185 lines
22 KiB
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185 lines
22 KiB
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<document id="D126CA78F6100C0340B97C1F0925E882" ID-DOI="http://dx.doi.org/10.3897/BDJ.2.e4117" ID-PMC="PMC4238084" ID-Pensoft-Pub="1314-2828--4117" ID-PubMed="25425938" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1314-2828--e4117" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="The Afrotropical Miomantiscaffra Saussure 1871 and M.paykullii Stal 1871: first records of alien mantid species in Portugal and Europe, with an updated checklist of Mantodea in Portugal (Insecta: Mantodea)" checkinTime="1553126635119" checkinUser="pensoft" docAuthor="Marabuto, Eduardo" docDate="2014" docId="E2C500871464B4CB012F2E29891F4261" docLanguage="en" docName="BiodivDatJour 2: e4117" docOrigin="Biodiversity Data Journal 2" docSource="http://dx.doi.org/10.3897/BDJ.2.e4117" docTitle="Miomantis paykullii Stal 1871" docType="treatment" docVersion="5" lastPageNumber="4117" masterDocId="E354FFB1DF588A09D215B52A0513FFB7" masterDocTitle="The Afrotropical Miomantiscaffra Saussure 1871 and M. paykullii Stal 1871: first records of alien mantid species in Portugal and Europe, with an updated checklist of Mantodea in Portugal (Insecta: Mantodea)" masterLastPageNumber="4117" masterPageNumber="4117" pageNumber="4117" updateTime="1668122760864" updateUser="ExternalLinkService">
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<mods:title id="5881A8D912B8A1B4D05136CC61367703">The Afrotropical Miomantiscaffra Saussure 1871 and M. paykullii Stal 1871: first records of alien mantid species in Portugal and Europe, with an updated checklist of Mantodea in Portugal (Insecta: Mantodea)</mods:title>
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<mods:name id="0BE272583EC3A17820790D7108039DAE" type="personal">
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<mods:namePart id="C710F606CE94BC1E3C919EB2B7306DA6">Marabuto, Eduardo</mods:namePart>
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<mods:typeOfResource id="B252EA19F6E54A0A6EE1ADEE7968EB2D">text</mods:typeOfResource>
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<mods:title id="8A7AE0D7F91C8F9E7ACEE7A8514FDC3D">Biodiversity Data Journal</mods:title>
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<mods:date id="4B355AB1936869BD0F53C2A62BCF45C7">2014</mods:date>
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<mods:detail id="5E62B720609E7FCB43C0B49EC6467160" type="volume">
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<mods:number id="A5AFBC2BB4C2DDB79E9F8D801E669B19">2</mods:number>
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<mods:start id="4BF9E426498CF46E3ED90B37230A4C58">4117</mods:start>
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<mods:location id="37AB5C82CF276A1F47628F12C7F5D7B8">
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<mods:url id="F368126CC86889954E733B2728701C3A">http://dx.doi.org/10.3897/BDJ.2.e4117</mods:url>
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<mods:classification id="19E80575807A75C1A96B5A013FF9F08A">journal article</mods:classification>
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<mods:identifier id="E098ABEF928C9032BC49B7E93AFED540" type="DOI">http://dx.doi.org/10.3897/BDJ.2.e4117</mods:identifier>
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<mods:identifier id="18798A70E7A55F85D2D955C7774FC3AC" type="Pensoft-Pub">1314-2828--4117</mods:identifier>
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</mods:mods>
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<treatment id="E2C500871464B4CB012F2E29891F4261" LSID="urn:lsid:plazi:treatment:E2C500871464B4CB012F2E29891F4261" httpUri="http://treatment.plazi.org/id/E2C500871464B4CB012F2E29891F4261" lastPageNumber="4117" pageId="0" pageNumber="4117">
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<subSubSection id="C63F371F6F5FB36D4388F230DB9BB4E8" pageId="0" pageNumber="4117" type="nomenclature">
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<paragraph id="CF3DAB967A952B3A9CF69B258A74E99C" pageId="0" pageNumber="4117">
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<taxonomicName id="5BE89A3284DFA92D271FB6FB7B6A224A" ID-CoL="43L4F" ID-ENA="627754" authority="Stal, 1871" authorityName="Stal" authorityYear="1871" class="Insecta" family="Miomantidae" genus="Miomantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miomantis paykullii" order="Mantodea" pageId="0" pageNumber="4117" phylum="Arthropoda" rank="species" species="paykullii">Miomantis paykullii Stal, 1871</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3882FA69C0215AC7D9B17D309593F29" pageId="0" pageNumber="4117" type="materials_examined">
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<paragraph id="98F732E4495DDE61259F07B1E9B1FB1A" pageId="0" pageNumber="4117">Materials</paragraph>
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<paragraph id="633FF4B03244280BEB28D9FD1ED510B2" pageId="0" pageNumber="4117">
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<materialsCitation id="E337BEC545A2C4D0D19BAE7B96E01A9B" collectingDate="2014-05-08" collectingMethod="ad hoc observation" collectorName="Eduardo Marabuto" country="Portugal" latitude="37.09" location="Vila Sol" longitude="-8.093" pageId="0" pageNumber="4117" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
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Type status:
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<typeStatus id="FA836D82159DD8007A153038480D5263" pageId="0" pageNumber="4117">Other material</typeStatus>
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. Occurrence: recordedBy:
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<collectorName id="7E48328A2287DBC01D71E9819D93A326" pageId="0" pageNumber="4117">Eduardo Marabuto</collectorName>
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; individualCount:
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<specimenCount id="47B08B62FCE0A5E43A8AD65F0A0BEDE4" pageId="0" pageNumber="4117">1</specimenCount>
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; sex:
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<specimenType id="AF4350F82B6BA107B5E9BA754C75548D" pageId="0" pageNumber="4117">male</specimenType>
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; Taxon: genus: Miomantis; specificEpithet: paykullii; taxonRank: species; scientificNameAuthorship: Stal, 1871; vernacularName: Egyptian pygmy mantis; Location: continent: Europe; country:
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<collectingCountry id="D011052A0255AECB67B4FF6534CAA0C3" pageId="0" pageNumber="4117">Portugal</collectingCountry>
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; countryCode: PT; stateProvince: Faro; county:
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<normalizedToken id="246DCE8E492FF5FAF2EA4578713E3508" originalValue="Loulé">Loule</normalizedToken>
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; municipality: Quarteira; locality:
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<location id="2FB7C33AD877685B54FF2096F5241B4E" LSID="urn:lsid:plazi:treatment:E2C500871464B4CB012F2E29891F4261:2FB7C33AD877685B54FF2096F5241B4E" country="Portugal" latitude="37.09" longitude="-8.093" name="Vila Sol" pageId="0" pageNumber="4117">Vila Sol</location>
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; verbatimElevation:
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<elevation id="A831783647A7B2111166E97F4C796942" pageId="0" pageNumber="4117">40</elevation>
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; decimalLatitude:
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<geoCoordinate id="B0834854651CD76006000B5917EB43DF" orientation="latitude" pageId="0" pageNumber="4117" value="37.090">37.090</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate id="613946481EE9A3E39AACCF4D17AC201E" orientation="longitude" pageId="0" pageNumber="4117" value="-08.093">-08.093</geoCoordinate>
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; Event: samplingProtocol:
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<collectingMethod id="D1B3E1852B892EAE7EA1EA15F4791AF6" pageId="0" pageNumber="4117">ad hoc observation</collectingMethod>
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; eventDate:
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<collectingDate id="AB4CB07938864CC88D69B608D29FBF58" pageId="0" pageNumber="4117" value="2014-05-08">2014-08-05</collectingDate>
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; habitat: garden
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</materialsCitation>
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<materialsCitation id="0F547BB84527DFDE0D1958ACFFA17AD4" collectingDate="2014-08-08" collectingMethod="vegetation beating" collectorName="Eduardo Marabuto" country="Portugal" latitude="37.09" location="Vila Sol" longitude="-8.093" pageId="0" pageNumber="4117" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
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Type status:
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<typeStatus id="2CDBAF1F5643762C42F198A78768D601" pageId="0" pageNumber="4117">Other material</typeStatus>
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. Occurrence: recordedBy:
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<collectorName id="235495E9F014F0939B3E6936BE0DC58C" pageId="0" pageNumber="4117">Eduardo Marabuto</collectorName>
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; individualCount:
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<specimenCount id="0F0606B7033523EAE1A1700C0B34A904" pageId="0" pageNumber="4117">1</specimenCount>
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; sex:
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<specimenType id="5C596490399BAA83469B0AAA83FDCC53" pageId="0" pageNumber="4117">male</specimenType>
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; Taxon: genus: Miomantis; specificEpithet: paykullii; taxonRank: species; scientificNameAuthorship: Stal, 1871; vernacularName: Egyptian pygmy mantis; Location: continent: Europe; country:
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<collectingCountry id="BEF019D0D0BB5AD92F7A982C208480F0" pageId="0" pageNumber="4117">Portugal</collectingCountry>
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; countryCode: PT; stateProvince: Faro; county:
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<normalizedToken id="1F842E812500770F7B84944DE0D61C1E" originalValue="Loulé">Loule</normalizedToken>
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; municipality: Quarteira; locality:
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<location id="699BF20D6DAF18830D8F0C8799FE23C3" LSID="urn:lsid:plazi:treatment:E2C500871464B4CB012F2E29891F4261:699BF20D6DAF18830D8F0C8799FE23C3" country="Portugal" latitude="37.09" longitude="-8.093" name="Vila Sol" pageId="0" pageNumber="4117">Vila Sol</location>
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; verbatimElevation:
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<elevation id="641DF7F4987BF44EB9BD123C3E62A4BF" pageId="0" pageNumber="4117">40</elevation>
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; decimalLatitude:
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<geoCoordinate id="1BC0BC11CB64D08F11603F8046A8652C" orientation="latitude" pageId="0" pageNumber="4117" value="37.090">37.090</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate id="8BC929944357FEADD18023D87C9F81A6" orientation="longitude" pageId="0" pageNumber="4117" value="-08.093">-08.093</geoCoordinate>
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; Event: samplingProtocol:
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<collectingMethod id="9AB34E05FE6B8AD7ED5D33263D53DCD7" pageId="0" pageNumber="4117">vegetation beating</collectingMethod>
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; eventDate:
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<collectingDate id="8947B8F0956D2AF3DD4ED32F250A7FD8" pageId="0" pageNumber="4117" value="2014-08-08">2014-08-08</collectingDate>
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; habitat: garden
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9081587BDD8A05C4A2F21090F1D32555" pageId="0" pageNumber="4117" type="description">
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<paragraph id="0B53F8017F0AC73C7FBBC02B69AB0456" pageId="0" pageNumber="4117">Description</paragraph>
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<paragraph id="C930C590EE19B7079AC5120651342FC9" pageId="0" pageNumber="4117">
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Adapted from
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<bibRefCitation id="6674836672ABC64E1305DA9EC0001FB5" author="Giglio-Tos, E." journalOrPublisher="Das Tierreich" pageId="0" pageNumber="4117" pagination="1 - 707" title="Orthoptera. Mantidae" volume="50" year="1927">Giglio-Tos 1927</bibRefCitation>
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,
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<bibRefCitation id="C79B4A5C2C76C9A13ACD17AD5F71392D" author="Ehrmann, R" journalOrPublisher="Natur und Tier - Verlag, Muenster" pageId="0" pageNumber="4117" title="Mantodea: Gottesanbeterinnen der Welt." year="2002">Ehrmann 2002</bibRefCitation>
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and
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<bibRefCitation id="145102493FF35425F9A67568613169EA" author="Agabiti, Barbara" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa (S. E. A)" pageId="0" pageNumber="4117" pagination="1 - 20" title="The Mediterranean species of the genus Ameles Burmeister, 1838 (Insecta, Mantodea: Amelinae) with a biogeographic and phylogenetic evaluation" volume="47" year="2010">Agabiti et al. 2010</bibRefCitation>
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: Small or medium sized species. Green or light-brown coloured, patternless except occasional obscured humeral vein. Head broader than the pronotum, especially in the male. Eye prominent, slightly conical, more apparent in male but not ending in a spine. Pronotum slender with weakly developed supracoxal dilation, as long as or longer than fore coxa and smooth in male, finely toothed in female. Forewing hyaline in male, more opaque in female, distally dilated and slightly exceeding the tip of the abdomen in the former, only reaching the base of the cerci in the latter. Male hindwing hyaline, female yellowish, crossed by yellow veins. Supra-anal plate longer than broad, triangular. Foreleg unpatterned, coxa finely toothed with 5-6 spaced small spines stronger in female. Femur with 4 discoidal spines and 4 external. Fore tibia with 7 external spines. Body length: 36-39mm; pronotum length: 11-12mm in male, to 14mm in female; forewing length 23mm in male, 19-21mm in female.
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</paragraph>
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<subSection id="CF4EF5579985E8EB9172D92D3A312CA3" pageId="0" pageNumber="4117" type="portuguese specimens">
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<paragraph id="E80E234DF3A5777D43F536A56F4F8B14" pageId="0" pageNumber="4117">Portuguese specimens</paragraph>
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<paragraph id="E35939CA0D2958A495A0345F007EF2B9" pageId="0" pageNumber="4117">
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Both observed specimens are adult-stage males conforming well with the descriptions of the species available (
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<bibRefCitation id="77A256773C9F405A2ABC64D53F8DF143" author="Ehrmann, R" journalOrPublisher="Natur und Tier - Verlag, Muenster" pageId="0" pageNumber="4117" title="Mantodea: Gottesanbeterinnen der Welt." year="2002">Ehrmann 2002</bibRefCitation>
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,
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<bibRefCitation id="3A33F830CDBBD9F87BE318F41F7F2DB7" author="Battiston, Roberto" journalOrPublisher="WBA Handbooks, Verona" pageId="0" pageNumber="4117" title="Mantids of the Euro-Mediterranean Area" year="2010">Battiston et al. 2010</bibRefCitation>
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). The first is illustrated in Fig. 1. This specimen is a straw-coloured and patternless mantis with a darker forewing radial vein. Both forewings and hindwings are hyaline, forewings being slightly less transparent and brownish. Head is short with prominent conical eyes. The raptorial forelegs bear the typical spine scheme of the genus and species: 5-6, 4, 7. Biometrics: pronotum length: 10mm; whole body length: 38mm; forewing length: 26mm; fore-leg length measurements - coxa: 6.6mm, femur: 8.2mm, tibia: 4.7mm; ratio forewing/ pronotum length: 2.6.
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</paragraph>
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</subSection>
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</subSubSection>
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<subSubSection id="3A37C418D5D60393DED2110649783348" pageId="0" pageNumber="4117" type="distribution">
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<paragraph id="A68D03A96C86DD8CA08791208437E2E6" pageId="0" pageNumber="4117">Distribution</paragraph>
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<paragraph id="DB7D00B4754A235B2DB99DFC3B32A07A" pageId="0" pageNumber="4117">
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An Afrotropical species cited throughout the biogeographical area, with some island populations. Countries where it has been found are: Burkina Faso, Cameroon, Chad, Egypt, Ghana, Israel, Ivory Coast, Kenya, Mauritania, Mauritius, Mozambique, Niger, Senegal, Togo, Uganda and Zimbabwe (
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<bibRefCitation id="7530A45B6742828686A86746323890DC" author="Ehrmann, R" journalOrPublisher="Natur und Tier - Verlag, Muenster" pageId="0" pageNumber="4117" title="Mantodea: Gottesanbeterinnen der Welt." year="2002">Ehrmann 2002</bibRefCitation>
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,
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<bibRefCitation id="440AFDF3C006E4CE7B332461B2AF7D7F" author="Battiston, Roberto" journalOrPublisher="WBA Handbooks, Verona" pageId="0" pageNumber="4117" title="Mantids of the Euro-Mediterranean Area" year="2010">Battiston et al. 2010</bibRefCitation>
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).
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</paragraph>
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<subSection id="3FD496E60E8BE2976C53E89FBF6232C7" pageId="0" pageNumber="4117" type="recorded habitats and distribution in portugal">
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<paragraph id="D6B2B883C319D2778FDD33EA1BE96E1C" pageId="0" pageNumber="4117">Recorded habitats and distribution in Portugal</paragraph>
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<paragraph id="34FC08C4315E2B07F39F44E5349BB0B2" pageId="0" pageNumber="4117">
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The area is a private property garden of the resort Vila Sol where many exotic plant species are planted. Among them, hedges of
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<taxonomicName id="62DE2307718D2BDDBCCD717F341F8BFB" class="Liliopsida" family="Poaceae" genus="Phyllostachys" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phyllostachys aurea" order="Poales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="aurea">Phyllostachys aurea</taxonomicName>
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and
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<taxonomicName id="22FA2A410553BD52ED3B42A1414B1DE0" class="Magnoliopsida" family="Verbenaceae" genus="Lantana" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lantana camara" order="Lamiales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="camara">Lantana camara</taxonomicName>
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are typical while stands of
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<taxonomicName id="04EB57275AE866C83695135B8DD1A8BB" class="Liliopsida" family="Poaceae" genus="Pennisetum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pennisetum alopecuroides" order="Poales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="alopecuroides">Pennisetum alopecuroides</taxonomicName>
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,
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<taxonomicName id="DF38930AF76C3B9D08D53B99384737F5" class="Liliopsida" family="Cyperaceae" genus="Cyperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cyperus papyrus" order="Poales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="papyrus">Cyperus papyrus</taxonomicName>
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and garden turf are widespread with smaller amounts of other exotic species. The only natives are isolated
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<taxonomicName id="2C468726FC0A559D2049B57678093E4C" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus pinea" order="Pinales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="pinea">Pinus pinea</taxonomicName>
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trees, remnants from the open woodland pre-resort and a hedge of
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<taxonomicName id="021E1CA49AEC3005C320A42C35CAB8A4" class="Magnoliopsida" family="Ericaceae" genus="Arbutus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arbutus unedo" order="Ericales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="unedo">Arbutus unedo</taxonomicName>
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. At dusk, several garden lights at ground level are regularly lit and during the night, an automatic irrigation system maintains humidity levels high, even during the summer. Prior to the installment of the resort, the whole area was a dry thermomediterranean open
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<taxonomicName id="EEEBD2658820E194461E1750AA957E68" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus pinea" order="Pinales" pageId="0" pageNumber="4117" phylum="Tracheophyta" rank="species" species="pinea">Pinus pinea</taxonomicName>
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woodland with mild winters and warm summers on an arenite substrate. The biogeographic province is the Gaditano-Onubo-Algarviense in its Algarviense coastal sector with some notable endemics (
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<bibRefCitation id="392CFFB1D56EBCB03792016A54412678" author="Rivas-Martinez, S." journalOrPublisher="Itinera Geobotanica" pageId="0" pageNumber="4117" pagination="5 - 126" title="La vegetacion del sur de Portugal (Sado, Alentejo y Algarve)" volume="3" year="1990">Rivas-Martinez et al. 1990</bibRefCitation>
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,
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<bibRefCitation id="E05CAACD1A757BA69F03A426493923F5" author="Costa, J. C." journalOrPublisher="Quercetea" pageId="0" pageNumber="4117" pagination="5 - 56" title="Biogeografia de Portugal Continental" volume="0" year="1999">Costa et al. 1999</bibRefCitation>
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) and seriously under threat from urban development. This particular site is one already lost as a natural habitat and is home to only the most resilient and human-adapted species.
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</paragraph>
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</subSection>
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</subSubSection>
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<subSubSection id="56087320258CE11AE9B120B27CF60228" pageId="0" pageNumber="4117" type="ecology">
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<paragraph id="429B5AAFE8271C9FCF7498DECE6027B7" pageId="0" pageNumber="4117">Ecology</paragraph>
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<paragraph id="5E8E20BEB1F11DA0E1023E6F5B3C35D1" pageId="0" pageNumber="4117">
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Ecological plasticity in this species over a broad temperature range accounts for its wide distribution. According to
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<bibRefCitation id="4A9DAC55ECF862A84AE4EE77C9B8FBE7" author="Prete, F. R." journalOrPublisher="The Johns Hopkins University Press, Baltimore and London" pageId="0" pageNumber="4117" title="The Praying Mantids" year="1999">Prete et al. (1999)</bibRefCitation>
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, in Ghana
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<taxonomicName id="0CC059B177E366D883C39A275E9B1C23" lsidName="M. paykullii" pageId="0" pageNumber="4117" rank="species" species="paykullii">M. paykullii</taxonomicName>
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mostly inhabits grasslands, the colour of which (a proxy for humidity levels) determines the final coloration and proportion between brown and green morphs. Presumably, a brown morph develops in a less moist environment. In Ghana, adults display no deimatic behaviour and attempt to fly or walk away from danger (
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<bibRefCitation id="1E49465AC5398763E008226A43834D87" author="Edmunds, Malcolm" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="4117" pagination="1 - 32" title="Defensive behaviour in Ghanaian praying mantids" volume="51" year="1972">Edmunds 1972</bibRefCitation>
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). Adults are also presumably very mobile and active at night and especially sensitive to bat echolocation high frequency sounds (80-100 KHz), thus being able to evade predation (
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<bibRefCitation id="06DB9DF10D950A981477A499911FA9D0" author="Prete, F. R." journalOrPublisher="The Johns Hopkins University Press, Baltimore and London" pageId="0" pageNumber="4117" title="The Praying Mantids" year="1999">Prete et al. 1999</bibRefCitation>
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). This phenomenon has been experimentally tested with the closely related
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<taxonomicName id="C1229E828A5D518774327E28859B0713" lsidName="M. natalica" pageId="0" pageNumber="4117" rank="species" species="natalica">M. natalica</taxonomicName>
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Beier, 1930 by
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<bibRefCitation id="A804911394CE2E28BFCB8E75F751F90F" author="Cumming, Graeme S." journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="4117" pagination="1771 - 1774" title="Mantis movements by night and the interactions of sympatric bats and mantises" volume="74" year="1996">Cumming (1996)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="088D7764CFDD5C3A461D6170C44FC48C" pageId="0" pageNumber="4117" type="conservation">
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<paragraph id="0E3BD314D47A07F9808F2D69CEEC5A64" pageId="0" pageNumber="4117">Conservation</paragraph>
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<paragraph id="1CC83B975764FBB731419ADF04EE56CE" pageId="0" pageNumber="4117">
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Showing a wide distribution centred in the subsaharan African continent,
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<taxonomicName id="8DD1617122F6EFF53F8A457C0DA5D107" lsidName="M. paykullii" pageId="0" pageNumber="4117" rank="species" species="paykullii">M. paykullii</taxonomicName>
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should not be at risk of any kind. However, in the Euro-Mediterranean area, where it has only been found along the Nile valley and nearby areas of Israel, this species has recently been evaluated as at "Potential risk", because of sparse observations for a long time (
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<bibRefCitation id="930E7791C4515F2EB5ADDA7F655CB49C" author="Agabiti, Barbara" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa (S. E. A)" pageId="0" pageNumber="4117" pagination="1 - 20" title="The Mediterranean species of the genus Ameles Burmeister, 1838 (Insecta, Mantodea: Amelinae) with a biogeographic and phylogenetic evaluation" volume="47" year="2010">Agabiti et al. 2010</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |