis</emphasis>
</taxonomicName>
was for a long time confused with 
<taxonomicName id="E8854D6EFFC9FFE00C530E0108929267" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[840,956,544,570]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC9FFE00C530E0108929267" box="[840,956,544,570]" italics="true" pageId="93" pageNumber="94">S. proseni</emphasis>
</taxonomicName>
, much of what is published under the former name actually refers to the latter species (see the discussion of 
<taxonomicName id="E8854D6EFFC9FFE00B130E650F509200" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[1032,1150,580,606]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC9FFE00B130E650F509200" box="[1032,1150,580,606]" italics="true" pageId="93" pageNumber="94">S. proseni</emphasis>
</taxonomicName>
for more details). In particular, the geographical distribution cited in the literature for 
<taxonomicName id="E8854D6EFFC9FFE00CD40E460FBB92DD" authorityName="Harold" authorityYear="1868" box="[975,1173,614,641]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC9FFE00CD40E460FBB92DD" box="[975,1173,614,641]" italics="true" pageId="93" pageNumber="94">S. aequinoctialis</emphasis>
</taxonomicName>
suffered from this problem. In general, the species is cited as occurring in large parts of Central 
<taxonomicName id="E8854D6EFFC9FFE00B4D0EA80F9592FE" box="[1110,1211,649,675]" class="Insecta" family="Cerambycidae" genus="America" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="189" phylum="Arthropoda" rank="genus">America</taxonomicName>
and throughout the Amazon region, including 
<collectingCountry id="5792767DFFC9FFE00D590E8C09A3929A" box="[578,653,685,711]" name="Brazil" pageId="93" pageNumber="94">Brazil</collectingCountry>
, 
<collectingCountry id="5792767DFFC9FFE00DBA0E8C09F9929A" box="[673,727,685,711]" name="Peru" pageId="93" pageNumber="94">Peru</collectingCountry>
and 
<collectingCountry id="5792767DFFC9FFE00C3A0E8C0855929A" box="[801,891,685,711]" name="Bolivia" pageId="93" pageNumber="94">Bolivia</collectingCountry>
. In reality, however, in South 
<taxonomicName id="E8854D6EFFC9FFE00A110E8C0E5C929A" box="[1290,1394,685,711]" class="Insecta" family="Cerambycidae" genus="America" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="189" phylum="Arthropoda" rank="genus">America</taxonomicName>
, 
<taxonomicName id="E8854D6EFFC9FFE00FA60EF10A5192B7" authorityName="Harold" authorityYear="1868" box="[189,383,720,746]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC9FFE00FA60EF10A5192B7" box="[189,383,720,746]" italics="true" pageId="93" pageNumber="94">S. aequinoctialis</emphasis>
</taxonomicName>
is present only in the portion west of the Cordillera Oriental 
<superScript id="D8F09BA5FFC9FFE00B230EEF0F679281" attach="left" box="[1080,1097,718,732]" fontSize="6" pageId="93" pageNumber="94">20</superScript>
, in the Colombian Andes, specifically on the Caribbean coast (department of Atlántico), in the Chocó biogeographical region (maybe also present in 
<collectingCountry id="5792767DFFC9FFE00EC80F370919936D" box="[467,567,790,816]" name="Ecuador" pageId="93" pageNumber="94">Ecuador</collectingCountry>
, although not recorded from there yet) and in the valleys between the Central and Oriental Cordilleras, in places such as Tolima and Caldas. In Central 
<taxonomicName id="E8854D6EFFC9FFE00B9A0F180FC7930E" box="[1153,1257,825,851]" class="Insecta" family="Cerambycidae" genus="America" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="189" phylum="Arthropoda" rank="genus">America</taxonomicName>
, the species occurs at least from 
<collectingCountry id="5792767DFFC9FFE00EB10F7C0929932A" box="[426,519,861,887]" name="Panama" pageId="93" pageNumber="94">Panama</collectingCountry>
northwards to 
<collectingCountry id="5792767DFFC9FFE00DAC0F7C0807932A" box="[695,809,861,887]" name="Honduras" pageId="93" pageNumber="94">Honduras</collectingCountry>
(
<figureCitation id="B7BE2A68FFC9FFE00C220F7C08BE932A" box="[825,912,861,887]" captionStart="Fig" captionStartId="92.[189,232,2000,2026]" captionTargetBox="[189,1399,1037,1965]" captionTargetId="figure@92.[189,1399,1037,1965]" captionTargetPageId="92" captionText="Fig. 30. Distribution of the two species of the aequinoctialis subgroup." figureDoi="http://doi.org/10.5281/zenodo.3824841" httpUri="https://zenodo.org/record/3824841/files/figure.png" pageId="93" pageNumber="94">Fig. 30</figureCitation>
). 
<bibRefCitation id="4B144B1CFFC9FFE00CBE0F7C0FFF932A" author="Howden H. F. &amp; Young O. P." box="[933,1233,861,887]" pageId="93" pageNumber="94" pagination="1 - 204" refId="ref137827" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young (1981)</bibRefCitation>
, Kohlmann &amp; 
<bibRefCitation id="4B144B1CFFC9FFE00FA60FA10A6193C7" author="Solis A. &amp; Kohlmann B." box="[189,335,896,922]" pageId="93" pageNumber="94" pagination="1 - 68" refId="ref146463" refString="Solis A. &amp; Kohlmann B. 2002. El genero Canthon (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia 10: 1 - 68." type="journal article" year="2002">Solís (2002)</bibRefCitation>
and 
<bibRefCitation id="4B144B1CFFC9FFE00E970FA1096D93C7" author="Halffter G." box="[396,579,896,922]" pageId="93" pageNumber="94" pagination="21 - 43" refId="ref136039" refString="Halffter G. 2003. Tribu Scarabaeini, subtribu Canthonina. In: Moron M. A. (ed.) Atlas de los escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 21 - 43. Argania Editio, Barcelona." type="book chapter" year="2003">Halffter (2003)</bibRefCitation>
, without citing their primary sources, said that 
<taxonomicName id="E8854D6EFFC9FFE00B610FA10E1193C7" authorityName="Harold" authorityYear="1868" box="[1146,1343,896,922]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC9FFE00B610FA10E1193C7" box="[1146,1343,896,922]" italics="true" pageId="93" pageNumber="94">S. aequinoctialis</emphasis>
</taxonomicName>
was also present in 
<collectingCountry id="5792767DFFC9FFE00E760F820A9593E0" box="[365,443,931,957]" name="Belize" pageId="93" pageNumber="94">Belize</collectingCountry>
, but this information is probably incorrect, since we could not find any specimens with this provenance in none of the studied collections (including the former private collection of Henry Howden, today housed at the CMNC; François Génier, personal communication to MC, 2015), nor researchers having recently collected in that country found the species living there (Latha Thomas, University of 
<collectingCountry id="5792767DFFCAFFE30E7D0D570A9A91CD" box="[358,436,374,400]" name="Belize" pageId="94" pageNumber="95">Belize</collectingCountry>
, personal communication to MC, 2015). 
<bibRefCitation id="4B144B1CFFCAFFE30CBC0D570F7291CD" author="Halffter G." box="[935,1116,374,400]" pageId="94" pageNumber="95" pagination="21 - 43" refId="ref136039" refString="Halffter G. 2003. Tribu Scarabaeini, subtribu Canthonina. In: Moron M. A. (ed.) Atlas de los escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 21 - 43. Argania Editio, Barcelona." type="book chapter" year="2003">Halffter (2003)</bibRefCitation>
added that this species occurred in 
<collectingCountry id="5792767DFFCAFFE30E4A0DBB0AFE91E9" box="[337,464,410,436]" name="Guatemala" pageId="94" pageNumber="95">Guatemala</collectingCountry>
and most probably also in southern 
<collectingCountry id="5792767DFFCAFFE30C8D0DBB08DD91E9" box="[918,1011,410,436]" name="Mexico" pageId="94" pageNumber="95">Mexico</collectingCountry>
, but these records were equally unsupported by more detailed information.
</paragraph>
<caption id="7BFA6665FFC9FFE00FA60B440E6697B7" ID-DOI="http://doi.org/10.5281/zenodo.3824843" ID-Zenodo-Dep="3824843" httpUri="https://zenodo.org/record/3824843/files/figure.png" pageId="93" pageNumber="94" startId="93.[189,232,1893,1919]" targetBox="[189,1399,1019,1850]" targetPageId="93">
<paragraph id="2F3A36EDFFC9FFE00FA60B440E6697B7" blockId="93.[189,1398,1893,2026]" pageId="93" pageNumber="94">
<emphasis id="1DF1EAFFFFC9FFE00FA60B440A349722" bold="true" box="[189,282,1893,1919]" pageId="93" pageNumber="94">Fig. 31.</emphasis>
Metafemora of the two species of the 
<taxonomicName id="E8854D6EFFC9FFE00DF80B4408A79722" authorityName="Harold" authorityYear="1868" box="[739,905,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFC9FFE00DF80B4408A79722" box="[739,905,1893,1919]" italics="true" pageId="93" pageNumber="94">aequinoctialis</emphasis>
</taxonomicName>
subgroup. 
<emphasis id="1DF1EAFFFFC9FFE00B170B440F0F9722" bold="true" box="[1036,1057,1893,1919]" pageId="93" pageNumber="94">A</emphasis>
. 
<taxonomicName id="E8854D6EFFC9FFE00B2A0B440A4297FE" authority="(Harold, 1868)" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis" status="comb. nov.">
<emphasis id="1DF1EAFFFFC9FFE00B2A0B440E589722" box="[1073,1398,1893,1919]" italics="true" pageId="93" pageNumber="94">Sylvicanthon aequinoctialis</emphasis>
(Harold, 1868)
</taxonomicName>
<taxonomicNameLabel id="06C25784FFC9FFE00E6F0BA90AD897FE" box="[372,502,1928,1955]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="1DF1EAFFFFC9FFE00EE50BA8093C97FE" bold="true" box="[510,530,1929,1955]" pageId="93" pageNumber="94">B</emphasis>
. 
<taxonomicName id="E8854D6EFFC9FFE00D3A0BA8084A97FF" authority="(Martinez, 1949)" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[545,868,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC9FFE00D3A0BA809B897FF" box="[545,662,1929,1955]" italics="true" pageId="93" pageNumber="94">S. proseni</emphasis>
(Martínez, 1949)
</taxonomicName>
stat. et comb. nov. Arrow indicates the basal end of the posterior margin of the ventral face of metafemora (note that it is longer in 
<taxonomicName id="E8854D6EFFC9FFE00BA80B8D0E58979B" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1203,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="aequinoctialis" status="comb. nov.">
<emphasis id="1DF1EAFFFFC9FFE00BA80B8D0E58979B" box="[1203,1398,1964,1990]" italics="true" pageId="93" pageNumber="94">S. aequinoctialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06C25784FFC9FFE00FA60BEE0A1197B4" box="[189,319,1999,2025]" pageId="93" pageNumber="94" rank="species">comb. nov.</taxonomicNameLabel>
than in 
<taxonomicName id="E8854D6EFFC9FFE00E850BEE093C97B4" authorityName="Martinez" authorityYear="1949" baseAuthorityName="Martinez" baseAuthorityYear="1949" box="[414,530,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="proseni">
<emphasis id="1DF1EAFFFFC9FFE00E850BEE093C97B4" box="[414,530,1999,2025]" italics="true" pageId="93" pageNumber="94">S. proseni</emphasis>
</taxonomicName>
stat. et comb. nov. and may exceed the trochanter in the first species).
</paragraph>
</caption>
<paragraph id="2F3A36EDFFCAFFE30FA60DDE0A509244" blockId="94.[189,382,511,537]" box="[189,382,511,537]" pageId="94" pageNumber="95">
<heading id="74728181FFCAFFE30FA60DDE0A509244" bold="true" box="[189,382,511,537]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="1DF1EAFFFFCAFFE30FA60DDE0A509244" bold="true" box="[189,382,511,537]" pageId="94" pageNumber="95">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="2F3A36EDFFCAFFE30FA60E0F0F5894DC" blockId="94.[189,1399,558,1154]" pageId="94" pageNumber="95">
In contrast to the other species of 
<taxonomicName id="E8854D6EFFCAFFE30D490E0F09C49215" authorityName="Halffter &amp; Martinez" authorityYear="1977" box="[594,746,558,584]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="1DF1EAFFFFCAFFE30D490E0F09C49215" box="[594,746,558,584]" italics="true" pageId="94" pageNumber="95">Sylvicanthon</emphasis>
</taxonomicName>
, of which only fragmentary biological information has been published, it is possible to find far more detailed data about 
<taxonomicName id="E8854D6EFFCAFFE30CF80E730F849231" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[995,1194,594,620]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30CF80E730F849231" box="[995,1194,594,620]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
in the literature. 
<bibRefCitation id="4B144B1CFFCAFFE30FA60E540AC492D2" author="Howden H. F. &amp; Young O. P." box="[189,490,629,656]" pageId="94" pageNumber="95" pagination="1 - 204" refId="ref137827" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young (1981)</bibRefCitation>
, for example, in their important monograph on the Panamanian dung beetles, described several aspects of the life of this species based on populations from the island of Barro Colorado, in the Gatun Lake of the 
<collectingCountry id="5792767DFFCAFFE30EF30E9C096B928A" box="[488,581,701,727]" name="Panama" pageId="94" pageNumber="95">Panama</collectingCountry>
Canal. According to the authors, 
<taxonomicName id="E8854D6EFFCAFFE30CCF0E9C0FB6928B" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[980,1176,700,727]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30CCF0E9C0FB6928B" box="[980,1176,700,727]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
breeds throughout the year and is one of the most abundant dung beetle species on the island, not showing any seasonal variation in its population density. During the rainy season (May to December), they feed on both dung and carrion, with the consumption of carcasses of animals such as the agouti [
<taxonomicName id="E8854D6EFFCAFFE30B660F090A769338" authority="(Gray, 1842)" baseAuthorityName="Gray" baseAuthorityYear="1842" class="Mammalia" family="Dasyproctidae" genus="Dasyprocta" kingdom="Animalia" order="Rodentia" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="punctata">
<emphasis id="1DF1EAFFFFCAFFE30B660F090E59931C" box="[1149,1399,808,833]" italics="true" pageId="94" pageNumber="95">Dasyprocta punctata</emphasis>
(Gray, 1842)
</taxonomicName>
], collared peccary [
<taxonomicName id="E8854D6EFFCAFFE30D240F6A09F09339" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[575,734,843,868]" class="Mammalia" family="Tayassuidae" genus="Pecari" kingdom="Animalia" order="Artiodactyla" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="tajacu">
<emphasis id="1DF1EAFFFFCAFFE30D240F6A09F09339" box="[575,734,843,868]" italics="true" pageId="94" pageNumber="95">Pecari tajacu</emphasis>
</taxonomicName>
(Linnaeus, 1758]), and rat, as well as human faeces and coati [
<taxonomicName id="E8854D6EFFCAFFE30E1D0F4E095F93D4" authority="nasua (Linnaeus, 1758)" authorityName="nasua (Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[262,625,878,905]" class="Mammalia" family="Procyonidae" genus="Nasua" kingdom="Animalia" order="Carnivora" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="nasua">
<emphasis id="1DF1EAFFFFCAFFE30E1D0F4E0AB093D5" box="[262,414,879,904]" italics="true" pageId="94" pageNumber="95">Nasua nasua</emphasis>
(Linnaeus, 1758)
</taxonomicName>
], jaguarundi [
<taxonomicName id="E8854D6EFFCAFFE30C090F4E0E4193D4" authority="(Geoffroy Saint-Hilaire, 1803)" baseAuthorityName="Geoffroy Saint-Hilaire" baseAuthorityYear="1803" box="[786,1391,878,905]" class="Mammalia" family="Felidae" genus="Puma" kingdom="Animalia" order="Carnivora" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="yagouaroundi">
<emphasis id="1DF1EAFFFFCAFFE30C090F4E0F2F93D5" box="[786,1025,878,904]" italics="true" pageId="94" pageNumber="95">Puma yagouaroundi</emphasis>
(Geoffroy Saint-Hilaire, 1803)
</taxonomicName>
] and tapir [
<taxonomicName id="E8854D6EFFCAFFE30E2F0FB2095B93F1" authority="(Gill, 1865)" baseAuthorityName="Gill" baseAuthorityYear="1865" box="[308,629,913,940]" class="Mammalia" family="Tapiridae" genus="Tapirus" kingdom="Animalia" order="Perissodactyla" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="bairdii">
<emphasis id="1DF1EAFFFFCAFFE30E2F0FB20ACC93F1" box="[308,482,914,940]" italics="true" pageId="94" pageNumber="95">Tapirus bairdii</emphasis>
(Gill, 1865)
</taxonomicName>
] dung, being reported. During the dry season (December to May), however, when carcasses are scarcer and consumed mainly by vertebrates or specialist necrophagous dung beetles (
<bibRefCitation id="4B144B1CFFCAFFE30E7F0FF80AD793AE" author="Young O. P." box="[356,505,985,1011]" pageId="94" pageNumber="95" refId="ref148329" refString="Young O. P. 1978. Resource Partitioning in a Neotropical Necrophagous Scarab Guild. PhD thesis, University of Maryland, College Park, MD." type="book" year="1978">Young 1978</bibRefCitation>
, 
<bibRefCitation id="4B144B1CFFCAFFE30D130FF8096993AE" author="Young O. P." box="[520,583,985,1011]" pageId="94" pageNumber="95" pagination="253 - 255" refId="ref148357" refString="Young O. P. 1980. Bone burial by a Neotropical dung beetle (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 34 (2): 253 - 255. Available from http: // www. jstor. org / stable / 4000139 [accessed 30 May 2018]." type="journal article" year="1980">1980</bibRefCitation>
), 
<taxonomicName id="E8854D6EFFCAFFE30D460FF8080D93AE" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[605,803,985,1011]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30D460FF8080D93AE" box="[605,803,985,1011]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
is found only on dung. Regarding their circadian cycle, 
<taxonomicName id="E8854D6EFFCAFFE30E110FDC0AE3944B" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[266,461,1020,1047]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30E110FDC0AE3944B" box="[266,461,1020,1047]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
, as the other species in the genus, is crepuscular and nocturnal, with an activity period between 5 and 9 pm (Howden &amp; 
<bibRefCitation id="4B144B1CFFCAFFE30D81080008039467" author="Young O. P." box="[666,813,1056,1083]" pageId="94" pageNumber="95" pagination="253 - 255" refId="ref148357" refString="Young O. P. 1980. Bone burial by a Neotropical dung beetle (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 34 (2): 253 - 255. Available from http: // www. jstor. org / stable / 4000139 [accessed 30 May 2018]." type="journal article" year="1980">Young 1980</bibRefCitation>
). In Barro Colorado, 
<taxonomicName id="E8854D6EFFCAFFE30B3108000FC09467" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1066,1262,1056,1083]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30B3108000FC09467" box="[1066,1262,1056,1083]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
is the most abundant roller dung beetle, having been responsible for the consumption of at least one-fourth of the total volume of tapir dung utilized in experiments by 
<bibRefCitation id="4B144B1CFFCAFFE30C35084908FF94DC" author="Young O. P." box="[814,977,1127,1154]" pageId="94" pageNumber="95" pagination="319 - 324" refId="ref148410" refString="Young O. P. 2009. Observations on burrowing behavior by Panamanian Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 63 (3): 319 - 324. https: // doi. org / 10.1649 / 1164.1" type="journal article" year="2009">Young (2009)</bibRefCitation>
on the island.
</paragraph>
<paragraph id="2F3A36EDFFCAFFE30FA608880F4F95C5" blockId="94.[189,1399,1193,1433]" pageId="94" pageNumber="95">
Farther north, we have information about some populations of 
<taxonomicName id="E8854D6EFFCAFFE30CBA08880F4B949E" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[929,1125,1193,1219]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30CBA08880F4B949E" box="[929,1125,1193,1219]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
from 
<collectingCountry id="5792767DFFCAFFE30BB608880E00949E" box="[1197,1326,1193,1219]" name="Costa Rica" pageId="94" pageNumber="95">Costa Rica</collectingCountry>
given by Kohlmann &amp; 
<bibRefCitation id="4B144B1CFFCAFFE30E9E08EC093494BA" author="Solis A. &amp; Kohlmann B." box="[389,538,1229,1255]" pageId="94" pageNumber="95" pagination="1 - 68" refId="ref146463" refString="Solis A. &amp; Kohlmann B. 2002. El genero Canthon (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia 10: 1 - 68." type="journal article" year="2002">Solís (2002)</bibRefCitation>
. Like in 
<collectingCountry id="5792767DFFCAFFE30D9A08EC09CF94BA" box="[641,737,1229,1255]" name="Panama" pageId="94" pageNumber="95">Panama</collectingCountry>
, Costa Rican adults of 
<taxonomicName id="E8854D6EFFCAFFE30CE808EC0F9994BA" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1011,1207,1229,1255]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30CE808EC0F9994BA" box="[1011,1207,1229,1255]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
from both sides of the Continental Divide (i.e., both on the Caribbean and the Pacific coasts of 
<collectingCountry id="5792767DFFCAFFE30B7408D10FDD9556" box="[1135,1267,1264,1291]" name="Costa Rica" pageId="94" pageNumber="95">Costa Rica</collectingCountry>
) are active throughout the year and are among the most abundant nocturnal dung beetles of humid lowland forests. There, they live up to 
<quantity id="E87D9B08FFCAFFE30EDB09160924950F" box="[448,522,1335,1362]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="94" pageNumber="95" unit="m" value="800.0">800 m</quantity>
altitude on the Atlantic side of the Continental Divide, and up to 
<quantity id="E87D9B08FFCAFFE30A1309160E7C950F" box="[1288,1362,1335,1362]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="94" pageNumber="95" unit="m" value="900.0">900 m</quantity>
on the Pacific side. Specimens were collected on human and equine dung, as well as using flight interception traps and through active collections on the ground or beneath fallen tree trunks.
</paragraph>
<paragraph id="2F3A36EDFFCAFFE30FA609E10F109634" blockId="94.[189,1399,1472,1641]" pageId="94" pageNumber="95">
Lastly, we have data for populations in 
<collectingCountry id="5792767DFFCAFFE30D9A09E109D79587" box="[641,761,1472,1498]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
. 
<bibRefCitation id="4B144B1CFFCAFFE30C1909E108C09587" author="Medina C. A. C. &amp; Medina-Uribe C. A. &amp; Quintero B. G. M. &amp; Escobar-Villa A. F. &amp; Chuaire L. M. C. &amp; Posada N. J. B." box="[770,1006,1472,1498]" pageId="94" pageNumber="95" refId="ref141571" refString="Medina C. A. C., Medina-Uribe C. A., Quintero B. G. M., Escobar-Villa A. F., Chuaire L. M. C. &amp; Posada N. J. B. 2012. Escarabajos coprofagos (Scarabaeinae) del eje cafetero: Guia para el estudio ecologico. No editor, Villa Maria." type="book" year="2012">
Medina 
<emphasis id="1DF1EAFFFFCAFFE30C7A09E008B49587" box="[865,922,1472,1498]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
(2012)
</bibRefCitation>
also highlighted 
<taxonomicName id="E8854D6EFFCAFFE30BAF09E00E589587" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1204,1398,1472,1499]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30BAF09E00E589587" box="[1204,1398,1472,1499]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
as one of the most common species in the 
<collectingRegion id="ED41F80FFFCAFFE30DA509C5086E95A3" box="[702,832,1508,1534]" country="Colombia" name="Magdalena" pageId="94" pageNumber="95">Magdalena</collectingRegion>
River Valley region at an altitude between 180 and 
<quantity id="E87D9B08FFCAFFE30FF50A290A14967F" box="[238,314,1544,1570]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="94" pageNumber="95" unit="m" value="300.0">300 m</quantity>
, and stated that the species was collected with traps baited with human faeces and cow dung, mushrooms and, occasionally, with rotten fish. Additionally, 
<bibRefCitation id="4B144B1CFFCAFFE30C940A0A0F4A9618" author="Solis C. &amp; Noriega J. A. &amp; Herrera G." box="[911,1124,1579,1605]" pageId="94" pageNumber="95" pagination="33 - 41" refId="ref146525" refString="Solis C., Noriega J. A. &amp; Herrera G. 2011. Escarabajos coprofagos (Coleoptera: Scarabaeinae) en tres bosques secos del departamento del Atlatico-Colombia. Boletin del Museo de Entomologia de la Universidad del Valle 12 (1): 33 - 41." type="journal article" year="2011">
Solís 
<emphasis id="1DF1EAFFFFCAFFE30CC90A0D0F209618" box="[978,1038,1579,1605]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
(2011)
</bibRefCitation>
stated that this species was present in the 
<collectingRegion id="ED41F80FFFCAFFE30E8D0A6E09A09634" box="[406,654,1615,1641]" country="Colombia" name="Atlantico" pageId="94" pageNumber="95">Atlántico department</collectingRegion>
in altitudes between 160 and 
<specimenCount id="3983FD64FFCAFFE30CF70A6F0F109634" box="[1004,1086,1614,1641]" pageId="94" pageNumber="95" type="generic">
<quantity id="E87D9B08FFCAFFE30CF70A6F0F149634" box="[1004,1082,1614,1641]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="94" pageNumber="95" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="2F3A36EDFFCAFFE20FA60AB108969114" blockId="94.[189,1399,1680,2027]" lastBlockId="95.[189,1399,267,330]" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95">
Several of the specimens examined also have bionomic information on their labels. There are records of specimens collected using traps baited with swine, human and howler monkey [
<taxonomicName id="E8854D6EFFCAFFE30B800A950AA796AF" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Atelidae" genus="Alouatta" kingdom="Animalia" order="Primates" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="seniculus">
<emphasis id="1DF1EAFFFFCAFFE30B800A950E589693" box="[1179,1398,1716,1742]" italics="true" pageId="94" pageNumber="95">Alouatta seniculus</emphasis>
(Linnaeus, 1758)
</taxonomicName>
] dung, as well as rotten flesh, and also in flight interception traps and with ultraviolet light. Specimens were collected in all the months of the year, but there is an apparent higher concentration during the rainiest months (March: 49; April: 87; May: 52; June: 76; July: 63), when 327 out of the 573 studied specimens were caught. In relation to the altitudinal amplitude, there are representatives of 
<taxonomicName id="E8854D6EFFCAFFE30FA60B470AAF97DD" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[189,385,1894,1920]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30FA60B470AAF97DD" box="[189,385,1894,1920]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
collected from at approximately the sea level up to areas as elevated as 
<quantity id="E87D9B08FFCAFFE30BC80B470E0397DD" box="[1235,1325,1894,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="94" pageNumber="95" unit="m" value="1000.0">1000 m</quantity>
in the Reserva Natural Cerro Kilambé (
<collectingRegion id="ED41F80FFFCAFFE30D580BA8098797FE" box="[579,681,1929,1955]" country="Nicaragua" name="Jinotega" pageId="94" pageNumber="95">Jinotega</collectingRegion>
, 
<collectingCountry id="5792767DFFCAFFE30DAD0BA8081F97F9" box="[694,817,1929,1956]" name="Nicaragua" pageId="94" pageNumber="95">Nicaragua</collectingCountry>
) and 
<quantity id="E87D9B08FFCAFFE30C680BAB08E297F9" box="[883,972,1930,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="94" pageNumber="95" unit="m" value="1440.0">1440 m</quantity>
in San Luis (
<collectingRegion id="ED41F80FFFCAFFE30B700BA80FCB97FE" box="[1131,1253,1929,1955]" country="Colombia" name="Antioquia" pageId="94" pageNumber="95">Antioquia</collectingRegion>
, 
<collectingCountry id="5792767DFFCAFFE30BE90BA80E4597F9" box="[1266,1387,1929,1956]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
). Therefore, after combining all the available data, what we see is that 
<taxonomicName id="E8854D6EFFCAFFE30CFD0B8F0F87979A" authorityName="Cupello &amp; Vaz-De" authorityYear="2018" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[998,1193,1965,1992]" class="Insecta" family="Scarabaeidae" genus="Sylvicanthon" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aequinoctialis">
<emphasis id="1DF1EAFFFFCAFFE30CFD0B8F0F87979A" box="[998,1193,1965,1992]" italics="true" pageId="94" pageNumber="95">S. aequinoctialis</emphasis>
</taxonomicName>
is, throughout its distribution, an opportunistic species that explores several food sources and habitats and that is resistant to the seasonal changes, characteristics that make it one of the most abundant and dominant species among the roller 
<taxonomicName id="E8854D6EFFCBFFE20E9D0D0E090F9114" baseAuthorityName="Tarasov &amp; Genier" baseAuthorityYear="2015" box="[390,545,303,329]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
of the New World tropical forests.
</paragraph>
</subSubSection>
</treatment>
</document>