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<document id="4BF88084E2B81B7A3791315F58ED42C0" ID-CLB-Dataset="42479" ID-DOI="10.11646/zootaxa.3866.4.2" ID-GBIF-Dataset="4115b984-e9fd-44a6-ad86-05e88ab3b439" ID-ISSN="1175-5326" ID-Zenodo-Dep="249698" ID-ZooBank="D06E5477-4D5C-4402-B909-09A2AAFFB556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" checkinTime="1460919741933" checkinUser="plazi" docAuthor="Gapp, I. Wesley &amp; Lieberman, Bruce S." docDate="2014" docId="DC11878B3B47FFCFB1AF38C5E093241B" docLanguage="en" docName="zt03866p498.pdf" docOrigin="Zootaxa 3866 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holmiella taurus Gapp &amp; Lieberman, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="489" masterDocId="2028FFF33B4DFFC5B1383913E4032219" masterDocTitle="New olenelloid trilobites from the Northwest Territories, Canada" masterLastPageNumber="498" masterPageNumber="479" pageNumber="489" updateTime="1728481819383" updateUser="valdenar">
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<mods:title id="8A3C30140CE1825DBBA4E0949287AB3E">New olenelloid trilobites from the Northwest Territories, Canada</mods:title>
</mods:titleInfo>
<mods:name id="96E247166942EC623AC5F20F2DE4C994" type="personal">
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<mods:namePart id="2BCDD4CD02C9A57813F4CB63F755B3D8">Gapp, I. Wesley</mods:namePart>
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<mods:namePart id="54B31CCF51992FDF99EB89A575EEB31A">Lieberman, Bruce S.</mods:namePart>
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<mods:typeOfResource id="9CBA64A0EF7A8A179814E9D2B7D7F23E">text</mods:typeOfResource>
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<mods:title id="DAF2671C2D39B88C18B3406379DA2ED0">Zootaxa</mods:title>
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<mods:date id="1F15A79FA0B16E257E9E5F1B7A288129">2014</mods:date>
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<mods:identifier id="379CF07783DC1CCF62412C71C79F2768" type="DOI">10.11646/zootaxa.3866.4.2</mods:identifier>
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<treatment id="DC11878B3B47FFCFB1AF38C5E093241B" ID-DOI="http://doi.org/10.5281/zenodo.6136086" ID-GBIF-Taxon="119588943" ID-Zenodo-Dep="6136086" LSID="urn:lsid:plazi:treatment:DC11878B3B47FFCFB1AF38C5E093241B" httpUri="http://treatment.plazi.org/id/DC11878B3B47FFCFB1AF38C5E093241B" lastPageNumber="489" pageId="10" pageNumber="489">
<subSubSection id="1CA265163B47FFCFB1AF38C5E4DA200B" pageId="10" pageNumber="489" type="nomenclature">
<paragraph id="5407369D3B47FFCFB1AF38C5E5CA23E9" blockId="10.[151,457,470,530]" box="[151,457,470,496]" pageId="10" pageNumber="489">
<heading id="0F4F81F13B47FFCFB1AF38C5E5CA23E9" bold="true" box="[151,457,470,496]" fontSize="11" level="1" pageId="10" pageNumber="489" reason="1">
<emphasis id="66CCEA8F3B47FFCFB1AF38C5E5CA23E9" bold="true" box="[151,457,470,496]" pageId="10" pageNumber="489">
<taxonomicName id="93B84D1E3B47FFCFB1AF38C5E56623E9" box="[151,357,470,496]" class="Trilobita" family="Olenellidae" genus="Holmiella" higherTaxonomySource="CoL" kingdom="Animalia" order="Redlichiida" pageId="10" pageNumber="489" phylum="Arthropoda" rank="species" species="taurus" status="sp. nov.">
<emphasis id="66CCEA8F3B47FFCFB1AF38C5E56623E9" bold="true" box="[151,357,470,496]" italics="true" pageId="10" pageNumber="489">Holmiella taurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B47FFCFB05438C5E5CA23E9" box="[364,457,470,496]" pageId="10" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5407369D3B47FFCFB1AF38EAE4DA200B" blockId="10.[151,457,470,530]" box="[151,217,505,530]" pageId="10" pageNumber="489">
<figureCitation id="CC832A183B47FFCFB1AF38EAE4DA200B" box="[151,217,505,530]" captionStart="FIGURE 6" captionStartId="11.[151,250,1953,1975]" captionTargetBox="[171,1416,193,1930]" captionTargetId="figure@11.[171,1417,193,1932]" captionTargetPageId="11" captionText="FIGURE 6. Holmiella taurus sp. nov. 1, holotype KUMIP 355545, x 1.4; 2, PWNHC 2013.23. 21 (latex cast), x 2.1; 3, KUMIP 355546, x 1.7; 4, PWNHC- 2013.23.22, x 0.9." httpUri="https://zenodo.org/record/249704/files/figure.png" pageId="10" pageNumber="489">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1AF3B52E1582040" box="[151,1371,577,602]" pageId="10" pageNumber="489" type="materials_examined">
<paragraph id="5407369D3B47FFCFB1AF3B52E1582040" blockId="10.[151,1437,577,1538]" box="[151,1371,577,602]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1AF3B52E5462043" bold="true" box="[151,325,577,602]" pageId="10" pageNumber="489">
<typeStatus id="8B03883F3B47FFCFB1AF3B52E4D12043" box="[151,210,577,602]" pageId="10" pageNumber="489">Type</typeStatus>
material.
</emphasis>
<typeStatus id="8B03883F3B47FFCFB0743B52E5B72043" box="[332,436,577,602]" pageId="10" pageNumber="489" type="holotype">Holotype</typeStatus>
<collectionCode id="32A9AE583B47FFCFB0833B51E6162040" box="[443,533,578,601]" pageId="10" pageNumber="489">KUMIP</collectionCode>
355545.
<typeStatus id="8B03883F3B47FFCFB3463B51E6E82043" box="[638,747,578,602]" pageId="10" pageNumber="489" type="paratype">Paratypes</typeStatus>
<collectionCode id="32A9AE583B47FFCFB3CA3B51E74F2040" box="[754,844,578,601]" pageId="10" pageNumber="489">KUMIP</collectionCode>
355546355547 and
<collectionCode id="32A9AE583B47FFCFB5783B52E0A62040" box="[1088,1189,577,601]" pageId="10" pageNumber="489">PWNHC</collectionCode>
2013.23.2122.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1FF3B76E13B2067" box="[199,1336,613,638]" pageId="10" pageNumber="489" type="etymology">
<paragraph id="5407369D3B47FFCFB1FF3B76E13B2067" blockId="10.[151,1437,577,1538]" box="[199,1336,613,638]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3B76E54D2067" bold="true" box="[199,334,613,638]" pageId="10" pageNumber="489">Etymology.</emphasis>
The name
<taxonomicName id="93B84D1E3B47FFCFB0F33B75E6112064" box="[459,530,614,637]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="species" species="taurus">
<emphasis id="66CCEA8F3B47FFCFB0F33B75E6112064" box="[459,530,614,637]" italics="true" pageId="10" pageNumber="489">taurus</emphasis>
</taxonomicName>
was chosen because the large genal spines resemble the horns of a bull.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1FF3B9AE6732117" pageId="10" pageNumber="489" type="diagnosis">
<paragraph id="5407369D3B47FFCFB1FF3B9AE6732117" blockId="10.[151,1437,577,1538]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3B9AE54220BB" bold="true" box="[199,321,649,674]" pageId="10" pageNumber="489">Diagnosis.</emphasis>
Anterior constriction of cephalic border between intergenal spine and genal spine, where border meets genal spine; strongly incised S3; short ocular lobes (terminate opposite S1); anterior most part of genal spine is anterior of LA; anterior cephalic margin is approximately parallel to a sagittal line; the posterolateral margin of LA is constricted to subparallel anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1FF3A0AE60C2683" pageId="10" pageNumber="489" type="description">
<paragraph id="5407369D3B47FFCFB1FF3A0AE60C2683" blockId="10.[151,1437,577,1538]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3A0AE55A212B" bold="true" box="[199,345,793,818]" pageId="10" pageNumber="489">Description.</emphasis>
Anterior cephalic margin relatively straight, extending to genal spines, parallel to the transverse line, developed as a flattened ledge; anterior cephalic border directly anterior of LA length approximately equal to length L3 and expands greatly abaxially; genal spine where attached subparallel to transverse line and then further abaxially deflected back strongly, directed slightly inward, posterior of the posterior cephalic border; length (sag.) of LA approximately equal to combined length (sag.) of L1, L2, and L3; anterior margin of LA semicircular in shape; lateral margins of LA abaxial of lateral margins of LO; anterior margin of ocular lobe separated from LA by a furrow; S3 convex anteriorly; lateral margin of L3 convex abaxially; length (sag.) of L3 equal to length of L2; S2 convex anteriorly; lateral margin of L2 convex abaxially; S1 convex anteriorly; length (sag.) of L1 is approximately 1.5 times the length of L2; lateral margins of L1 adaxial of lateral margins of LO; length (sag.) of LO is approximately length (sag.) L2 and L3; small occipital spine present; small intergenal spines present on posterior border angled abaxially.
</paragraph>
</subSubSection>
<subSubSection id="1CA265163B47FFCFB1FF3DB6E093241B" pageId="10" pageNumber="489" type="discussion">
<paragraph id="5407369D3B47FFCFB1FF3DB6E137278F" blockId="10.[151,1437,577,1538]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3DB6E54F26A7" bold="true" box="[199,332,1189,1214]" pageId="10" pageNumber="489">Discussion.</emphasis>
This species differs from numerous other species of
<taxonomicName id="93B84D1E3B47FFCFB2933DB6E01826A4" box="[939,1051,1189,1213]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="genus">
<emphasis id="66CCEA8F3B47FFCFB2933DB6E01826A4" box="[939,1051,1189,1213]" italics="true" pageId="10" pageNumber="489">Holmiella</emphasis>
</taxonomicName>
by various characters such as the anterior cephalic margin being nearly parallel to a transverse line, short ocular lobes that terminate posteriorly at S1, and the posterolateral margin of LA being constricted to subparallel to a sagittal line. The cephalic border between the intergenal spine and genal spine constricts anteriorly, similarly to
<taxonomicName id="93B84D1E3B47FFCFB5323C01E0552730" box="[1034,1110,1297,1321]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="species" species="falx">
<emphasis id="66CCEA8F3B47FFCFB5323C01E0552730" box="[1034,1110,1297,1321]" italics="true" pageId="10" pageNumber="489">H. falx</emphasis>
</taxonomicName>
. Like
<taxonomicName id="93B84D1E3B47FFCFB5A73C01E1372730" box="[1183,1332,1297,1321]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="species" species="domackae" status="sp. nov.">
<emphasis id="66CCEA8F3B47FFCFB5A73C01E1372730" box="[1183,1332,1297,1321]" italics="true" pageId="10" pageNumber="489">H. domackae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DFF57F43B47FFCFB4033C02E1962733" box="[1339,1429,1297,1322]" pageId="10" pageNumber="489" rank="species">
<emphasis id="66CCEA8F3B47FFCFB4033C02E1962733" bold="true" box="[1339,1429,1297,1322]" pageId="10" pageNumber="489">sp. nov.</emphasis>
</taxonomicNameLabel>
, the genal spines are prominently directed posteriorly, however the ends of the genal spines in
<taxonomicName id="93B84D1E3B47FFCFB5913C25E1102754" box="[1193,1299,1334,1357]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="species" species="taurus">
<emphasis id="66CCEA8F3B47FFCFB5913C25E1102754" box="[1193,1299,1334,1357]" italics="true" pageId="10" pageNumber="489">H. taurus</emphasis>
</taxonomicName>
are directed more adaxially. Intergenal and occipital spines are present as they are in other species from the Sekwi Formation (
<taxonomicName id="93B84D1E3B47FFCFB1983C6DE537278C" box="[160,308,1405,1429]" class="Dothideomycetes" family="Patellariaceae" genus="Holmiella" kingdom="Fungi" order="Patellariales" pageId="10" pageNumber="489" phylum="Ascomycota" rank="species" species="domackae">
<emphasis id="66CCEA8F3B47FFCFB1983C6DE537278C" box="[160,308,1405,1429]" italics="true" pageId="10" pageNumber="489">H. domackae</emphasis>
</taxonomicName>
included), which is not the case for species recovered from Nevada (
<bibRefCitation id="30294B6C3B47FFCFB5013C6EE12B278F" author="Hollingsworth" box="[1081,1320,1405,1430]" pageId="10" pageNumber="489" refString="Hollingsworth, J. S. (2006) Holmiidae (Trilobita: Olenellina) of the Montezuman Stage (Early Cambrian) in western Nevada. Journal of Paleontology, 80, 309 - 332. http: // dx. doi. org / 10.1666 / 0022 - 3360 (2006) 080 [0309: HTOOTM] 2.0. CO; 2" type="journal article" year="2006">Hollingsworth, 2006</bibRefCitation>
).
</paragraph>
<paragraph id="5407369D3B47FFCFB1FF3CB2E093241B" blockId="10.[151,1437,577,1538]" pageId="10" pageNumber="489">
<emphasis id="66CCEA8F3B47FFCFB1FF3CB2E55A27A3" bold="true" box="[199,345,1441,1466]" pageId="10" pageNumber="489">Occurrence.</emphasis>
Early Cambrian, Sekwi Formation, Mackenzie Mountains, Northwest Territories,
<collectingCountry id="2CAF760D3B47FFCFB4793CB2E19B27A3" box="[1345,1432,1441,1466]" name="Canada" pageId="10" pageNumber="489">Canada</collectingCountry>
, Section 14, in float. Given that this material occurs in float it was impossible to determine whether it came from the Dyeran or underlying Montezuman stages (or
<taxonomicName id="93B84D1E3B47FFCFB3A43CFAE70B2418" box="[668,776,1513,1537]" class="Trilobita" family="Olenellidae" genus="Olenellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="10" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B47FFCFB3A43CFAE70B2418" box="[668,776,1513,1537]" italics="true" pageId="10" pageNumber="489">Olenellus</emphasis>
</taxonomicName>
or
<taxonomicName id="93B84D1E3B47FFCFB2153CFAE79C2418" box="[813,927,1513,1537]" class="Trilobita" family="Holmiidae" genus="Nevadella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Redlichiida" pageId="10" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="66CCEA8F3B47FFCFB2153CFAE79C2418" box="[813,927,1513,1537]" italics="true" pageId="10" pageNumber="489">Nevadella</emphasis>
</taxonomicName>
stages, respectively).
</paragraph>
</subSubSection>
</treatment>
</document>