520 lines
62 KiB
XML
520 lines
62 KiB
XML
<document id="4C05BB5B0B96A0961F7DF34E2B94913A" ID-DOI="10.11646/zootaxa.3841.3.4" ID-GBIF-Dataset="fd4f8193-3005-4d9b-ae7f-304b847215a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="225778" ID-ZooBank="E52D2F6B-631D-45FE-97E0-DD26137757FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460462027068" checkinUser="plazi" docAuthor="Bianco, Pier Giorgio & Ketmaier, Valerio" docDate="2014" docId="8E3D87DAFFC9FFA066A3F948FAF9FE3C" docLanguage="en" docName="zt03841p402.pdf" docOrigin="Zootaxa 3841 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucos basak Heckel 1843" docType="treatment" docVersion="8" lastPageNumber="387" masterDocId="7204FFA2FFCCFFA86634FFEBFF9AFFB4" masterDocTitle="A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)" masterLastPageNumber="402" masterPageNumber="379" pageNumber="384" updateTime="1698304242431" updateUser="plazi">
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<mods:titleInfo id="3C64BA216106AE6F13F3C6D75263B68B">
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<mods:title id="C9D137B15A008298EC0962D089B2974A">A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)</mods:title>
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<mods:namePart id="23E9287542CC03E319021D61467DB9FD">Bianco, Pier Giorgio</mods:namePart>
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<mods:namePart id="45ECA31BCF670E0CBE21AD54A6B88FE5">Ketmaier, Valerio</mods:namePart>
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<mods:title id="24A788A09E44A06FF3AD70371939EA8B">Zootaxa</mods:title>
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<mods:part id="34CB91B1A745C0D99D619E1CF380BA7E">
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<mods:date id="A31EF17C33335C278D0667006941A672">2014</mods:date>
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<mods:detail id="6E1E0E3FADB136CBC071A79721AD247B" type="volume">
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<mods:number id="29264FAC5152F8188A030F3775DFA0E0">3841</mods:number>
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<mods:identifier id="4642EEA8CAD8D2F600A6C1C3D38C79EC" type="DOI">10.11646/zootaxa.3841.3.4</mods:identifier>
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<treatment id="8E3D87DAFFC9FFA066A3F948FAF9FE3C" ID-DOI="http://doi.org/10.5281/zenodo.6140503" ID-GBIF-Taxon="119512643" ID-Zenodo-Dep="6140503" LSID="urn:lsid:plazi:treatment:8E3D87DAFFC9FFA066A3F948FAF9FE3C" httpUri="http://treatment.plazi.org/id/8E3D87DAFFC9FFA066A3F948FAF9FE3C" lastPageId="8" lastPageNumber="387" pageId="5" pageNumber="384">
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<subSubSection id="4E8E6547FFC9FFAD66A3F948FEADF96B" pageId="5" pageNumber="384" type="nomenclature">
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<paragraph id="062B36CCFFC9FFAD66A3F948FE78F90A" blockId="5.[151,482,1699,1759]" box="[151,482,1699,1726]" pageId="5" pageNumber="384">
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<heading id="5D6381A0FFC9FFAD66A3F948FE78F90A" bold="true" box="[151,482,1699,1726]" fontSize="11" level="1" pageId="5" pageNumber="384" reason="1">
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<taxonomicName id="C1944D4FFFC9FFAD66A3F948FE78F90A" ID-CoL="725Z3" authority="Heckel, 1843" authorityName="Heckel" authorityYear="1843" box="[151,482,1699,1726]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="species" species="basak">
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<emphasis id="34E0EADEFFC9FFAD66A3F948FE78F90A" bold="true" box="[151,482,1699,1726]" pageId="5" pageNumber="384">
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<emphasis id="34E0EADEFFC9FFAD66A3F948FEA1F909" bold="true" box="[151,315,1699,1725]" italics="true" pageId="5" pageNumber="384">Leucos basak</emphasis>
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Heckel, 1843
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="062B36CCFFC9FFAD66A3F92DFEADF96B" blockId="5.[151,482,1699,1759]" box="[151,311,1734,1759]" pageId="5" pageNumber="384">
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(
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<figureCitation id="9EAF2A49FFC9FFAD66ABF92DFF77F96B" box="[159,237,1734,1759]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="5" pageNumber="384">Figs. 2</figureCitation>
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B, 3B)
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</paragraph>
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</subSubSection>
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<subSubSection id="4E8E6547FFC9FFAE66A3F8FBFA8CF9DC" lastPageId="6" lastPageNumber="385" pageId="5" pageNumber="384" type="reference_group">
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<paragraph id="062B36CCFFC9FFAD66A3F8FBFB38F890" blockId="5.[151,1436,1806,2013]" box="[151,1186,1806,1829]" pageId="5" pageNumber="384">
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<taxonomicName id="C1944D4FFFC9FFAD66A3F8FBFE74F890" authority="Heckel, 1843: 1006" authorityName="Heckel" authorityPageNumber="1006" authorityYear="1843" box="[151,494,1806,1829]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="species" species="basak">
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<emphasis id="34E0EADEFFC9FFAD66A3F8FBFEBAF890" box="[151,288,1806,1829]" italics="true" pageId="5" pageNumber="384">Leucos basak</emphasis>
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Heckel, 1843: 1006
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</taxonomicName>
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(
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<typeStatus id="D92F886EFFC9FFAD67CEF8E5FDB4F890" box="[506,558,1806,1828]" pageId="5" pageNumber="384">Type</typeStatus>
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locality: Vergoraz and Lake Drusino, near Imotschi,
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<collectingCountry id="7E83765CFFC9FFAD627CF8E5FB02F890" box="[1096,1176,1806,1828]" name="Croatia" pageId="5" pageNumber="384">Croatia</collectingCountry>
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).
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</paragraph>
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<paragraph id="062B36CCFFC9FFAD66A3F8C6FB37F80A" blockId="5.[151,1436,1806,2013]" pageId="5" pageNumber="384">
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<treatmentCitationGroup id="268411E2FFC9FFAD66A3F8C6FB37F80A" infered="true" pageId="5" pageNumber="384">
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<taxonomicName id="C1944D4FFFC9FFAD66A3F8C6FD2FF8F7" ID-CoL="4TTH9" authority="Vladikov & Petit, 1930: 391" authorityName="Vladikov & Petit" authorityPageNumber="391" authorityYear="1930" box="[151,693,1837,1859]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="subSpecies" species="aula" subSpecies="karamani">
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<emphasis id="34E0EADEFFC9FFAD66A3F8C6FE1AF8F7" box="[151,384,1837,1859]" italics="true" pageId="5" pageNumber="384">Rutilus aula karamani</emphasis>
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<treatmentCitation id="873510DDFFC9FFAD67BDF8C6FD2FF8F7" author="Vladikov" box="[393,693,1837,1859]" page="391" pageId="5" pageNumber="384" year="1930">
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<bibRefCitation id="62054B3DFFC9FFAD67BDF8C6FD18F8F7" author="Vladikov" box="[393,642,1837,1859]" pageId="5" pageNumber="384" refString="Vladikov, V. D. & Petit, G. (1930) Sur quelques poisons d'eau douce d'Albanie. Bulletin de la Societe Zoologique de France, 55, 383 - 409." type="journal article" year="1930">Vladikov & Petit, 1930</bibRefCitation>
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: 391
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</treatmentCitation>
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</taxonomicName>
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(
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<typeStatus id="D92F886EFFC9FFAD64F2F8C6FD69F8F7" box="[710,755,1837,1859]" pageId="5" pageNumber="384">type</typeStatus>
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locality, Lake Ohrid,
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<collectingCountry id="7E83765CFFC9FFAD65E9F8C6FBA8F8F7" box="[989,1074,1837,1859]" name="Albania" pageId="5" pageNumber="384">Albania</collectingCountry>
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). The authorship of this species is reported as Fowler,
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<quantity id="C16C9B29FFC9FFAD67A2F8A7FE7FF8D6" box="[406,485,1868,1890]" metricMagnitude="1" metricUnit="m" metricValue="5.02158" pageId="5" pageNumber="384" unit="in" value="1977.0">1977 in</quantity>
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<bibRefCitation id="62054B3DFFC9FFAD67D9F8A7FD10F8D6" author="Kottelat" box="[493,650,1868,1890]" pageId="5" pageNumber="384" refString="Kottelat, M. (1997) European freshwater fish. Biologia, 52 (Supplement), 1 - 271." type="journal article" year="1997">Kottelat (1997)</bibRefCitation>
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because Kottelat assumed that the original description was infrasubspecific (
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<taxonomicName id="C1944D4FFFC9FFAD66FBF881FED1F834" ID-CoL="4TTGW" box="[207,331,1898,1920]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="species" species="aula">
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<emphasis id="34E0EADEFFC9FFAD66FBF881FED1F834" box="[207,331,1898,1920]" italics="true" pageId="5" pageNumber="384">Rutilus aula</emphasis>
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</taxonomicName>
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natio
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<taxonomicName id="C1944D4FFFC9FFAD67B8F881FE77F834" ID-CoL="4TTH9" box="[396,493,1898,1920]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="species" species="karamani">
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<emphasis id="34E0EADEFFC9FFAD67B8F881FE77F834" box="[396,493,1898,1920]" italics="true" pageId="5" pageNumber="384">karamani</emphasis>
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</taxonomicName>
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), which should not be made available (ICZN, 1999; art. 45.5). It should be noted, however, that
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<bibRefCitation id="62054B3DFFC9FFAD66C7F862FE70F82B" author="Vladikov" box="[243,490,1929,1951]" pageId="5" pageNumber="384" refString="Vladikov, V. D. & Petit, G. (1930) Sur quelques poisons d'eau douce d'Albanie. Bulletin de la Societe Zoologique de France, 55, 383 - 409." type="journal article" year="1930">Vladikov & Petit (1930)</bibRefCitation>
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in the text of their original description used the name
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<taxonomicName id="C1944D4FFFC9FFAD6225F862FB6EF82B" ID-CoL="4TTH9" box="[1041,1268,1929,1951]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="subSpecies" species="aula" subSpecies="karamani">
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<emphasis id="34E0EADEFFC9FFAD6225F862FB6EF82B" box="[1041,1268,1929,1951]" italics="true" pageId="5" pageNumber="384">Rutilus aula karamani</emphasis>
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</taxonomicName>
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, without the use of “natio”, making clear their intention to describe a subspecific taxon (ICZN, 1999: art. 45.6.4.1).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="062B36CCFFC9FFAD66A3F823FC7EF868" blockId="5.[151,1436,1806,2013]" box="[151,996,1990,2013]" pageId="5" pageNumber="384">
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<treatmentCitationGroup id="268411E2FFC9FFAD66A3F823FC7EF868" box="[151,996,1990,2013]" pageId="5" pageNumber="384">
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||
<taxonomicName id="C1944D4FFFC9FFAD66A3F823FE88F868" ID-CoL="725YR" box="[151,274,1990,2013]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="384" phylum="Chordata" rank="species" species="aula">
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<emphasis id="34E0EADEFFC9FFAD66A3F823FE88F868" box="[151,274,1990,2013]" italics="true" pageId="5" pageNumber="384">Leucos aula</emphasis>
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</taxonomicName>
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<taxonomicName id="C1944D4FFFC9FFAD672CF82CFDF3F868" authority="Karaman, 1924: 56" authorityName="Karaman" authorityPageNumber="56" authorityYear="1924" box="[280,617,1990,2012]" pageId="5" pageNumber="384" rank="variety" variety="ohridana">
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var.
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<emphasis id="34E0EADEFFC9FFAD6770F82DFE3BF868" box="[324,417,1990,2012]" italics="true" pageId="5" pageNumber="384">ohridana</emphasis>
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<treatmentCitation id="873510DDFFC9FFAD6793F82CFDF3F868" author="Karaman" box="[423,617,1990,2012]" page="56" pageId="5" pageNumber="384" year="1924">
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||
<bibRefCitation id="62054B3DFFC9FFAD6793F82CFDDCF868" author="Karaman" box="[423,582,1990,2012]" pageId="5" pageNumber="384" refString="Karaman, S. L. (1924) Pisces macedoniae. Hrvatska Stamparija, Split, 90 pp." type="book" year="1924">Karaman, 1924</bibRefCitation>
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||
: 56
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</treatmentCitation>
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||
</taxonomicName>
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||
(
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<typeStatus id="D92F886EFFC9FFAD6443F82DFD3EF868" box="[631,676,1990,2012]" pageId="5" pageNumber="384">type</typeStatus>
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locality, Lake Ohrid, FYROM)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="062B36CCFFCAFFAE66A3FF72FA8CF9DC" blockId="6.[151,1018,152,205]" lastBlockId="6.[151,1436,1364,2037]" pageId="6" pageNumber="385">
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||
<taxonomicName id="C1944D4FFFCAFFAE66A3FF72FE88FF1A" box="[151,274,152,174]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="aula">
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<emphasis id="34E0EADEFFCAFFAE66A3FF72FE88FF1A" box="[151,274,152,174]" italics="true" pageId="6" pageNumber="385">Leucos aula</emphasis>
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||
</taxonomicName>
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||
<taxonomicName id="C1944D4FFFCAFFAE672CFF72FDECFF1A" authority="Karaman, 1924: 57" authorityName="Karaman" authorityPageNumber="57" authorityYear="1924" box="[280,630,152,174]" pageId="6" pageNumber="385" rank="variety" variety="prespensis">
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||
var.
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<emphasis id="34E0EADEFFCAFFAE6770FF72FE34FF1A" box="[324,430,153,174]" italics="true" pageId="6" pageNumber="385">prespensis</emphasis>
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||
<bibRefCitation id="62054B3DFFCAFFAE6780FF72FDC9FF1A" author="Karaman" box="[436,595,152,174]" pageId="6" pageNumber="385" refString="Karaman, S. L. (1924) Pisces macedoniae. Hrvatska Stamparija, Split, 90 pp." type="book" year="1924">Karaman, 1924</bibRefCitation>
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||
: 57
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||
</taxonomicName>
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||
(
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||
<typeStatus id="D92F886EFFCAFFAE64B0FF73FD2BFF1A" box="[644,689,152,174]" pageId="6" pageNumber="385">type</typeStatus>
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||
locality, Lake Prespa, FYROM)
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<taxonomicName id="C1944D4FFFCAFFAE66A3FF5DFDC8FF78" ID-CoL="5KYNJ" authority="Maric, 1989: 65" authorityName="Maric" authorityPageNumber="65" authorityYear="1989" box="[151,594,182,205]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="subSpecies" species="prespensis" subSpecies="vukovici">
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||
<emphasis id="34E0EADEFFCAFFAE66A3FF5DFE33FF78" box="[151,425,182,205]" italics="true" pageId="6" pageNumber="385">Rutilus prespensis vukovici</emphasis>
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||
Marić, 1989: 65
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</taxonomicName>
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||
(
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<typeStatus id="D92F886EFFCAFFAE646BFF5DFD16FF78" box="[607,652,182,204]" pageId="6" pageNumber="385">type</typeStatus>
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||
locality, Lake Skadar,
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<collectingCountry id="7E83765CFFCAFFAE6540FF5CFC6BFF78" box="[884,1009,183,204]" name="Serbia" pageId="6" pageNumber="385">Montenegro</collectingCountry>
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||
)
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<emphasis id="34E0EADEFFCAFFAE66A3FABFFE14FAD9" bold="true" box="[151,398,1364,1389]" pageId="6" pageNumber="385">Examined materials.</emphasis>
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||
IZA 83120, 33,
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<collectingCountry id="7E83765CFFCAFFAE6473FABFFD06FAD9" box="[583,668,1364,1389]" name="Croatia" pageId="6" pageNumber="385">Croatia</collectingCountry>
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, R. Krupa (R. Neretva basin), P.G. Bianco,
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<date id="722A100CFFCAFFAE624AFABFFA88FAD8" box="[1150,1298,1364,1388]" pageId="6" pageNumber="385" value="1983-05-28">28 May 1983</date>
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.—IZA 0 48, 5,
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<collectingCountry id="7E83765CFFCAFFAE6680FA9CFE90FA24" box="[180,266,1399,1424]" name="Croatia" pageId="6" pageNumber="385">Croatia</collectingCountry>
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, L. Bačinska (R. Neretva basin), M. Mrakovic,
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<date id="722A100CFFCAFFAE651CFA93FC48FA24" box="[808,978,1399,1424]" pageId="6" pageNumber="385" value="1997-04-10">10 Aprile 1997</date>
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.—IZA 0 0 164, 5, FYROM, L. Ohrid, R. Gruptché,
|
||
<date id="722A100CFFCAFFAE6739FA77FE30FA00" box="[269,426,1436,1461]" pageId="6" pageNumber="385" value="1970-04-10">10 April 1970</date>
|
||
.—IZA 0 429, 30 (out of 122), FYROM; L. Ohrid, P.G. Bianco,
|
||
<date id="722A100CFFCAFFAE625AFA77FAD3FA00" box="[1134,1353,1436,1460]" pageId="6" pageNumber="385" value="1987-08-28" valueMax="1987-08-29" valueMin="1987-08-28">28–29 August 1987</date>
|
||
.—IZA 0 430, 10, FYROM, L. Ohrid, R. Gruptché,
|
||
<date id="722A100CFFCAFFAE64B6FA2BFCB4FA6D" box="[642,814,1472,1497]" pageId="6" pageNumber="385" value="1969-03-13">13 March 1969</date>
|
||
.- IZA 85481, 30 (out of 232),
|
||
<collectingCountry id="7E83765CFFCAFFAE62A5FA2AFA87FA6D" box="[1169,1309,1473,1497]" name="Serbia" pageId="6" pageNumber="385">Montenegro</collectingCountry>
|
||
, L. Skadar, P.G. Bianco and B. Knezevic,
|
||
<date id="722A100CFFCAFFAE67D1FA0FFD04FA48" box="[485,670,1508,1533]" pageId="6" pageNumber="385" value="1984-07-24" valueMax="1984-07-25" valueMin="1984-07-24">24–25 July 1984</date>
|
||
.—IZA 0 441, 4,
|
||
<collectingCountry id="7E83765CFFCAFFAE6563FA0EFC79FA49" box="[855,995,1509,1533]" name="Serbia" pageId="6" pageNumber="385">Montenegro</collectingCountry>
|
||
, L. Skadar, P.G. Bianco and D. Marić,
|
||
<date id="722A100CFFCAFFAE63BAFA0FFED1F994" pageId="6" pageNumber="385" value="1999-11-09">9 November 1999</date>
|
||
.—IZA 0 0 55, 5, FYROM: L. Prespa, P. Banarescu,
|
||
<date id="722A100CFFCAFFAE65A4F9E3FBFDF995" box="[912,1127,1544,1569]" pageId="6" pageNumber="385" value="1975-11-29">29 November 1975</date>
|
||
.—IZA 0 435, 10, FYROM, L. Prespa, R. Gruptché,
|
||
<date id="722A100CFFCAFFAE6787F9C7FDD7F9F1" box="[435,589,1580,1605]" pageId="6" pageNumber="385" value="1968-05-15">15 May 1968</date>
|
||
.—IZA 0 422, 12,
|
||
<collectingCountry id="7E83765CFFCAFFAE6515F9C7FCE9F9F0" box="[801,883,1580,1604]" name="Greece" pageId="6" pageNumber="385">Greece</collectingCountry>
|
||
, lakes Prespa and Micraprespa, P.G. Bianco,
|
||
<date id="722A100CFFCAFFAE63B4F9C7FEB0F9DC" pageId="6" pageNumber="385" value="1998-08-22">22 August 1998</date>
|
||
. – MNHN 1977.281, 3,
|
||
<collectingCountry id="7E83765CFFCAFFAE640DF9BBFD08F9DD" box="[569,658,1616,1641]" name="Albania" pageId="6" pageNumber="385">Albania</collectingCountry>
|
||
(
|
||
<typeStatus id="D92F886EFFCAFFAE6495F9BBFC9FF9DC" box="[673,773,1616,1640]" pageId="6" pageNumber="385" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="C1944D4FFFCAFFAE651DF9BBFA90F9DC" authority="Vladikov & Petit, 1930" authorityName="Vladikov & Petit" authorityYear="1930" box="[809,1290,1616,1641]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="karamanni">
|
||
<emphasis id="34E0EADEFFCAFFAE651DF9BBFC63F9DD" box="[809,1017,1616,1641]" italics="true" pageId="6" pageNumber="385">Rutilus karamanni</emphasis>
|
||
<bibRefCitation id="62054B3DFFCAFFAE6234F9BBFA90F9DC" author="Vladikov" box="[1024,1290,1616,1641]" pageId="6" pageNumber="385" refString="Vladikov, V. D. & Petit, G. (1930) Sur quelques poisons d'eau douce d'Albanie. Bulletin de la Societe Zoologique de France, 55, 383 - 409." type="journal article" year="1930">Vladikov & Petit, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="52EB6644FFCAFFAE66A3FB32FD38FA98" httpUri="https://zenodo.org/record/225781/files/figure.png" pageId="6" pageNumber="385" targetBox="[151,1436,257,1219]" targetPageId="6">
|
||
<paragraph id="062B36CCFFCAFFAE66A3FB32FD38FA98" blockId="6.[151,1436,1240,1324]" pageId="6" pageNumber="385">
|
||
<emphasis id="34E0EADEFFCAFFAE66A3FB32FE94FB5A" bold="true" box="[151,270,1241,1263]" pageId="6" pageNumber="385">FIGURE 3</emphasis>
|
||
.
|
||
<emphasis id="34E0EADEFFCAFFAE6728FB33FEACFB5B" bold="true" box="[284,310,1240,1263]" pageId="6" pageNumber="385">A)</emphasis>
|
||
Lectotype of
|
||
<taxonomicName id="C1944D4FFFCAFFAE67FCFB31FDDEFB5B" box="[456,580,1241,1263]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="aula">
|
||
<emphasis id="34E0EADEFFCAFFAE67FCFB31FDDEFB5B" box="[456,580,1241,1263]" italics="true" pageId="6" pageNumber="385">Leucos aula</emphasis>
|
||
</taxonomicName>
|
||
, 74 mm SL (ANSP 6434), Padua Province, northern Italy;
|
||
<emphasis id="34E0EADEFFCAFFAE629DFB33FB58FB5B" bold="true" box="[1193,1218,1240,1263]" pageId="6" pageNumber="385">B)</emphasis>
|
||
Lectotype of
|
||
<emphasis id="34E0EADEFFCAFFAE6360FB31FF43FABA" italics="true" pageId="6" pageNumber="385">
|
||
<taxonomicName id="C1944D4FFFCAFFAE6360FB31FF4FFABA" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="basak">Leucos basak</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
117 mm SL (NMW 50723-1), Vergoraz and Lake Drusino near Imotsky, Croatia;
|
||
<emphasis id="34E0EADEFFCAFFAE6229FB1CFBA2FABA" bold="true" box="[1053,1080,1271,1294]" pageId="6" pageNumber="385">C)</emphasis>
|
||
Lectotype of
|
||
<emphasis id="34E0EADEFFCAFFAE62F2FB13FA06FABA" box="[1222,1436,1272,1294]" italics="true" pageId="6" pageNumber="385">
|
||
<taxonomicName id="C1944D4FFFCAFFAE62F2FB13FA02FABA" box="[1222,1432,1272,1294]" class="Actinopterygii" family="Cyprinidae" genus="Sarmarutilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="rubilio">Sarmarutilus rubilio</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
93 mm SL (ANSP 6509), Lake Nemi, Central Italy.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4E8E6547FFCAFFAE66F3F99FFC6DF818" pageId="6" pageNumber="385" type="diagnosis">
|
||
<paragraph id="062B36CCFFCAFFAE66F3F99FFC6DF818" blockId="6.[151,1436,1364,2037]" pageId="6" pageNumber="385">
|
||
<emphasis id="34E0EADEFFCAFFAE66F3F99FFEC4F939" bold="true" box="[199,350,1652,1677]" pageId="6" pageNumber="385">Diagnosis. A</emphasis>
|
||
species of
|
||
<taxonomicName id="C1944D4FFFCAFFAE67D4F99EFDB5F938" box="[480,559,1653,1676]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFCAFFAE67D4F99EFDB5F938" box="[480,559,1653,1676]" italics="true" pageId="6" pageNumber="385">Leucos</emphasis>
|
||
</taxonomicName>
|
||
characterized by moderate size, usually
|
||
<quantity id="C16C9B29FFCAFFAE6234F99FFB0DF938" box="[1024,1175,1652,1676]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="6" pageNumber="385" unit="mm" value="130.0" valueMax="140.0" valueMin="120.0">120–140 mm</quantity>
|
||
SL; absence of a midlateral band; body uniformly silvery in life; peritoneal membrane black.; usually 36–38 pored scales on LL, as opposed to
|
||
<quantity id="C16C9B29FFCAFFAE672DF957FEE7F961" box="[281,381,1724,1749]" metricMagnitude="0" metricUnit="m" metricValue="1.0287" metricValueMax="1.0922" metricValueMin="0.9652" pageId="6" pageNumber="385" unit="in" value="40.5" valueMax="43.0" valueMin="38.0">38–43 in</quantity>
|
||
the other
|
||
<taxonomicName id="C1944D4FFFCAFFAE67DBF956FDA4F960" box="[495,574,1725,1748]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFCAFFAE67DBF956FDA4F960" box="[495,574,1725,1748]" italics="true" pageId="6" pageNumber="385">Leucos</emphasis>
|
||
</taxonomicName>
|
||
species. It may be distinguished from
|
||
<taxonomicName id="C1944D4FFFCAFFAE65C0F956FBE7F961" box="[1012,1149,1724,1749]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="aula">
|
||
<emphasis id="34E0EADEFFCAFFAE65C0F956FBE7F961" box="[1012,1149,1724,1749]" italics="true" pageId="6" pageNumber="385">Leucos aula</emphasis>
|
||
</taxonomicName>
|
||
for the absence of lateral band and the color of the peritoneal membrane, black in
|
||
<taxonomicName id="C1944D4FFFCAFFAE652EF90AFCE0F94D" box="[794,890,1760,1785]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCAFFAE652EF90AFCE0F94D" box="[794,890,1760,1785]" italics="true" pageId="6" pageNumber="385">L. basak</emphasis>
|
||
</taxonomicName>
|
||
, and silvery in
|
||
<taxonomicName id="C1944D4FFFCAFFAE621FF90AFBE6F94D" box="[1067,1148,1760,1785]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="aula">
|
||
<emphasis id="34E0EADEFFCAFFAE621FF90AFBE6F94D" box="[1067,1148,1760,1785]" italics="true" pageId="6" pageNumber="385">L. aula</emphasis>
|
||
</taxonomicName>
|
||
. It differs from
|
||
<taxonomicName id="C1944D4FFFCAFFAE6307F90AFA06F94C" box="[1331,1436,1761,1784]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="panosi">
|
||
<emphasis id="34E0EADEFFCAFFAE6307F90AFA06F94C" box="[1331,1436,1761,1784]" italics="true" pageId="6" pageNumber="385">L. panosi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C1944D4FFFCAFFAE66FCF8EEFECBF8A9" box="[200,337,1796,1821]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFCAFFAE66FCF8EEFECBF8A9" box="[200,337,1796,1821]" italics="true" pageId="6" pageNumber="385">L. ylikiensis</emphasis>
|
||
</taxonomicName>
|
||
mainly by the number of GR, which are usually
|
||
<quantity id="C16C9B29FFCAFFAE65B3F8EFFC44F8A9" box="[903,990,1796,1821]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="6" pageNumber="385" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE65D3F8EEFBD3F8A9" box="[999,1097,1796,1821]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCAFFAE65D3F8EEFBD3F8A9" box="[999,1097,1796,1821]" italics="true" pageId="6" pageNumber="385">L. basak</emphasis>
|
||
</taxonomicName>
|
||
and respectively,
|
||
<quantity id="C16C9B29FFCAFFAE632CF8EFFAE7F8A9" box="[1304,1405,1796,1821]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="6" pageNumber="385" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE63B1F8EEFE9AF8F5" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="ylikiensis">
|
||
<emphasis id="34E0EADEFFCAFFAE63B1F8EEFE9AF8F5" italics="true" pageId="6" pageNumber="385">L. ylikiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="C16C9B29FFCAFFAE6701F8C3FE03F8F5" box="[309,409,1832,1857]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="6" pageNumber="385" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">13–14 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE67ABF8C2FD92F8F4" box="[415,520,1833,1856]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="panosi">
|
||
<emphasis id="34E0EADEFFCAFFAE67ABF8C2FD92F8F4" box="[415,520,1833,1856]" italics="true" pageId="6" pageNumber="385">L. panosi</emphasis>
|
||
</taxonomicName>
|
||
. The most closely related species, according to molecular analyses (
|
||
<figureCitation id="9EAF2A49FFCAFFAE6332F8C3FADCF8F4" box="[1286,1350,1832,1857]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1610,1632]" captionTargetBox="[182,1420,597,1530]" captionTargetId="figure@2.[151,1436,591,1588]" captionTargetPageId="2" captionText="FIGURE 1. A) Collecting localities (L. = lakes, R. = rivers) of Rutilus complex species. 1, R. Thames; 2, R. Danube (Slovakia); 3, R. Danube (Romania); 4, L. Como; 5, L. Maggiore; 6, R. Arno and R. Serchio; 7, R. Albegna and R. Cecina; 8, R. Ombrone and R. Fiora; 9, R. Tiber and R. Mignone; 10, L. Trasimeno; 11, R. Liri and R. Garigliano; 12, R. Volturno and R. Sele; 13, R. Pescara and R. Tavo; 14, R. Trigno; 15, R. Tronto; 16, R. Livenza; 17, L. Pernica; 18, R. Sava; 19, L. Drusino; 20, R. Krupa (R. Neretva basin); 21, L. Besanka (R. Neretva basin); 22, R. Moraca and R. Zeta; 23, L. Skadar; 24, L. Dojran; 25, L. Horid; 26, L. Prespa and L. Micraprespa; 27, L. Vegoritis; 28, L. Volvi; 29, L. Joannina; 30, L. Yliki and L. Paralimni; 31, L. Ambrakia; 32, L. Trichonis. Bold numbers are the different peri-Mediterranean ichthyogeographic districts described in Bianco (1990). B) A schematic temporal reconstruction of the diversification of the genus in southern Mediterranean Europe based on cyt- b is shown (modified from Ketmaier et al., 2008). For each taxon the corresponding number of the district of origin is shown; triangles indicate that multiple populations were analyzed. Numbers at nodes are divergence times in million years; all labeled nodes were robustly supported in the phylogenetic reconstructions (see Ketmaier et al., 2008 for further details)." httpUri="https://zenodo.org/record/225779/files/figure.png" pageId="6" pageNumber="385">Fig. 1</figureCitation>
|
||
B) is
|
||
<taxonomicName id="C1944D4FFFCAFFAE63B1F8C2FF4EF8D1" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="albus">
|
||
<emphasis id="34E0EADEFFCAFFAE63B1F8C2FF4EF8D1" italics="true" pageId="6" pageNumber="385">L. albus</emphasis>
|
||
</taxonomicName>
|
||
. The two species can be identified on the basis of the number of LL scales, usually
|
||
<quantity id="C16C9B29FFCAFFAE6295F8A7FA9CF8D1" box="[1185,1286,1868,1893]" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.0668" metricValueMin="1.0413999999999999" pageId="6" pageNumber="385" unit="in" value="41.5" valueMax="42.0" valueMin="41.0">41–42 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE633AF8A6FAF1F8D1" box="[1294,1387,1868,1893]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="albus">
|
||
<emphasis id="34E0EADEFFCAFFAE633AF8A6FAF1F8D1" box="[1294,1387,1868,1893]" italics="true" pageId="6" pageNumber="385">L. albus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="C16C9B29FFCAFFAE66A3F89BFF61F83D" box="[151,251,1904,1929]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.652" metricValueMin="9.144" pageId="6" pageNumber="385" unit="in" value="37.0" valueMax="38.0" valueMin="36.0">36–38 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE6736F89AFEFBF83D" box="[258,353,1904,1929]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCAFFAE6736F89AFEFBF83D" box="[258,353,1904,1929]" italics="true" pageId="6" pageNumber="385">L. basak</emphasis>
|
||
</taxonomicName>
|
||
, the number of D branched rays,
|
||
<quantity id="C16C9B29FFCAFFAE64E9F89BFC93F83D" box="[733,777,1904,1929]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="6" pageNumber="385" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE6524F89AFCEAF83D" box="[784,880,1904,1929]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCAFFAE6524F89AFCEAF83D" box="[784,880,1904,1929]" italics="true" pageId="6" pageNumber="385">L. basak</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="C16C9B29FFCAFFAE6593F89BFC49F83D" box="[935,979,1904,1929]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="385" unit="in" value="8.0">8 in</quantity>
|
||
<taxonomicName id="C1944D4FFFCAFFAE65EEF89AFBAEF83D" box="[986,1076,1904,1929]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="albus">
|
||
<emphasis id="34E0EADEFFCAFFAE65EEF89AFBAEF83D" box="[986,1076,1904,1929]" italics="true" pageId="6" pageNumber="385">L. albus</emphasis>
|
||
</taxonomicName>
|
||
, and apparently for the color of the peritoneal membrane, nearly silvery in
|
||
<taxonomicName id="C1944D4FFFCAFFAE644CF87EFD48F819" box="[632,722,1940,1965]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="albus">
|
||
<emphasis id="34E0EADEFFCAFFAE644CF87EFD48F819" box="[632,722,1940,1965]" italics="true" pageId="6" pageNumber="385">L. albus</emphasis>
|
||
</taxonomicName>
|
||
, and blackish in
|
||
<taxonomicName id="C1944D4FFFCAFFAE65A5F87EFC75F819" box="[913,1007,1940,1965]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="385" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCAFFAE65A5F87EFC75F819" box="[913,1007,1940,1965]" italics="true" pageId="6" pageNumber="385">L. basak</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFCAFFAF66F3F853FAD9FD88" lastPageId="7" lastPageNumber="386" pageId="6" pageNumber="385" type="description">
|
||
<paragraph id="062B36CCFFCAFFAF66F3F853FAD9FD88" blockId="6.[151,1436,1364,2037]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="386" pageId="6" pageNumber="385">
|
||
<emphasis id="34E0EADEFFCAFFAE66F3F853FEC3F865" bold="true" box="[199,345,1976,2001]" pageId="6" pageNumber="385">Description.</emphasis>
|
||
A small-medium sized species. In
|
||
<collectingCountry id="7E83765CFFCAFFAE64CEF853FCD7F865" box="[762,845,1976,2001]" name="Croatia" pageId="6" pageNumber="385">Croatia</collectingCountry>
|
||
may reach
|
||
<quantity id="C16C9B29FFCAFFAE65E8F853FBA7F864" box="[988,1085,1976,2000]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="385" unit="mm" value="220.0">220 mm</quantity>
|
||
TL and
|
||
<quantity id="C16C9B29FFCAFFAE629DF853FB76F865" box="[1193,1260,1976,2001]" metricMagnitude="-1" metricUnit="kg" metricValue="1.8" pageId="6" pageNumber="385" unit="g" value="180.0">180 g</quantity>
|
||
of weight, but usually less than
|
||
<quantity id="C16C9B29FFCAFFAE676CF837FE2FF840" box="[344,437,2012,2037]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="385" unit="mm" value="150.0">150 mm</quantity>
|
||
TL (
|
||
<bibRefCitation id="62054B3DFFCAFFAE67D9F837FD6BF841" author="Macrovcic" box="[493,753,2012,2037]" pageId="6" pageNumber="385" refString="Macrovcic, M., Brigic, A., Buj, I., Caleta, M., Mustafic, P. & Zanella, D. (2006) Red book of freshwater fishes of Croatia. Ministry of Culture, State Institute for Nature Protection, Zagreb, 253 pp." type="book" year="2006">
|
||
Mačrovcić
|
||
<emphasis id="34E0EADEFFCAFFAE645AF836FD37F841" box="[622,685,2012,2037]" italics="true" pageId="6" pageNumber="385">et al.,</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
). Body uniformly silvery without longitudinal band; color of the eye in living animals is yellowish; fins yellowish or pale grayish in preserved specimens; snout pointed, preorbital distance near equal to the horizontal diameter of the eye; lips smooth; mouth opening oblique, the corner of maxillae at the same level of the vertical crossing the anterior border of the orbit; mouth terminal or slightly inferior; profile of dorsum convex sometimes humped in large specimens; paired and un-paired fins yellowish with few scattered melanophores, free margin of D and A concave; P1 longer in males, were may reach the origin of P2; caudal fin forked; peritoneal membrane blackened by several melanophores addensed and fused; head length about 3.6–3.8 times in the SL; body deep about 3.5–3.7 times in the SL; origin of the D at same level of the insertion of the P2; LL complete, and extending from the margin of opercular membrane, to the end of caudal peduncle, with 35–41 pored scales, usually 36–38; 7.5 above and 3.5 below the LL; constantly 3 un-branched rays followed by 9 branched rays in the D; constantly 3 un-branched and usually 8 branched rays in the anal fin; P1 with 1 unbranched and 14–15 branched rays; P2 with 1 un-branched ray and constantly 8 branched rays; usually 9–10 total GR; in adult males pearl organ absent. For additional description and shape, see
|
||
<figureCitation id="9EAF2A49FFCBFFAF6228FDCFFBC7FD88" box="[1052,1117,548,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="7" pageNumber="386">Fig. 2</figureCitation>
|
||
B and Tables 1, 2, 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFCBFFAF66F3FDACFDA7FD79" pageId="7" pageNumber="386" type="distribution">
|
||
<paragraph id="062B36CCFFCBFFAF66F3FDACFDA7FD79" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="386">
|
||
<emphasis id="34E0EADEFFCBFFAF66F3FDACFEFAFDD4" bold="true" box="[199,352,583,608]" pageId="7" pageNumber="386">Distribution.</emphasis>
|
||
The species distribution includes lakes and mostly still water of rivers of the Adriatic drainages, from
|
||
<collectingCountry id="7E83765CFFCBFFAF66EDFD87FEB5FD31" box="[217,303,620,645]" name="Croatia" pageId="7" pageNumber="386">Croatia</collectingCountry>
|
||
, till to Albany, and
|
||
<collectingCountry id="7E83765CFFCBFFAF6414FD86FD37FD31" box="[544,685,621,645]" name="Serbia" pageId="7" pageNumber="386">Montenegro</collectingCountry>
|
||
. In Lake Skadar the species is sympatric with
|
||
<emphasis id="34E0EADEFFCBFFAF62EFFD86FAE0FD31" box="[1243,1402,620,645]" italics="true" pageId="7" pageNumber="386">
|
||
<taxonomicName id="C1944D4FFFCBFFAF62EFFD86FAEFFD31" box="[1243,1397,620,645]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="albus">Leucos albus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In
|
||
<collectingCountry id="7E83765CFFCBFFAF66A3FD64FF70FD1C" box="[151,234,655,680]" name="Croatia" pageId="7" pageNumber="386">Croatia</collectingCountry>
|
||
its distribution is limited to basins of south-eastern part: lakes Crven, Modro, Bačinska and rivers Matica and Neretva (Mracovčić
|
||
<emphasis id="34E0EADEFFCBFFAF679AFD5EFE7CFD79" box="[430,486,692,717]" italics="true" pageId="7" pageNumber="386">et al.</emphasis>
|
||
, 2006).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFCBFFAF66F3FD3CFDDCFBEC" pageId="7" pageNumber="386" type="discussion">
|
||
<paragraph id="062B36CCFFCBFFAF66F3FD3CFDDCFBEC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="386">
|
||
<emphasis id="34E0EADEFFCBFFAF66F3FD3CFE7FFD44" bold="true" box="[199,485,727,752]" pageId="7" pageNumber="386">Remarks on synonyms.</emphasis>
|
||
According to mtDNA (
|
||
<bibRefCitation id="62054B3DFFCBFFAF64CAFD3CFC61FD44" author="Ketmaier" box="[766,1019,727,752]" pageId="7" pageNumber="386" refString="Ketmaier, V., Bianco, P. G. & Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article" year="2008">
|
||
Ketmaier
|
||
<emphasis id="34E0EADEFFCBFFAF6547FD32FC2CFD44" box="[883,950,727,752]" italics="true" pageId="7" pageNumber="386">et al.,</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
) and to the morphological analyses presented here on
|
||
<taxonomicName id="C1944D4FFFCBFFAF6753FD16FD9BFCA1" box="[359,513,764,789]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCBFFAF6753FD16FD9BFCA1" box="[359,513,764,789]" italics="true" pageId="7" pageNumber="386">Leucos basak</emphasis>
|
||
</taxonomicName>
|
||
(Table 3) from the Dalmatian, Albanian, and Aegean ichthyogeographic districts (
|
||
<emphasis id="34E0EADEFFCBFFAF6694FCCAFF44FC8C" box="[160,222,801,824]" italics="true" pageId="7" pageNumber="386">sensu</emphasis>
|
||
<bibRefCitation id="62054B3DFFCBFFAF66DFFCF4FE19FC8C" author="Bianco" box="[235,387,799,824]" pageId="7" pageNumber="386" refString="Bianco, P. G. (1990) Potential role of the palaeohistory of the Mediterranean and Parathetys basin on the early dispersal of Europe-Mediterranean freshwater fishes. Ichthyological Exploration of Freshwaters, 1, 167 - 184." type="journal article" year="1990">Bianco 1990</bibRefCitation>
|
||
), the populations from lakes Ohrid, Prespa and Skadar and River Krupa (River Neretva drainage) cannot be unequivocally distinguished from one another and should be regarded as a single species. This would be partially at odds with the results of
|
||
<bibRefCitation id="62054B3DFFCBFFAF64BAFC8CFC15FC34" box="[654,911,871,896]" pageId="7" pageNumber="386" refString="Milosevic, D., Winkler, K. A., Maric, D. & Weiss, S. (2011) Genotypic and phenotypic evaluation of Rutilus spp. from Skadar, Ohrid and Prespa Lakes supports revision of endemic as well as taxonomic status of several taxa. Journal of Fish Biology, 79, 1094 - 1100. http: // dx. doi. org / 10.1111 / j. 1095 - 8649.2011.03090. x" type="journal article">
|
||
Milošević
|
||
<emphasis id="34E0EADEFFCBFFAF6530FC82FCA6FC34" box="[772,828,871,896]" italics="true" pageId="7" pageNumber="386">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
. These authors, based on mitochondrial (cyt-
|
||
<emphasis id="34E0EADEFFCBFFAF63B0FC8CFA08FC34" box="[1412,1426,871,896]" italics="true" pageId="7" pageNumber="386">b</emphasis>
|
||
) and nuclear (five polymorphic microsatellites) markers coupled with morphology proposed the existence of three species in the area (
|
||
<taxonomicName id="C1944D4FFFCBFFAF6742FC44FDA4FC7C" box="[374,574,943,968]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="ohridanus">
|
||
<emphasis id="34E0EADEFFCBFFAF6742FC44FDA4FC7C" box="[374,574,943,968]" italics="true" pageId="7" pageNumber="386">Rutilus ohridanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C1944D4FFFCBFFAF6478FC5AFD7BFC7C" box="[588,737,945,968]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="prespensis">
|
||
<emphasis id="34E0EADEFFCBFFAF6478FC5AFD7BFC7C" box="[588,737,945,968]" italics="true" pageId="7" pageNumber="386">R. prespensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C1944D4FFFCBFFAF652DFC5AFCEFFC7C" box="[793,885,943,968]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="albus">
|
||
<emphasis id="34E0EADEFFCBFFAF652DFC5AFCEFFC7C" box="[793,885,943,968]" italics="true" pageId="7" pageNumber="386">R. albus</emphasis>
|
||
</taxonomicName>
|
||
). It should be noted, however, that divergence at the mtDNA level was shallow (up to a maximum of five substitutions) and that the lack of gene flow at the nuclear loci could reflect the geographical isolation of these lakes rather than a specific status of the taxa. The authors themselves claimed that “
|
||
<taxonomicName id="C1944D4FFFCBFFAF6788FBF6FDCBFB80" box="[444,593,1053,1076]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="prespensis">
|
||
<emphasis id="34E0EADEFFCBFFAF6788FBF6FDCBFB80" box="[444,593,1053,1076]" italics="true" pageId="7" pageNumber="386">R. prespensis</emphasis>
|
||
</taxonomicName>
|
||
should be further evaluated to determine if the taxon is synonymous with other
|
||
<taxonomicName id="C1944D4FFFCBFFAF66E3FBD4FEBCFBEC" box="[215,294,1087,1112]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFCBFFAF66E3FBD4FEBCFBEC" box="[215,294,1087,1112]" italics="true" pageId="7" pageNumber="386">Rutilus</emphasis>
|
||
</taxonomicName>
|
||
from the Adriatic basin”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFCBFFA066F3FB8FFE76FEA8" lastPageId="8" lastPageNumber="387" pageId="7" pageNumber="386" type="reference_group">
|
||
<paragraph id="062B36CCFFCBFFAF66F3FB8FFD42F954" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="386">
|
||
<taxonomicName id="C1944D4FFFCBFFAF66F3FB8FFE0BFBC9" ID-CoL="4TTHJ" box="[199,401,1124,1149]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="ohridanus">
|
||
<emphasis id="34E0EADEFFCBFFAF66F3FB8FFE0BFBC9" box="[199,401,1124,1149]" italics="true" pageId="7" pageNumber="386">Rutilus ohridanus</emphasis>
|
||
</taxonomicName>
|
||
is a problematic species as the diagnosis of meristic characters given in Karaman’s (1924) original description (D with 10–11 branched rays, A with 11 branched rays and 44 LL scales) overlaps with that given for
|
||
<taxonomicName id="C1944D4FFFCBFFAF6730FB46FE3DFB70" ID-CoL="4TTJ3" box="[260,423,1197,1220]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="subSpecies" species="pygus" subSpecies="virgo">
|
||
<emphasis id="34E0EADEFFCBFFAF6730FB46FE3DFB70" box="[260,423,1197,1220]" italics="true" pageId="7" pageNumber="386">R. pygus virgo</emphasis>
|
||
</taxonomicName>
|
||
for Lake Dojran (FYROM,
|
||
<collectingCountry id="7E83765CFFCBFFAF64D3FB47FCA2FB70" box="[743,824,1196,1220]" name="Greece" pageId="7" pageNumber="386">Greece</collectingCountry>
|
||
) in the very same paper. The overlapping in biometric characters with
|
||
<taxonomicName id="C1944D4FFFCBFFAF677AFB3AFE6CFB5C" ID-CoL="4TTJ3" box="[334,502,1233,1256]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="subSpecies" species="pygus" subSpecies="virgo">
|
||
<emphasis id="34E0EADEFFCBFFAF677AFB3AFE6CFB5C" box="[334,502,1233,1256]" italics="true" pageId="7" pageNumber="386">R. pygus virgo</emphasis>
|
||
</taxonomicName>
|
||
is very likely due to a printing error; also the maximum size reported
|
||
<bibRefCitation id="62054B3DFFCBFFAF6301FB3AFF79FAB8" author="Karaman" pageId="7" pageNumber="386" refString="Karaman, S. L. (1924) Pisces macedoniae. Hrvatska Stamparija, Split, 90 pp." type="book" year="1924">Karaman (1924)</bibRefCitation>
|
||
, (about
|
||
<quantity id="C16C9B29FFCBFFAF6709FB1FFE03FAB8" box="[317,409,1268,1293]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="386" unit="mm" value="150.0">150 mm</quantity>
|
||
TL) would classify the species as a small sized
|
||
<taxonomicName id="C1944D4FFFCBFFAF659AFB1FFC67FAB9" ID-CoL="643L5" box="[942,1021,1268,1293]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFCBFFAF659AFB1FFC67FAB9" box="[942,1021,1268,1293]" italics="true" pageId="7" pageNumber="386">Rutilus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="62054B3DFFCBFFAF6238FB1FFB22FAB8" author="Ivanovic" box="[1036,1208,1268,1293]" pageId="7" pageNumber="386" refString="Ivanovic, B. M. (1973) Ichthyofauna of Skadar Lake. Montenegro, Titograd, 146 pp." type="book" year="1973">Ivanović, 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="62054B3DFFCBFFAF62F7FB1FFADDFAB8" author="Soric" box="[1219,1351,1268,1293]" pageId="7" pageNumber="386" refString="Soric, V. M. (1983) Rutilus rubilio (Cyprinidae, Pisces) U Ohrid-Drim-Skadar sistemu. Acta Biologica Jugoslavica, Biosistematika, 9, 61 - 70." type="journal article" year="1983">Soric, 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="62054B3DFFCBFFAF6366FB1FFF48FA84" author="Maric" pageId="7" pageNumber="386" refString="Maric, D. (1988) The species revision of genus Rutilus Raf. from western part of Balkan peninsula. Prirode Prirodnjackog Museja Titograd, 21, 55 - 79." type="journal article" year="1988">Marić, 1988</bibRefCitation>
|
||
), (see
|
||
<tableCitation id="4B160377FFCBFFAF672AFAFCFEFBFA84" box="[286,353,1303,1328]" captionStart="TABLE 2" captionStartId="10.[119,200,619,640]" captionText="TABLE 2. Meristic counts and several general features in five species of Leucos. CV: most common values; N = sample size; L. = Lake; R. = River." pageId="7" pageNumber="386">Tab 2</tableCitation>
|
||
). In our extensive survey of samples of
|
||
<taxonomicName id="C1944D4FFFCBFFAF651AFAF2FC0BFA84" box="[814,913,1303,1328]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCBFFAF651AFAF2FC0BFA84" box="[814,913,1303,1328]" italics="true" pageId="7" pageNumber="386">R. basak</emphasis>
|
||
</taxonomicName>
|
||
(see above) we found neither 10/11 branched rays in D nor
|
||
<quantity id="C16C9B29FFCBFFAF6701FAD7FEEAFAE1" box="[309,368,1340,1365]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="386" unit="in" value="10.0">10 in</quantity>
|
||
A.
|
||
<taxonomicName id="C1944D4FFFCBFFAF67AFFAD6FDB4FAE1" ID-CoL="4TTHJ" box="[411,558,1340,1365]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="ohridanus">
|
||
<emphasis id="34E0EADEFFCBFFAF67AFFAD6FDB4FAE1" box="[411,558,1340,1365]" italics="true" pageId="7" pageNumber="386">R. ohridanus</emphasis>
|
||
</taxonomicName>
|
||
should be then placed as synonym for
|
||
<taxonomicName id="C1944D4FFFCBFFAF65D8FAD6FBD7FAE1" box="[1004,1101,1340,1365]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCBFFAF65D8FAD6FBD7FAE1" box="[1004,1101,1340,1365]" italics="true" pageId="7" pageNumber="386">R. basak</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="C1944D4FFFCBFFAF6268FAD6FB68FAE0" ID-CoL="4TTHQ" box="[1116,1266,1341,1364]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="prespensis">
|
||
<emphasis id="34E0EADEFFCBFFAF6268FAD6FB68FAE0" box="[1116,1266,1341,1364]" italics="true" pageId="7" pageNumber="386">R. prespensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C1944D4FFFCBFFAF6335FAD6FA06FAE1" ID-CoL="5KYNJ" box="[1281,1436,1340,1365]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="subSpecies" species="prespensis" subSpecies="vukovici">
|
||
<emphasis id="34E0EADEFFCBFFAF6335FAD6FA06FAE1" box="[1281,1436,1340,1365]" italics="true" pageId="7" pageNumber="386">R. p. vukovici</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="34E0EADEFFCBFFAF66F8FA8AFEEAFACC" box="[204,368,1375,1400]" italics="true" pageId="7" pageNumber="386">
|
||
<taxonomicName id="C1944D4FFFCBFFAF66F8FA8AFEF6FACC" box="[204,364,1375,1400]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="karamanni">R. karamanni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
cannot be separated from
|
||
<taxonomicName id="C1944D4FFFCBFFAF6487FA8AFC83FACC" box="[691,793,1375,1400]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCBFFAF6487FA8AFC83FACC" box="[691,793,1375,1400]" italics="true" pageId="7" pageNumber="386">R. basak</emphasis>
|
||
</taxonomicName>
|
||
neither molecularly nor morphologically and are here regarded as junior synonyms for
|
||
<taxonomicName id="C1944D4FFFCBFFAF6424FA6EFDEBFA29" box="[528,625,1412,1437]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="species" species="basak">
|
||
<emphasis id="34E0EADEFFCBFFAF6424FA6EFDEBFA29" box="[528,625,1412,1437]" italics="true" pageId="7" pageNumber="386">R. basak</emphasis>
|
||
</taxonomicName>
|
||
. Thus all the populations of the synonymized nominal taxa represent a single species; this would also reflect the hydrography of the Albanian district as these lakes are connected to one another: the lakes Prespa and Ohrid through a subterranean karstic channel, lakes Ohrid and Skadar through the Bojana-Drina river system. Lake Ohrid is considered to be the oldest continuously existing lake in Europe with a likely age of three to five million years. An extraordinarily high degree of endemism, including more than 210 described endemic species, renders the lake a unique aquatic ecosystem of worldwide importance (
|
||
<bibRefCitation id="62054B3DFFCBFFAF632CF9DCFEBFF9C0" author="Albrecht" pageId="7" pageNumber="386" refString="Albrecht, C. & Wilke, T. (2008) Ancient Lake Ohrid: biodiversity and evolution. Hydrobiologia, 615, 103 - 140. http: // dx. doi. org / 10.1007 / s 10750 - 008 - 9558 - y" type="journal article" year="2008">Albrecht & Wilke, 2008</bibRefCitation>
|
||
), and the center of origins of native fish and biodiversity in the Albanian district. Lake Skadar, on the contrary, is relatively young and originated through flood of a karstic field occupying a crypto-depression of recent origin (
|
||
<bibRefCitation id="62054B3DFFCBFFAF66D8F94FFE5FF908" author="Lasca" box="[236,453,1700,1725]" pageId="7" pageNumber="386" refString="Lasca, N. P., Radulovic, V., Ristic, R. J. & Cherkauer, D. S. (1981) Geology, hydrology, climate and bathymetry of Lake Skadar. In: Karaman, G. S. (Ed.), The biota and Limnology of Lake Skadar. Smithsonian Institution D. C. USA, printed in Yugoslavia, Titograd, pp. 17 - 38." type="book chapter" year="1981">
|
||
Lasca
|
||
<emphasis id="34E0EADEFFCBFFAF670FF94EFEEAF909" box="[315,368,1700,1725]" italics="true" pageId="7" pageNumber="386">et al</emphasis>
|
||
., 1981
|
||
</bibRefCitation>
|
||
); it is thus reasonable to hypothesize that its fish fauna originated through multiple colonization events from surrounding water bodies.
|
||
</paragraph>
|
||
<paragraph id="062B36CCFFCBFFA066F3F907FE76FEA8" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,392]" lastPageId="8" lastPageNumber="387" pageId="7" pageNumber="386">
|
||
<emphasis id="34E0EADEFFCBFFAF66F3F907FE54F8B1" bold="true" box="[199,462,1772,1797]" pageId="7" pageNumber="386">
|
||
<typeStatus id="D92F886EFFCBFFAF66F3F907FEA7F8B1" box="[199,317,1772,1797]" pageId="7" pageNumber="386" type="lectotype">Lectotype</typeStatus>
|
||
designation
|
||
</emphasis>
|
||
. Due to the high diversity of the genus
|
||
<taxonomicName id="C1944D4FFFCBFFAF65ACF907FC7DF8B1" ID-CoL="643L5" box="[920,999,1772,1797]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="386" phylum="Chordata" rank="genus">
|
||
<emphasis id="34E0EADEFFCBFFAF65ACF907FC7DF8B1" box="[920,999,1772,1797]" italics="true" pageId="7" pageNumber="386">Rutilus</emphasis>
|
||
</taxonomicName>
|
||
complex in the Balkan Peninsula (see Introduction and
|
||
<bibRefCitation id="62054B3DFFCBFFAF676AF8E4FDC3F89C" author="Ketmaier" box="[350,601,1807,1832]" pageId="7" pageNumber="386" refString="Ketmaier, V., Bianco, P. G. & Durand, J. D. (2008) Molecular systematics, phylogeny and biogeography of roaches (Rutilus, Teleostei, Cyprinidae). Molecular Phylogenetics and Evolution, 49, 362 - 367. http: // dx. doi. org / 10.1016 / j. ympev. 2008.07.012" type="journal article" year="2008">
|
||
Ketmaier
|
||
<emphasis id="34E0EADEFFCBFFAF67E4F8FAFD88F89C" box="[464,530,1807,1832]" italics="true" pageId="7" pageNumber="386">et al.,</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
), a
|
||
<typeStatus id="D92F886EFFCBFFAF64B2F8E4FD77F89C" box="[646,749,1807,1832]" pageId="7" pageNumber="386" type="lectotype">lectotype</typeStatus>
|
||
is here selected among the four
|
||
<typeStatus id="D92F886EFFCBFFAF625AF8FBFB4AF89C" box="[1134,1232,1808,1832]" pageId="7" pageNumber="386" type="syntype">syntypes</typeStatus>
|
||
housed in NMW 50723, and 50725. The specimen,
|
||
<quantity id="C16C9B29FFCBFFAF641EF8DFFD13F8F8" box="[554,649,1844,1868]" metricMagnitude="-1" metricUnit="m" metricValue="1.17" pageId="7" pageNumber="386" unit="mm" value="117.0">117 mm</quantity>
|
||
SL (
|
||
<quantity id="C16C9B29FFCBFFAF64F3F8DFFCB2F8F8" box="[711,808,1844,1869]" metricMagnitude="-1" metricUnit="m" metricValue="1.51" pageId="7" pageNumber="386" unit="mm" value="151.0">151 mm</quantity>
|
||
TL), bearing the catalogue number NMW 50723-1 is designed as the
|
||
<typeStatus id="D92F886EFFCBFFAF677FF8BCFE2CF8C4" box="[331,438,1879,1904]" pageId="7" pageNumber="386" type="lectotype">lectotype</typeStatus>
|
||
. A picture of that specimen was published in
|
||
<bibRefCitation id="62054B3DFFCBFFAF658EF8BCFA02F8C4" author="Bianco" box="[954,1432,1879,1904]" pageId="7" pageNumber="386" refString="Bianco, P. G. & Taraborelli, T. (1985) Contributo alla conoscenza del genere Rutilus R. in Italia e nei balcani occidentali (Pisces, Cyprinidae). Bollettino del Museo Regionale di Scienze naturali di Torino, 3, 131 - 172." type="journal article" year="1985" yearSuffix="p">Bianco & Taraborelli (1985: fig. 6, p.149)</bibRefCitation>
|
||
. According to ICZN (1999), other things being equal, preference for selection of a
|
||
<typeStatus id="D92F886EFFCBFFAF6271F897FB31F821" box="[1093,1195,1916,1941]" pageId="7" pageNumber="386" type="lectotype">lectotype</typeStatus>
|
||
should be given to a
|
||
<typeStatus id="D92F886EFFCBFFAF66A3F84AFF74F80D" box="[151,238,1953,1977]" pageId="7" pageNumber="386" type="syntype">syntype</typeStatus>
|
||
that has been illustrated in a publication (art 74. recommendation 74A). The
|
||
<typeStatus id="D92F886EFFCBFFAF627BF84BFB2CF80D" box="[1103,1206,1952,1977]" pageId="7" pageNumber="386" type="lectotype">lectotype</typeStatus>
|
||
is in good condition (
|
||
<figureCitation id="9EAF2A49FFCBFFAF66ABF82FFF78F868" box="[159,226,1988,2013]" captionStart="FIGURE 3. A" captionStartId="6.[151,250,1241,1263]" captionTargetBox="[151,1436,257,1219]" captionTargetId="figure@6.[151,1436,257,1220]" captionTargetPageId="6" captionText="FIGURE 3. A) Lectotype of Leucos aula, 74 mm SL (ANSP 6434), Padua Province, northern Italy; B) Lectotype of Leucos basak, 117 mm SL (NMW 50723 - 1), Vergoraz and Lake Drusino near Imotsky, Croatia; C) Lectotype of Sarmarutilus rubilio, 93 mm SL (ANSP 6509), Lake Nemi, Central Italy." httpUri="https://zenodo.org/record/225781/files/figure.png" pageId="7" pageNumber="386">Fig. 3</figureCitation>
|
||
B). It has 40 pored scales on LL; 9 above and 4 below LL; 9 branched dorsal rays and 8 branched anal rays; pharyngeal teeth formula, 5-5. Head length about 3.8 times in SL; head depth 4.8 times in SL; body depth about 4.0 times in SL, least body depth about 2.3 times in body depth; snout length about 3.4 times in head length. Dorsal and anal fins slightly concave; caudal fin forked; origin of D at same level as origin of P2; mouth terminal and slightly oblique; dorsum quite convex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E8E6547FFC4FFA066F3FECCFAF9FE3C" pageId="8" pageNumber="387" type="discussion">
|
||
<paragraph id="062B36CCFFC4FFA066F3FECCFAF9FE3C" blockId="8.[151,1436,151,392]" pageId="8" pageNumber="387">
|
||
<emphasis id="34E0EADEFFC4FFA066F3FECCFD61FEF4" bold="true" box="[199,763,295,320]" pageId="8" pageNumber="387">Remarks on ecology, biology and conservation.</emphasis>
|
||
It is a still-water adapted species, invasive in lakes Skadar, Ohrid and lakes of Adriatic drainages of
|
||
<collectingCountry id="7E83765CFFC4FFA06469FEA7FD28FED1" box="[605,690,332,357]" name="Croatia" pageId="8" pageNumber="387">Croatia</collectingCountry>
|
||
, and low course of rivers. Spawning takes place between April and May (
|
||
<bibRefCitation id="62054B3DFFC4FFA066ECFE84FE46FE3C" author="Macrovcic" box="[216,476,367,392]" pageId="8" pageNumber="387" refString="Macrovcic, M., Brigic, A., Buj, I., Caleta, M., Mustafic, P. & Zanella, D. (2006) Red book of freshwater fishes of Croatia. Ministry of Culture, State Institute for Nature Protection, Zagreb, 253 pp." type="book" year="2006">
|
||
Mačrovcić
|
||
<emphasis id="34E0EADEFFC4FFA0676DFE9AFE13FE3C" box="[345,393,367,392]" italics="true" pageId="8" pageNumber="387">et al</emphasis>
|
||
., 2006
|
||
</bibRefCitation>
|
||
). Its conservation status is of “Low Concern” according to IUCN (2013) red list.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |