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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.70.762" ID-GBIF-Dataset="b8fceb5a-9f78-4dc4-8fa2-92fe20bb2c42" ID-PMC="PMC3088446" ID-Pensoft-Pub="1313-2970-70-1" ID-PubMed="21594041" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-70-1" ModsDocOrigin="ZooKeys 70" ModsDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an “oceanic” island" checkinTime="1451250880545" checkinUser="pensoft" docAuthor="Rowson, B, Warren, B. H. &amp; Ngereza, C. F." docDate="2010" docId="ACCDA2231D1679BAFD1071877F2BC44B" docLanguage="en" docName="ZooKeys 70: 1-39" docOrigin="ZooKeys 70" docSource="http://dx.doi.org/10.3897/zookeys.70.762" docTitle="Tropidophora zanguebarica Petit 1850" docType="treatment" docVersion="5" lastPageNumber="11" masterDocId="FFB2FFB6591BFFBAFFFF0472FFE6A56D" masterDocTitle="Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island" masterLastPageNumber="39" masterPageNumber="1" pageNumber="11" updateTime="1668164835731" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Terrestrial molluscs of Pemba Island, Zanzibar, Tanzania, and its status as an &quot; oceanic &quot; island</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rowson, B</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Warren, B. H.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ngereza, C. F.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>70</mods:number>
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<mods:start>1</mods:start>
<mods:end>39</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.70.762</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.70.762</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-70-1</mods:identifier>
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<treatment ID-GBIF-Taxon="159360777" LSID="urn:lsid:plazi:treatment:ACCDA2231D1679BAFD1071877F2BC44B" httpUri="http://treatment.plazi.org/id/ACCDA2231D1679BAFD1071877F2BC44B" lastPageId="12" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection pageId="10" pageNumber="11" type="nomenclature">
<paragraph pageId="10" pageNumber="11">
5.
<taxonomicName authority="Petit, 1850" authorityName="Petit" authorityYear="1850" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidophora zanguebarica" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="zanguebarica">Tropidophora zanguebarica (Petit, 1850)</taxonomicName>
Fig. 3
</paragraph>
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<subSubSection pageId="10" pageNumber="11" type="reference_group">
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Gastropoda" family="Pomatiidae" genus="Cyclostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclostoma zanguebarica" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="zanguebarica">Cyclostoma zanguebarica</taxonomicName>
<bibRefCitation author="Petit de la Saussaye, M" journalOrPublisher="Journal de Conchyliologie" pageId="23" pageNumber="24" pagination="36 - 55" title="Notice sur le genre Cyclostoma, et Catalogue des especes appartenant a ce genre." volume="1" year="1850">Petit de la Saussaye 1850</bibRefCitation>
: 53; pl. III, fig. 5
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="notes">
<paragraph pageId="10" pageNumber="11">Notes.</paragraph>
<paragraph lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
This group needs revision. Pemba shells are almost identical to those from Jozani Forest, Unguja, including in microsculpture and the range of colour patterns, differing mainly
<pageBreakToken pageId="11" pageNumber="12" start="start">in</pageBreakToken>
a smaller maximum size and (Unguja 14.0
<normalizedToken originalValue="×">x</normalizedToken>
12.5 mm; Pemba 12.0
<normalizedToken originalValue="×">x</normalizedToken>
11.5 mm). There is slight variation in the strength of the spiral raised ridges, though none of the shells are as smooth as zanguebarica Petit, 1850 or letourneuxi Bourguignat, 1889 in historical collections (e.g. NMW, MNHN). It is hard to know whether this is infraspecific variation or not.
<bibRefCitation author="Voeltzkow, A" journalOrPublisher="Witu-Inseln und Zanzibar-Archipel. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart" pageId="24" pageNumber="25" title="Reise in Ostafrika in den Jahren 1903 - 1905. Wissenschaftliche Ergebnisse" year="1923">Voeltzkow (1923)</bibRefCitation>
recorded &quot;
<taxonomicName genus="Lygatella" lsidName="Lygatella letourneuxi" pageId="11" pageNumber="12" rank="species" species="letourneuxi">Lygatella letourneuxi</taxonomicName>
(Bgl.)&quot; (sic) from Pemba;
<bibRefCitation author="Haas, F" journalOrPublisher="Zoologische Jahrbuecher" pageId="23" pageNumber="24" pagination="387 - 430" title="Die Binnenmollusken der Voeltzkow'schen Reisen in Ostafrika und den ostafrikanischen Inseln" volume="57" year="1929">Haas (1929)</bibRefCitation>
recorded it from Chake Chake. As well as zanguebarica and letourneuxi,
<bibRefCitation author="Verdcourt, B" journalOrPublisher="B Verdcourt, Maidenhead, UK" pageId="24" pageNumber="25" title="A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi)" year="2006">Verdcourt (2006)</bibRefCitation>
lists two unnamed
<normalizedToken originalValue="“species”">&quot;species&quot;</normalizedToken>
from
<normalizedToken originalValue="“Zanzibar”">&quot;Zanzibar&quot;</normalizedToken>
. Either could correspond
<pageBreakToken pageId="12" pageNumber="13" start="start">to</pageBreakToken>
the Pemba taxon; possibly one biological species encompasses all four. One is said to have an affinity to letourneuxi and to
<taxonomicName class="Gastropoda" family="Pomatiidae" genus="Tropidophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidophora scabra" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="scabra">Tropidophora scabra</taxonomicName>
(H. Adams, 1867), an extinct Mauritian species that varies in sculpture according to
<bibRefCitation author="Griffiths, OL" journalOrPublisher="Bioculture Press, Mauritius" pageId="23" pageNumber="24" title="A field guide to the non-marine molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion) and the Northern Dependencies of Mauritius" year="2006">Griffiths and Florens (2006)</bibRefCitation>
. Intriguingly, these authors describe (p.53) a population of a smooth species of
<taxonomicName class="Gastropoda" family="Pomatiidae" genus="Tropidophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidophora" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Tropidophora</taxonomicName>
being replaced by a rough one over recent decades. Note: Verdcourt treats all East African
<taxonomicName class="Gastropoda" family="Pomatiidae" genus="Tropidophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidophora" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Tropidophora</taxonomicName>
in subgenus
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Otopoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otopoma" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Otopoma</taxonomicName>
Gray, 1850, but the Asian type species of this belongs in Cyclophoridae not Pomatiidae (=Pomatiasidae) (see
<bibRefCitation author="Neubert, E" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="23" pageNumber="24" pagination="93 - 96" title="Otopoma Gray 1850 - a few words to add to a 150 years old debate" volume="132" year="2003">Neubert 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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