241 lines
30 KiB
XML
241 lines
30 KiB
XML
<document id="401E586889C334EFA0C1849B715825D7" ID-DOI="10.11646/zootaxa.3866.4.2" ID-GBIF-Dataset="4115b984-e9fd-44a6-ad86-05e88ab3b439" ID-ISSN="1175-5326" ID-Zenodo-Dep="249698" ID-ZooBank="D06E5477-4D5C-4402-B909-09A2AAFFB556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460919741933" checkinUser="plazi" docAuthor="Gapp, I. Wesley & Lieberman, Bruce S." docDate="2014" docId="DC11878B3B4BFFCDB1AF3AD4E64623ED" docLanguage="en" docName="zt03866p498.pdf" docOrigin="Zootaxa 3866 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesonacis wileyi Gapp & Lieberman, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="486" masterDocId="2028FFF33B4DFFC5B1383913E4032219" masterDocTitle="New olenelloid trilobites from the Northwest Territories, Canada" masterLastPageNumber="498" masterPageNumber="479" pageNumber="485" updateTime="1698362066151" updateUser="plazi">
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<mods:mods id="A39A8776561B6F2F8C032BFBB168E90E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="44F719B934C2FDE482844BB827453E59">
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<mods:title id="6679FFD3090D7C8360ED8941F1DFECB6">New olenelloid trilobites from the Northwest Territories, Canada</mods:title>
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</mods:titleInfo>
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<mods:name id="3737A77F8BC3160E7EBA161EDF1CAC33" type="personal">
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<mods:role id="EA66CF9442E14C689A94CC7A3C8EBC40">
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<mods:roleTerm id="1494FE607732C6609B2E8171F315642D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A048E74F56D527C5D3BC7736C3391313">Gapp, I. Wesley</mods:namePart>
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</mods:name>
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<mods:name id="A0CDF90B8AF45CF88464496AC3045EDC" type="personal">
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<mods:role id="A70D818C4635B1C2403EE635B19EB44A">
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<mods:roleTerm id="01CC51C938BA8172BBC326E7500B2420">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7FB28EF8E5AA4C8847BA6348B1B85BC5">Lieberman, Bruce S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DED76B958D988661E611EAD162BC8213">text</mods:typeOfResource>
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<mods:titleInfo id="EFD6CBB1AFC048EBF74C4DBBC0B01054">
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<mods:title id="9524D31A0D53F567260E665187A0DBB1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="29AA6CCA917C6F8738C92D49369460B9">
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<mods:date id="169D95EB4D9152E36CFDBBB572921E0E">2014</mods:date>
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<mods:detail id="F49F54938E5918E466B5305B7BA5FE61" type="volume">
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<mods:number id="CB6C39A3658F91FC494B94A271C110F0">3866</mods:number>
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</mods:detail>
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<mods:detail id="19EB14EFE7814BF061E00AFD33A99C33" type="issue">
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<mods:number id="5990AC39496BD8040C126C750311A7A2">4</mods:number>
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<mods:extent id="554721985452BA0332E2DB17945113B4" unit="page">
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<mods:start id="B96FDA538F61276970D1F75BD3635972">479</mods:start>
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<mods:end id="B793234F1C22A42F5B9256E37314154D">498</mods:end>
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</mods:relatedItem>
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<mods:classification id="75E284947E0BF8FBF8F393094E879309">journal article</mods:classification>
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<mods:identifier id="479AB18B6E39F4B7B3CEC62DC084AAEF" type="DOI">10.11646/zootaxa.3866.4.2</mods:identifier>
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<mods:identifier id="58E547F16B57E9D1692096B9572B629D" type="GBIF-Dataset">4115b984-e9fd-44a6-ad86-05e88ab3b439</mods:identifier>
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<mods:identifier id="8DE01BB0ED02DDA54E9EFF4AB60BEE1B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D1CAA2090130EED28567DB6B0BB2E011" type="Zenodo-Dep">249698</mods:identifier>
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<mods:identifier id="F88A2AC9A16A8D8324A3E0E03B28924F" type="ZooBank">D06E5477-4D5C-4402-B909-09A2AAFFB556</mods:identifier>
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</mods:mods>
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<treatment id="DC11878B3B4BFFCDB1AF3AD4E64623ED" ID-DOI="http://doi.org/10.5281/zenodo.6136072" ID-GBIF-Taxon="119588925" ID-Zenodo-Dep="6136072" LSID="urn:lsid:plazi:treatment:DC11878B3B4BFFCDB1AF3AD4E64623ED" httpUri="http://treatment.plazi.org/id/DC11878B3B4BFFCDB1AF3AD4E64623ED" lastPageId="8" lastPageNumber="486" pageId="6" pageNumber="485">
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<subSubSection id="1CA265163B4BFFC3B1AF3AD4E4DA261A" pageId="6" pageNumber="485" type="nomenclature">
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<paragraph id="5407369D3B4BFFC3B1AF3AD4E5CA21FB" blockId="6.[151,457,967,1027]" box="[151,457,967,994]" pageId="6" pageNumber="485">
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<heading id="0F4F81F13B4BFFC3B1AF3AD4E5CA21FB" bold="true" box="[151,457,967,994]" fontSize="11" level="1" pageId="6" pageNumber="485" reason="1">
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<emphasis id="66CCEA8F3B4BFFC3B1AF3AD4E5CA21FB" bold="true" box="[151,457,967,994]" pageId="6" pageNumber="485">
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<taxonomicName id="93B84D1E3B4BFFC3B1AF3AD4E56621F8" box="[151,357,967,993]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="6" pageNumber="485" phylum="Arthropoda" rank="species" species="wileyi" status="sp. nov.">
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<emphasis id="66CCEA8F3B4BFFC3B1AF3AD4E56621F8" bold="true" box="[151,357,967,993]" italics="true" pageId="6" pageNumber="485">Mesonacis wileyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7DFF57F43B4BFFC3B0543ADBE5CA21FB" box="[364,457,968,994]" pageId="6" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="5407369D3B4BFFC3B1AF3AF9E4DA261A" blockId="6.[151,457,967,1027]" box="[151,217,1002,1027]" pageId="6" pageNumber="485">
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<figureCitation id="CC832A183B4BFFC3B1AF3AF9E4DA261A" box="[151,217,1002,1027]" captionStart="FIGURE 4" captionStartId="7.[151,250,1617,1639]" captionTargetBox="[151,1436,373,1596]" captionTargetId="figure@7.[151,1436,373,1596]" captionTargetPageId="7" captionText="FIGURE 4. Mesonacis wileyi sp. nov. 1, holotype KUMIP 355550, x 1.9; 2, KUMIP 355551, x 2.4; 3, PWNHC- 2013.23.4, x 2.6; 4, PWNHC- 2013.23.5, x 2.4." httpUri="https://zenodo.org/record/249702/files/figure.png" pageId="6" pageNumber="485">Fig. 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1CA265163B4BFFC3B1AF3D21E6D426AE" pageId="6" pageNumber="485" type="materials_examined">
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<paragraph id="5407369D3B4BFFC3B1AF3D21E6D426AE" blockId="6.[151,1437,1074,2035]" pageId="6" pageNumber="485">
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<emphasis id="66CCEA8F3B4BFFC3B1AF3D21E5442652" bold="true" box="[151,327,1074,1099]" pageId="6" pageNumber="485">
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<typeStatus id="8B03883F3B4BFFC3B1AF3D21E4D12652" box="[151,210,1074,1099]" pageId="6" pageNumber="485">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="8B03883F3B4BFFC3B0773D21E5B42652" box="[335,439,1074,1099]" pageId="6" pageNumber="485" type="holotype">Holotype</typeStatus>
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<collectionCode id="32A9AE583B4BFFC3B0F83D20E6192653" box="[448,538,1075,1098]" pageId="6" pageNumber="485">KUMIP</collectionCode>
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355550.
