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<document id="7963403CAA8FCF213ADF9F082CFAD733" ID-DOI="10.11646/zootaxa.3866.4.2" ID-GBIF-Dataset="4115b984-e9fd-44a6-ad86-05e88ab3b439" ID-ISSN="1175-5326" ID-Zenodo-Dep="249698" ID-ZooBank="D06E5477-4D5C-4402-B909-09A2AAFFB556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460919741933" checkinUser="plazi" docAuthor="Gapp, I. Wesley &amp; Lieberman, Bruce S." docDate="2014" docId="DC11878B3B4FFFC6B1AF3FF2E69021D1" docLanguage="en" docName="zt03866p498.pdf" docOrigin="Zootaxa 3866 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Olenellus fowleri Palmer 1998" docType="treatment" docVersion="8" lastPageNumber="482" masterDocId="2028FFF33B4DFFC5B1383913E4032219" masterDocTitle="New olenelloid trilobites from the Northwest Territories, Canada" masterLastPageNumber="498" masterPageNumber="479" pageNumber="481" updateTime="1698362066151" updateUser="plazi">
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<mods:title id="3F26E97508D323F9C6314DE2A6940961">New olenelloid trilobites from the Northwest Territories, Canada</mods:title>
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<mods:namePart id="F43E2233FAB5438DF6AE7D36352297EA">Gapp, I. Wesley</mods:namePart>
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<mods:namePart id="B7EE419DBCD76A5A7FCF617924BDEB09">Lieberman, Bruce S.</mods:namePart>
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<treatment id="DC11878B3B4FFFC6B1AF3FF2E69021D1" ID-DOI="http://doi.org/10.5281/zenodo.6136062" ID-GBIF-Taxon="119588927" ID-Zenodo-Dep="6136062" LSID="urn:lsid:plazi:treatment:DC11878B3B4FFFC6B1AF3FF2E69021D1" httpUri="http://treatment.plazi.org/id/DC11878B3B4FFFC6B1AF3FF2E69021D1" lastPageId="3" lastPageNumber="482" pageId="2" pageNumber="481">
<subSubSection id="1CA265163B4FFFC7B1AF3FF2E4ED2505" pageId="2" pageNumber="481" type="nomenclature">
<paragraph id="5407369D3B4FFFC7B1AF3FF2E61D24E2" blockId="2.[151,542,1760,1820]" box="[151,542,1760,1787]" pageId="2" pageNumber="481">
<heading id="0F4F81F13B4FFFC7B1AF3FF2E61D24E2" bold="true" box="[151,542,1760,1787]" fontSize="11" level="1" pageId="2" pageNumber="481" reason="1">
<emphasis id="66CCEA8F3B4FFFC7B1AF3FF2E61D24E2" bold="true" box="[151,542,1760,1787]" pageId="2" pageNumber="481">
?
<taxonomicName id="93B84D1E3B4FFFC7B19D3FF3E61D24E2" authority="Palmer, 1998" authorityName="Palmer" authorityYear="1998" box="[165,542,1760,1787]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4FFFC7B19D3FF3E57624E3" bold="true" box="[165,373,1760,1786]" italics="true" pageId="2" pageNumber="481">Olenellus fowleri</emphasis>
<bibRefCitation id="30294B6C3B4FFFC7B0443FF2E61D24E2" author="Palmer" box="[380,542,1761,1787]" pageId="2" pageNumber="481" refString="Palmer, A. R. (1998) Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada, Journal of Paleontology, 72, 650 - 672." type="journal article" year="1998">Palmer, 1998</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="5407369D3B4FFFC7B1AF3E10E4ED2505" blockId="2.[151,542,1760,1820]" box="[151,238,1795,1820]" pageId="2" pageNumber="481">Fig. 2.3</paragraph>
</subSubSection>
<subSubSection id="1CA265163B4FFFC7B1AF3E58E09F2578" box="[151,1180,1867,1889]" pageId="2" pageNumber="481" type="reference_group">
<paragraph id="5407369D3B4FFFC7B1AF3E58E09F2578" blockId="2.[151,1180,1867,1889]" box="[151,1180,1867,1889]" pageId="2" pageNumber="481">
<taxonomicName id="93B84D1E3B4FFFC7B1AF3E58E4FB2578" ID-CoL="67KY" box="[151,248,1867,1889]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="2" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B4FFFC7B1AF3E58E4FB2578" box="[151,248,1867,1889]" italics="true" pageId="2" pageNumber="481">Olenellus</emphasis>
</taxonomicName>
(
<taxonomicName id="93B84D1E3B4FFFC7B03E3E58E5642578" ID-CoL="67KY" box="[262,359,1867,1889]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="2" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B4FFFC7B03E3E58E5642578" box="[262,359,1867,1889]" italics="true" pageId="2" pageNumber="481">Olenellus</emphasis>
</taxonomicName>
?)
<taxonomicName id="93B84D1E3B4FFFC7B0BA3E58E65B2578" authority="Palmer, 1998" authorityName="Palmer" authorityYear="1998" box="[386,600,1867,1889]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4FFFC7B0BA3E58E5C92578" box="[386,458,1867,1889]" italics="true" pageId="2" pageNumber="481">fowleri</emphasis>
<bibRefCitation id="30294B6C3B4FFFC7B0E83E58E65B2578" author="Palmer" box="[464,600,1867,1889]" pageId="2" pageNumber="481" refString="Palmer, A. R. (1998) Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada, Journal of Paleontology, 72, 650 - 672." type="journal article" year="1998">Palmer, 1998</bibRefCitation>
</taxonomicName>
, p. 669, figs. 5.5, 10.110.6; see for complete synonymy.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B4FFFC7B1AF3E9EE63A25F6" pageId="2" pageNumber="481" type="materials_examined">
<paragraph id="5407369D3B4FFFC7B1AF3E9EE63A25F6" blockId="2.[151,1436,1933,2031]" pageId="2" pageNumber="481">
<emphasis id="66CCEA8F3B4FFFC7B1AF3E9EE58725BF" bold="true" box="[151,388,1933,1958]" pageId="2" pageNumber="481">Material examined.</emphasis>
<collectionCode id="32A9AE583B4FFFC7B0B63E9CE5EB25BF" box="[398,488,1935,1958]" pageId="2" pageNumber="481">KUMIP</collectionCode>
355555 likely from the
<taxonomicName id="93B84D1E3B4FFFC7B2313E9DE7B725BF" authority="Zone" authorityName="Zone" box="[777,948,1934,1958]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="2" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B4FFFC7B2313E9DE77725BF" box="[777,884,1934,1958]" italics="true" pageId="2" pageNumber="481">Olenellus</emphasis>
zone
</taxonomicName>
or Waucoban Series, Dyeran stage,
<emphasis id="66CCEA8F3B4FFFC7B4663E9CE19F25BF" box="[1374,1436,1935,1958]" italics="true" pageId="2" pageNumber="481">sensu</emphasis>
<bibRefCitation id="30294B6C3B4FFFC7B1AF3EA1E54825D3" author="Webster" box="[151,331,1970,1995]" pageId="2" pageNumber="481" refString="Webster, M. (2011 a) Trilobite biostratigraphy and sequence stratigraphy of the Upper Dyeran (traditional Laurentian &quot; Lower Cambrian &quot;) in the southern Great Basin, U. S. A. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 121 - 154." type="journal article" year="2011" yearSuffix="a">Webster (2011a</bibRefCitation>
,
<bibRefCitation id="30294B6C3B4FFFC7B0603EA1E56425D2" author="Webster" box="[344,359,1970,1995]" pageId="2" pageNumber="481" refString="Webster, M. (2011 b) Stops 7 A, 7 B, and 7 C, Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 236 - 246." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
) and Webster
<emphasis id="66CCEA8F3B4FFFC7B3363EA0E64A25D3" box="[526,585,1970,1994]" italics="true" pageId="2" pageNumber="481">et al.</emphasis>
(2011), early Cambrian, Sekwi Formation, Northwest Territories,
<collectingCountry id="2CAF760D3B4FFFC7B4793EA1E19B25D2" box="[1345,1432,1970,1995]" name="Canada" pageId="2" pageNumber="481">Canada</collectingCountry>
, but occurring from Section
<quantity id="93409B783B4FFFC7B0F43EC4E5F425F6" box="[460,503,2006,2031]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="481" unit="in" value="2.0">2 in</quantity>
float.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B4EFFC6B1FF3984E57C2359" pageId="3" pageNumber="482" type="description">
<paragraph id="5407369D3B4EFFC6B1FF3984E57C2359" blockId="3.[151,1437,151,968]" pageId="3" pageNumber="482">
<emphasis id="66CCEA8F3B4EFFC6B1FF3984E55A22A9" bold="true" box="[199,345,151,176]" pageId="3" pageNumber="482">Description.</emphasis>
The addition of the new specimen figured here does not warrant a redescription of the species as only a partial thorax is present. At least six prothoracic segments and thirty-six opisthothoracic segments are visible in this specimen. The end of a spine found most abaxially to the right on the specimen is likely part of a macropleural T3 spine. The length of the prothoracic segments are long, some are longer than the length of the entire opisthothorax.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B4EFFC6B1FF385FE6812145" pageId="3" pageNumber="482" type="discussion">
<paragraph id="5407369D3B4EFFC6B1FF385FE02323ED" blockId="3.[151,1437,151,968]" pageId="3" pageNumber="482">
<emphasis id="66CCEA8F3B4EFFC6B1FF385FE54F237C" bold="true" box="[199,332,332,357]" pageId="3" pageNumber="482">Discussion.</emphasis>
Assigning this specimen to
<taxonomicName id="93B84D1E3B4EFFC6B3BF385FE732237D" box="[647,817,332,356]" class="Trilobita" family="Olenellidae" genus="Nephrolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B4EFFC6B3BF385FE732237D" box="[647,817,332,356]" italics="true" pageId="3" pageNumber="482">Nephrolenellus</emphasis>
</taxonomicName>
(a genus known for having greater than 22 segments in the opisthothorax) is ruled out because the prothorax pleural spines are much longer in this specimen than in species of
<taxonomicName id="93B84D1E3B4EFFC6B0353887E5B423B5" box="[269,439,404,428]" class="Trilobita" family="Olenellidae" genus="Nephrolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B4EFFC6B0353887E5B423B5" box="[269,439,404,428]" italics="true" pageId="3" pageNumber="482">Nephrolenellus</emphasis>
</taxonomicName>
(and have the condition typical of
<taxonomicName id="93B84D1E3B4EFFC6B2073887E7B323B5" box="[831,944,404,428]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B2073887E7B323B5" box="[831,944,404,428]" italics="true" pageId="3" pageNumber="482">O. fowleri</emphasis>
</taxonomicName>
). Also, this specimen is much larger than is common for other
<taxonomicName id="93B84D1E3B4EFFC6B05538ABE61423C9" box="[365,535,440,464]" class="Trilobita" family="Olenellidae" genus="Nephrolenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B4EFFC6B05538ABE61423C9" box="[365,535,440,464]" italics="true" pageId="3" pageNumber="482">Nephrolenellus</emphasis>
</taxonomicName>
species, which are known for having the cephalon generally
<quantity id="93409B783B4EFFC6B5ED38ABE12523D6" box="[1237,1318,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="482" unit="mm" value="12.0">12 mm</quantity>
or shorter (sag.) (
<bibRefCitation id="30294B6C3B4EFFC6B1DC38CFE57B23ED" author="Palmer" box="[228,376,476,501]" pageId="3" pageNumber="482" refString="Palmer, A. R. (1998) Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada, Journal of Paleontology, 72, 650 - 672." type="journal article" year="1998">Palmer 1998</bibRefCitation>
). This genus is discussed in greater detail in
<bibRefCitation id="30294B6C3B4EFFC6B25438CFE01F23ED" author="Webster" box="[876,1052,476,501]" pageId="3" pageNumber="482" refString="Webster, M. (2007) Ontogeny and evolution of the Early Cambrian trilobite genus Nephrolenellus (Olenelloidea). Journal of Paleontology, 81, 1168 - 1193. http: // dx. doi. org / 10.1666 / 06 - 092.1" type="journal article" year="2007">Webster (2007)</bibRefCitation>
.
</paragraph>
<paragraph id="5407369D3B4EFFC6B1FF3B13E018209C" blockId="3.[151,1437,151,968]" pageId="3" pageNumber="482">
As
<bibRefCitation id="30294B6C3B4EFFC6B1D638ECE5932001" author="Palmer" box="[238,400,511,536]" pageId="3" pageNumber="482" refString="Palmer, A. R. (1998) Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada, Journal of Paleontology, 72, 650 - 672." type="journal article" year="1998">Palmer (1998)</bibRefCitation>
noted, the thorax of
<taxonomicName id="93B84D1E3B4EFFC6B3473B13E7CF2001" authority="Burling, 1916" authorityName="Burling" authorityYear="1916" box="[639,972,511,536]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="robsonensis">
<emphasis id="66CCEA8F3B4EFFC6B3473B13E72B2001" box="[639,808,512,536]" italics="true" pageId="3" pageNumber="482">O. robsonensis</emphasis>
<bibRefCitation id="30294B6C3B4EFFC6B21738ECE7CF2001" author="Burling" box="[815,972,511,536]" pageId="3" pageNumber="482" refString="Burling, L. D. (1916) Paedeumias and the Mesonacidae, with description of a new species, having at least 44 segments, from the Lower Cambrian of British Columbia, The Ottawa Naturalist, 30, 53 - 58." type="journal article" year="1916">Burling, 1916</bibRefCitation>
</taxonomicName>
is very similar to that of
<taxonomicName id="93B84D1E3B4EFFC6B5D63B13E1622001" box="[1262,1377,512,536]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B5D63B13E1622001" box="[1262,1377,512,536]" italics="true" pageId="3" pageNumber="482">O. fowleri</emphasis>
</taxonomicName>
. The main difference that he observed in thoracic characters was a lack of granules on either pleural spines or the axial spine of T
<quantity id="93409B783B4EFFC6B0293B54E5482079" box="[273,331,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="3" pageNumber="482" unit="in" value="15.0">15 in</quantity>
<taxonomicName id="93B84D1E3B4EFFC6B06E3B5BE5C92079" box="[342,458,584,608]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B06E3B5BE5C92079" box="[342,458,584,608]" italics="true" pageId="3" pageNumber="482">O. fowleri</emphasis>
</taxonomicName>
, and a lack of axial nodes in
<taxonomicName id="93B84D1E3B4EFFC6B2153B5BE7D42079" box="[813,983,584,608]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="robsonensis">
<emphasis id="66CCEA8F3B4EFFC6B2153B5BE7D42079" box="[813,983,584,608]" italics="true" pageId="3" pageNumber="482">O. robsonensis</emphasis>
</taxonomicName>
. Unfortunately the preservation of this specimen is too poor to observe these details, especially since T15 is not visible.
</paragraph>
<paragraph id="5407369D3B4EFFC6B1FF3B83E6812145" blockId="3.[151,1437,151,968]" pageId="3" pageNumber="482">
However, the single specimen of
<taxonomicName id="93B84D1E3B4EFFC6B3063B83E6E520B1" box="[574,742,656,680]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="robsonensis">
<emphasis id="66CCEA8F3B4EFFC6B3063B83E6E520B1" box="[574,742,656,680]" italics="true" pageId="3" pageNumber="482">O. robsonensis</emphasis>
</taxonomicName>
appears to lack the long T3 spines (see
<bibRefCitation id="30294B6C3B4EFFC6B5913B9CE16120B1" author="Lieberman" box="[1193,1378,655,680]" pageId="3" pageNumber="482" refString="Lieberman, B. S. (1999) Systematic revision of the Olenelloidea (Trilobita, Cambrian). Bulletin of the Peabody Museum of Natural History, 45, 1 - 150. [Yale University]" type="journal article" year="1999">Lieberman 1999</bibRefCitation>
) that are present in specimens of
<taxonomicName id="93B84D1E3B4EFFC6B0F43BA7E63F20D5" box="[460,572,692,716]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B0F43BA7E63F20D5" box="[460,572,692,716]" italics="true" pageId="3" pageNumber="482">O. fowleri</emphasis>
</taxonomicName>
. Thus, the small part of the very large spine (present in the upper right part of Fig. 2.3) that we interpret as part of T3 is evidence for questionably assigning this specimen to
<taxonomicName id="93B84D1E3B4EFFC6B5C43BCBE17320E9" box="[1276,1392,728,752]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B5C43BCBE17320E9" box="[1276,1392,728,752]" italics="true" pageId="3" pageNumber="482">O. fowleri</emphasis>
</taxonomicName>
(as opposed to
<taxonomicName id="93B84D1E3B4EFFC6B0213BEFE5C2210D" box="[281,449,764,788]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="robsonensis">
<emphasis id="66CCEA8F3B4EFFC6B0213BEFE5C2210D" box="[281,449,764,788]" italics="true" pageId="3" pageNumber="482">O. robsonensis</emphasis>
</taxonomicName>
). Furthermore,
<taxonomicName id="93B84D1E3B4EFFC6B34C3BEFE6E5210D" box="[628,742,764,788]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B34C3BEFE6E5210D" box="[628,742,764,788]" italics="true" pageId="3" pageNumber="482">O. fowleri</emphasis>
</taxonomicName>
probably forms a clade with several species in addition to
<taxonomicName id="93B84D1E3B4EFFC6B4B93BEFE51F2121" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="robsonensis">
<emphasis id="66CCEA8F3B4EFFC6B4B93BEFE51F2121" italics="true" pageId="3" pageNumber="482">O. robsonensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="30294B6C3B4EFFC6B0603A0CE5E92121" author="Palmer" box="[344,490,799,824]" pageId="3" pageNumber="482" refString="Palmer, A. R. (1998) Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada, Journal of Paleontology, 72, 650 - 672." type="journal article" year="1998">Palmer 1998</bibRefCitation>
, p. 669) and it is conceivable that the material illustrated herein could represent one of those species (M. Webster, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B4EFFC6B1FF3A7BE69021D1" pageId="3" pageNumber="482" type="reference_group">
<paragraph id="5407369D3B4EFFC6B1FF3A7BE69021D1" blockId="3.[151,1437,151,968]" pageId="3" pageNumber="482">
<taxonomicName id="93B84D1E3B4EFFC6B1FF3A7BE58B2199" box="[199,392,872,896]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="fowleri">
<emphasis id="66CCEA8F3B4EFFC6B1FF3A7BE58B2199" box="[199,392,872,896]" italics="true" pageId="3" pageNumber="482">Olenellus fowleri</emphasis>
</taxonomicName>
has been recovered previously from the Pioche Formation, Nevada (
<bibRefCitation id="30294B6C3B4EFFC6B5B33A74E11D2199" author="Palmer" box="[1163,1310,871,896]" pageId="3" pageNumber="482" refString="Palmer, A. R. (1957) Ontogenetic development of two olenellid trilobites. Journal of Paleontology, 31, 1." type="journal article" year="1957">Palmer 1957</bibRefCitation>
;
<bibRefCitation id="30294B6C3B4EFFC6B4123A7BE1662199" author="Palmer" box="[1322,1381,872,896]" pageId="3" pageNumber="482" refString="Palmer, A. R. (1998) Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada, Journal of Paleontology, 72, 650 - 672." type="journal article" year="1998">1998</bibRefCitation>
), the Grapevine Mountains, California (Palmer in
<bibRefCitation id="30294B6C3B4EFFC6B3A93A9FE79021BD" author="Palmer" box="[657,915,908,933]" pageId="3" pageNumber="482" refString="Palmer, A. R. &amp; Halley, R. B. (1979) Physical stratigraphy and trilobite biostratigraphy of the Carrara Formation (Lower and Middle Cambrian) in the southern Great Basin. United States Geological Survey, Professional Papers, 1047, 1 - 131." type="journal article" year="1979">Palmer &amp; Halley 1979</bibRefCitation>
), and several additional localities described in
<bibRefCitation id="30294B6C3B4EFFC6B1AF3ABCE54B21D1" author="Webster" box="[151,328,943,968]" pageId="3" pageNumber="482" refString="Webster, M. (2011 a) Trilobite biostratigraphy and sequence stratigraphy of the Upper Dyeran (traditional Laurentian &quot; Lower Cambrian &quot;) in the southern Great Basin, U. S. A. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 121 - 154." type="journal article" year="2011" yearSuffix="a">Webster (2011a</bibRefCitation>
,
<bibRefCitation id="30294B6C3B4EFFC6B06C3ABCE56021D1" author="Webster" box="[340,355,943,968]" pageId="3" pageNumber="482" refString="Webster, M. (2011 b) Stops 7 A, 7 B, and 7 C, Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. In: Hollingsworth, J. S., Sundberg, F. A. &amp; Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 236 - 246." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
) and Webster
<emphasis id="66CCEA8F3B4EFFC6B33B3AA2E63F21D1" box="[515,572,944,968]" italics="true" pageId="3" pageNumber="482">et al.</emphasis>
(2011).
</paragraph>
</subSubSection>
</treatment>
</document>