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<typeStatus id="8B03883F3B4BFFC3B3BF3D20E6F72652" box="[647,756,1075,1099]" pageId="6" pageNumber="485" type="paratype">Paratypes</typeStatus>
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<collectionCode id="32A9AE583B4BFFC3B3C53D20E7542653" box="[765,855,1075,1098]" pageId="6" pageNumber="485">KUMIP</collectionCode>
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355551 and PWNHC-2013.23.4-5.
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<taxonomicName id="93B84D1E3B4BFFC3B5CB3D20E19F2653" authority="Zone" authorityName="Zone" box="[1267,1436,1075,1099]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="6" pageNumber="488" phylum="Arthropoda" rank="genus">
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<emphasis id="66CCEA8F3B4BFFC3B5CB3D20E15D2652" box="[1267,1374,1075,1099]" italics="true" pageId="6" pageNumber="485">Olenellus</emphasis>
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zone
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</taxonomicName>
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or Waucoban Series, Dyeran stage,
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<emphasis id="66CCEA8F3B4BFFC3B30B3D44E6722677" box="[563,625,1111,1134]" italics="true" pageId="6" pageNumber="485">sensu</emphasis>
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<bibRefCitation id="30294B6C3B4BFFC3B3433D45E72C2676" author="Webster" box="[635,815,1110,1135]" pageId="6" pageNumber="485" refString="Webster, M. (2011 a) Trilobite biostratigraphy and sequence stratigraphy of the Upper Dyeran (traditional Laurentian " Lower Cambrian ") in the southern Great Basin, U. S. A. In: Hollingsworth, J. S., Sundberg, F. A. & Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 121 - 154." type="journal article" year="2011" yearSuffix="a">Webster (2011a</bibRefCitation>
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,
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<bibRefCitation id="30294B6C3B4BFFC3B2053D45E74F2676" author="Webster" box="[829,844,1110,1135]" pageId="6" pageNumber="485" refString="Webster, M. (2011 b) Stops 7 A, 7 B, and 7 C, Upper Dyeran litho- and biostratigraphy of the Split Mountain area, Nevada. In: Hollingsworth, J. S., Sundberg, F. A. & Foster, J. R. (Eds.), Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada. Museum of Northern Arizona Bulletin, 67, 236 - 246." type="journal article" year="2011" yearSuffix="b">b</bibRefCitation>
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) and Webster
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<emphasis id="66CCEA8F3B4BFFC3B2CE3D44E0322677" box="[1014,1073,1110,1134]" italics="true" pageId="6" pageNumber="485">et al.</emphasis>
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(2011), early Cambrian, Sekwi Formation, Mackenzie Mountains, Northwest Territories,
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<collectingCountry id="2CAF760D3B4BFFC3B2183D69E774268A" box="[800,887,1146,1171]" name="Canada" pageId="6" pageNumber="485">Canada</collectingCountry>
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, Section 4, 430–
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<quantity id="93409B783B4BFFC3B50B3D69E071268A" box="[1075,1138,1146,1171]" metricMagnitude="2" metricUnit="m" metricValue="4.35" pageId="6" pageNumber="485" unit="m" value="435.0">435m</quantity>
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above the base of section; Section 3, 700–
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<quantity id="93409B783B4BFFC3B07E3D8CE58626AE" box="[326,389,1183,1207]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="485" unit="m" value="800.0">800m</quantity>
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above the base of the section.
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</paragraph>
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</subSubSection>
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<subSubSection id="1CA265163B4BFFC3B1FF3DD1E59026E6" pageId="6" pageNumber="485" type="etymology">
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<paragraph id="5407369D3B4BFFC3B1FF3DD1E59026E6" blockId="6.[151,1437,1074,2035]" pageId="6" pageNumber="485">
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<emphasis id="66CCEA8F3B4BFFC3B1FF3DD1E54D26C2" bold="true" box="[199,334,1218,1243]" pageId="6" pageNumber="485">Etymology.</emphasis>
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In honour of Edward O. Wiley, University of Kansas, for his fundamental contributions to the field of phylogenetics.
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</paragraph>
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</subSubSection>
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<subSubSection id="1CA265163B4BFFC3B1FF3C19E19F2772" pageId="6" pageNumber="485" type="diagnosis">
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<paragraph id="5407369D3B4BFFC3B1FF3C19E19F2772" blockId="6.[151,1437,1074,2035]" pageId="6" pageNumber="485">
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<emphasis id="66CCEA8F3B4BFFC3B1FF3C19E542273A" bold="true" box="[199,321,1290,1315]" pageId="6" pageNumber="485">Diagnosis.</emphasis>
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LA does not contact anterior cephalic border furrow; width of extraocular area approximately equal to width (tr.) of glabella at L1; posterior border between LO and intergenal angle parallel to a transverse line; intergenal angle 45 to 50 degrees relative to a transverse line; genal spines terminate posteriorly opposite T5 or T6.
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</paragraph>
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</subSubSection>
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<subSubSection id="1CA265163B4BFFC3B1FF3C65E04224EE" pageId="6" pageNumber="485" type="description">
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<paragraph id="5407369D3B4BFFC3B1FF3C65E04224EE" blockId="6.[151,1437,1074,2035]" pageId="6" pageNumber="485">
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<emphasis id="66CCEA8F3B4BFFC3B1FF3C65E55A2796" bold="true" box="[199,345,1398,1423]" pageId="6" pageNumber="485">Description.</emphasis>
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Anterior cephalic border developed as a raised ridge, length (sag.) directly anterior of LA is approximately 0.5 times length (sag.) LO; small preglabellar field; anterolateral margin of LA directed posteriorly at approximately 45 degrees; LA length (sag.) equal to the combined length (sag.) of LO and L1; ocular furrow present; ocular lobes extend back to SO; S3 convex anteriorly, conjoined adaxially; abaxial margins of L2 adaxial of abaxial margins of L3; S1 and SO both conjoined adaxially; L1 length (sag.) approximately length (sag.) of LO; abaxial margins of L1 adaxial to abaxial margins of LO; posterior margin of LO convex posteriorly; extraocular area relatively wide, with prominent anastomosing ridges; posterior border between intergenal angle and LO is parallel to a transverse line; intergenal angle deflected anteriorly at 45 to 50 degrees; genal spine narrow, developed opposite L1 or S1, extends back to T5 or T6; prothorax consists of 13(?) segments and opisthothorax about 5 or 6; pleural spines long and directed posteriorly; T3 is macropleural with spines extending posterior of pygidium; axial spine is present, questionably on T13, approximately the length (sag.) of the thorax.
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</paragraph>
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</subSubSection>
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<subSubSection id="1CA265163B4BFFC2B1FF3E11E5482359" lastPageId="7" lastPageNumber="486" pageId="6" pageNumber="485" type="discussion">
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<paragraph id="5407369D3B4BFFC2B1FF3E11E5482359" blockId="6.[151,1437,1074,2035]" lastBlockId="7.[151,1437,151,320]" lastPageId="7" lastPageNumber="486" pageId="6" pageNumber="485">
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<emphasis id="66CCEA8F3B4BFFC3B1FF3E11E54F2502" bold="true" box="[199,332,1794,1819]" pageId="6" pageNumber="485">Discussion.</emphasis>
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This species is assigned to
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<taxonomicName id="93B84D1E3B4BFFC3B3473E10E6F42503" box="[639,759,1795,1818]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="6" pageNumber="485" phylum="Arthropoda" rank="genus">
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<emphasis id="66CCEA8F3B4BFFC3B3473E10E6F42503" box="[639,759,1795,1818]" italics="true" pageId="6" pageNumber="485">Mesonacis</emphasis>
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</taxonomicName>
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due to the presence of several character states including: the anterolateral margins of LA are not prominently separated from the extraocular area by a furrow; the lateral margins of each prothoracic axial ring converge when proceeding from anterior to posterior. Further, this species can be distinguished from species of
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<taxonomicName id="93B84D1E3B4BFFC3B30D3E7DE6A3259F" box="[565,672,1902,1926]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="6" pageNumber="485" phylum="Arthropoda" rank="genus">
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<emphasis id="66CCEA8F3B4BFFC3B30D3E7DE6A3259F" box="[565,672,1902,1926]" italics="true" pageId="6" pageNumber="485">Olenellus</emphasis>
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</taxonomicName>
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based on several character states including: the transverse profile of the ocular lobe is convex dorsally; the intergenal angle is directed anteriorly approximately 45 degrees relative to a transverse line; and finally the thoracic pleural spines on all segments but the third are relatively narrow (tr.) compared with those of most species of
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<taxonomicName id="93B84D1E3B4BFFC3B3463EC8E6EA25EA" box="[638,745,2011,2035]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="6" pageNumber="485" phylum="Arthropoda" rank="genus">
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<emphasis id="66CCEA8F3B4BFFC3B3463EC8E6EA25EA" box="[638,745,2011,2035]" italics="true" pageId="6" pageNumber="485">Olenellus</emphasis>
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</taxonomicName>
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. Lastly, this species is assigned to
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<emphasis id="66CCEA8F3B4BFFC3B5AB3EC8E11125EB" box="[1171,1298,2011,2034]" italics="true" pageId="6" pageNumber="485">
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<taxonomicName id="93B84D1E3B4BFFC3B5AB3EC8E10D25EB" box="[1171,1294,2011,2034]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="6" pageNumber="485" phylum="Arthropoda" rank="genus">Mesonacis</taxonomicName>
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,
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</emphasis>
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rather than
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<taxonomicName id="93B84D1E3B4AFFC2B1AF398BE4FA22A9" box="[151,249,152,176]" class="Trilobita" family="Olenellidae" genus="Bristolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="genus">
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<emphasis id="66CCEA8F3B4AFFC2B1AF398BE4FA22A9" box="[151,249,152,176]" italics="true" pageId="7" pageNumber="486">Bristolia</emphasis>
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</taxonomicName>
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on the basis of several character states including: the lateral margins of L2, when proceeding anteriorly, do not bulge relative to LO; the surface of the interocular area does not slope evenly from the tip of the ocular lobe to the glabella; the width of the interocular area is approximately equal to the width of the ocular lobe at its midlength; and the posterior margin of the thoracic pleural furrow on the third segment is directed evenly posterolaterally.
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</paragraph>
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</subSubSection>
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<caption id="00C766153B4AFFC2B1AF3F42E5F0249F" httpUri="https://zenodo.org/record/249702/files/figure.png" pageId="7" pageNumber="486" targetBox="[151,1436,373,1596]" targetPageId="7">
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<paragraph id="5407369D3B4AFFC2B1AF3F42E5F0249F" blockId="7.[151,1436,1617,1670]" pageId="7" pageNumber="486">
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<emphasis id="66CCEA8F3B4AFFC2B1AF3F42E516247E" bold="true" box="[151,277,1617,1639]" pageId="7" pageNumber="486">FIGURE 4.</emphasis>
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<taxonomicName id="93B84D1E3B4AFFC2B0263F41E5CC247E" box="[286,463,1617,1639]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="wileyi" status="sp. nov.">
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<emphasis id="66CCEA8F3B4AFFC2B0263F41E5CC247E" box="[286,463,1617,1639]" italics="true" pageId="7" pageNumber="486">Mesonacis wileyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7DFF57F43B4AFFC2B0E03F42E628247E" box="[472,555,1617,1639]" pageId="7" pageNumber="486" rank="species">
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<emphasis id="66CCEA8F3B4AFFC2B0E03F42E628247E" bold="true" box="[472,555,1617,1639]" pageId="7" pageNumber="486">sp. nov.</emphasis>
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</taxonomicNameLabel>
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1, holotype KUMIP 355550, x1.9; 2, KUMIP 355551, x2.4; 3, PWNHC-2013.23.4, x2.6; 4, PWNHC-2013.23.5, x2.4.
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</paragraph>
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</caption>
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<subSubSection id="1CA265163B4AFFCDB1FF3FA7E64623ED" lastPageId="8" lastPageNumber="487" pageId="7" pageNumber="486" type="reference_group">
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<paragraph id="5407369D3B4AFFC2B1FF3FA7E03125BD" blockId="7.[151,1436,1715,2028]" pageId="7" pageNumber="486">
|
||
<taxonomicName id="93B84D1E3B4AFFC2B1FF3FA7E66624D5" authority="Hall, 1859" authorityName="Hall" authorityYear="1859" box="[199,613,1715,1740]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="vermontanus">
|
||
<emphasis id="66CCEA8F3B4AFFC2B1FF3FA7E5D524D2" box="[199,470,1716,1739]" italics="true" pageId="7" pageNumber="486">Mesonacis vermontanus</emphasis>
|
||
(
|
||
<bibRefCitation id="30294B6C3B4AFFC2B0DD3FA0E65E24D5" author="Hall" box="[485,605,1715,1740]" pageId="7" pageNumber="486" refString="Hall, J. (1859) Trilobites of the shales of the Hudson River Group. 12 th Annual Report of the New York Cabinet for Natural History. State of New York, Albany, New York, pp. 59 - 62." type="book chapter" year="1859">Hall, 1859</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has shorter thoracic spines and genal spines and possesses a much longer (sag.) thorax.
|
||
<taxonomicName id="93B84D1E3B4AFFC2B00D3FCBE6CF24F6" authority="Walcott, 1910" authorityName="Walcott" authorityYear="1910" box="[309,716,1750,1775]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="fremonti">
|
||
<emphasis id="66CCEA8F3B4AFFC2B00D3FCBE61024F6" box="[309,531,1751,1775]" italics="true" pageId="7" pageNumber="486">Mesonacis fremonti</emphasis>
|
||
(
|
||
<bibRefCitation id="30294B6C3B4AFFC2B31B3FC5E6C024F6" author="Walcott" box="[547,707,1750,1775]" pageId="7" pageNumber="486" refString="Walcott, C. D. (1910) Olenellus and other genera of the Mesonacidae. Smithsonian Miscellaneous Collections, 53 (6), 231 - 422." type="journal article" year="1910">Walcott, 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
possesses a genal angle further posterior than that of
|
||
<taxonomicName id="93B84D1E3B4AFFC2B40A3FCBE19F24F6" box="[1330,1436,1751,1775]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="wileyi" status="sp. nov.">
|
||
<emphasis id="66CCEA8F3B4AFFC2B40A3FCBE19F24F6" box="[1330,1436,1751,1775]" italics="true" pageId="7" pageNumber="486">M. wileyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7DFF57F43B4AFFC2B1AF3FE8E4F1250D" box="[151,242,1787,1812]" pageId="7" pageNumber="486" rank="species">
|
||
<emphasis id="66CCEA8F3B4AFFC2B1AF3FE8E4F1250D" bold="true" box="[151,242,1787,1812]" pageId="7" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, a posterior border directed posteriorly between LO and the intergenal angle, and a prominent intergenal ridge.
|
||
<taxonomicName id="93B84D1E3B4AFFC2B1E63E33E702252E" authority="Resser and Howell, 1938" authorityName="Resser and Howell" authorityYear="1938" box="[222,769,1822,1847]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="bonnensis">
|
||
<emphasis id="66CCEA8F3B4AFFC2B1E63E33E5CE252E" box="[222,461,1823,1847]" italics="true" pageId="7" pageNumber="486">Mesonacis bonnensis</emphasis>
|
||
(
|
||
<bibRefCitation id="30294B6C3B4AFFC2B0E43E0CE6FB252E" author="Resser" box="[476,760,1822,1847]" pageId="7" pageNumber="486" refString="Resser, C. E. & Howell, B. F. (1938) Lower Cambrian Olenellus zone of the Appalachians. Geological Society of America, Bulletin, 49, 195 - 248. http: // dx. doi. org / 10.1130 / GSAB- 49 - 195" type="journal article" year="1938">Resser and Howell, 1938</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has a narrower extraocular area and a prominent intergenal ridge.
|
||
<taxonomicName id="93B84D1E3B4AFFC2B1DC3E57E67D2545" authority="Best, 1952" authorityName="Best" authorityYear="1952" box="[228,638,1859,1884]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="eagerensis">
|
||
<emphasis id="66CCEA8F3B4AFFC2B1DC3E57E5E22542" box="[228,481,1860,1883]" italics="true" pageId="7" pageNumber="486">Mesonacis eagerensis</emphasis>
|
||
(
|
||
<bibRefCitation id="30294B6C3B4AFFC2B0CE3E50E6752545" author="Best" box="[502,630,1859,1884]" pageId="7" pageNumber="486" refString="Best, R. V. (1952) Two new species of Olenellus from British Columbia. Transactions of the Royal Society of Canada, 46, 13 - 22." type="journal article" year="1952">Best, 1952</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
possesses a narrower extraocular area and narrower pleural lobes.
|
||
<taxonomicName id="93B84D1E3B4AFFC2B1AF3E7BE6E32566" authority="Cowie & McNamara, 1978" authorityName="Cowie & McNamara" authorityYear="1978" box="[151,736,1894,1919]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="hamoculus">
|
||
<emphasis id="66CCEA8F3B4AFFC2B1AF3E7BE5932566" box="[151,400,1895,1919]" italics="true" pageId="7" pageNumber="486">Mesonacis hamoculus</emphasis>
|
||
(
|
||
<bibRefCitation id="30294B6C3B4AFFC2B0983E75E6DB2566" author="Cowie" box="[416,728,1894,1919]" pageId="7" pageNumber="486" refString="Cowie, J. W. & McNamara, K. J. (1978) Olenellus (Trilobita) from the Lower Cambrian strata of north-west Scotland. Palaeontology, 21, 615 - 634." type="journal article" year="1978">Cowie & McNamara, 1978</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has a prominent occipital spine and a narrower (tr.) posterior border between LO and the intergenal angle, which is directed slightly posteriorly.
|
||
</paragraph>
|
||
<paragraph id="5407369D3B4AFFCDB1FF3EA3E64623ED" blockId="7.[151,1436,1715,2028]" lastBlockId="8.[151,1437,151,501]" lastPageId="8" lastPageNumber="487" pageId="7" pageNumber="486">
|
||
<taxonomicName id="93B84D1E3B4AFFC2B1FF3EA3E5D225DE" box="[199,465,1967,1991]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="cylindricus">
|
||
<emphasis id="66CCEA8F3B4AFFC2B1FF3EA3E5D225DE" box="[199,465,1967,1991]" italics="true" pageId="7" pageNumber="486">Mesonacis cylindricus</emphasis>
|
||
</taxonomicName>
|
||
(Palmer in
|
||
<bibRefCitation id="30294B6C3B4AFFC2B3B83EBCE7B125DE" author="Palmer" box="[640,946,1967,1992]" pageId="7" pageNumber="486" refString="Palmer, A. R. & Halley, R. B. (1979) Physical stratigraphy and trilobite biostratigraphy of the Carrara Formation (Lower and Middle Cambrian) in the southern Great Basin. United States Geological Survey, Professional Papers, 1047, 1 - 131." type="journal article" year="1979">Palmer & Halley, 1979</bibRefCitation>
|
||
), which
|
||
<bibRefCitation id="30294B6C3B4AFFC2B5163EBCE10B25DE" author="Lieberman" box="[1070,1288,1967,1992]" pageId="7" pageNumber="486" refString="Lieberman, B. S. (1999) Systematic revision of the Olenelloidea (Trilobita, Cambrian). Bulletin of the Peabody Museum of Natural History, 45, 1 - 150. [Yale University]" type="journal article" year="1999">Lieberman (1999)</bibRefCitation>
|
||
determined phylogenetically to be a more derived species within the genus, appears to be most similar to
|
||
<taxonomicName id="93B84D1E3B4AFFC2B59D3EC7E10E25F2" box="[1189,1293,2003,2027]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="486" phylum="Arthropoda" rank="species" species="wileyi">
|
||
<emphasis id="66CCEA8F3B4AFFC2B59D3EC7E10E25F2" box="[1189,1293,2003,2027]" italics="true" pageId="7" pageNumber="486">M. wileyi</emphasis>
|
||
</taxonomicName>
|
||
. Similarities include: extraocular area that is wider than the width of the glabella (tr.); LA does not contact the anterior cephalic border furrow; the adaxial and abaxial tips of S2 are on a transverse line; the lateral border of L2 diverges anteriorly; SO is conjoined adaxially (although this is not observed in the smallest specimen of
|
||
<taxonomicName id="93B84D1E3B45FFCDB5DC39F2E14C22E1" box="[1252,1359,224,248]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="wileyi">
|
||
<emphasis id="66CCEA8F3B45FFCDB5DC39F2E14C22E1" box="[1252,1359,224,248]" italics="true" pageId="8" pageNumber="487">M. wileyi</emphasis>
|
||
</taxonomicName>
|
||
which represents an earlier ontogenetic stage); and the intergenal angle is 45 to 50 degrees relative to a transverse line. Some key differences between the species observed include: the posterior margin of the ocular lobe is opposite SO in
|
||
<taxonomicName id="93B84D1E3B45FFCDB18E385EE522237D" box="[182,289,332,356]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="wileyi">
|
||
<emphasis id="66CCEA8F3B45FFCDB18E385EE522237D" box="[182,289,332,356]" italics="true" pageId="8" pageNumber="487">M. wileyi</emphasis>
|
||
</taxonomicName>
|
||
and opposite the adaxial part of L
|
||
<quantity id="93409B783B45FFCDB38F385FE6E3237C" box="[695,736,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="487" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="93B84D1E3B45FFCDB3D2385EE78D237D" box="[746,910,332,356]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="cylindricus">
|
||
<emphasis id="66CCEA8F3B45FFCDB3D2385EE78D237D" box="[746,910,332,356]" italics="true" pageId="8" pageNumber="487">M. cylindricus</emphasis>
|
||
</taxonomicName>
|
||
; S
|
||
<quantity id="93409B783B45FFCDB296385FE7DE237C" box="[942,989,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="487" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="93B84D1E3B45FFCDB2DE385EE052237D" box="[998,1105,332,356]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="wileyi">
|
||
<emphasis id="66CCEA8F3B45FFCDB2DE385EE052237D" box="[998,1105,332,356]" italics="true" pageId="8" pageNumber="487">M. wileyi</emphasis>
|
||
</taxonomicName>
|
||
is more prominently incised adaxially; the adaxial part of the cephalic border between the lateral margins of LO and the intergenal angle parallels a transverse line rather than being directed posteriorly (though again this is not observed in the smallest specimen of
|
||
<emphasis id="66CCEA8F3B45FFCDB01038AAE59923C9" box="[296,410,440,464]" italics="true" pageId="8" pageNumber="487">
|
||
<taxonomicName id="93B84D1E3B45FFCDB01038AAE59123C9" box="[296,402,440,464]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="wileyi">M. wileyi</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
; and lastly the direction of the ocular lobes of
|
||
<taxonomicName id="93B84D1E3B45FFCDB29538AAE01423C9" box="[941,1047,440,464]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="wileyi">
|
||
<emphasis id="66CCEA8F3B45FFCDB29538AAE01423C9" box="[941,1047,440,464]" italics="true" pageId="8" pageNumber="487">M. wileyi</emphasis>
|
||
</taxonomicName>
|
||
do not form as large an angle to a sagittal line as those in
|
||
<taxonomicName id="93B84D1E3B45FFCDB0A538CEE63D23ED" box="[413,574,476,500]" class="Trilobita" family="Olenellidae" genus="Mesonacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="8" pageNumber="487" phylum="Arthropoda" rank="species" species="cylindricus">
|
||
<emphasis id="66CCEA8F3B45FFCDB0A538CEE63D23ED" box="[413,574,476,500]" italics="true" pageId="8" pageNumber="487">M. cylindricus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |