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<document id="3BD706F8B293FD33DDBAEF94DC7C4CDC" ID-DOI="10.11646/zootaxa.3841.3.4" ID-GBIF-Dataset="fd4f8193-3005-4d9b-ae7f-304b847215a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="225778" ID-ZooBank="E52D2F6B-631D-45FE-97E0-DD26137757FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460462027068" checkinUser="plazi" docAuthor="Bianco, Pier Giorgio &amp; Ketmaier, Valerio" docDate="2014" docId="8E3D87DAFFC0FFA466A3FF7CFD06F992" docLanguage="en" docName="zt03841p402.pdf" docOrigin="Zootaxa 3841 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucos panosi Bogutskaya &amp; Iliadou 2006" docType="treatment" docVersion="9" lastPageNumber="391" masterDocId="7204FFA2FFCCFFA86634FFEBFF9AFFB4" masterDocTitle="A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)" masterLastPageNumber="402" masterPageNumber="379" pageNumber="391" updateTime="1698304242431" updateUser="plazi">
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<mods:title id="B5106901D91550CD604AF37F13612C01">A revision of the Rutilus complex from Mediterranean Europe with description of a new genus, Sarmarutilus, and a new species, Rutilus stoumboudae (Teleostei: Cyprinidae)</mods:title>
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<mods:namePart id="8FD9EAEB1E2F8D628C94761B160B857D">Bianco, Pier Giorgio</mods:namePart>
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<mods:date id="41457FD351D0ABF30667D21147E5A011">2014</mods:date>
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<treatment id="8E3D87DAFFC0FFA466A3FF7CFD06F992" ID-DOI="http://doi.org/10.5281/zenodo.6140509" ID-GBIF-Taxon="119512649" ID-Zenodo-Dep="6140509" LSID="urn:lsid:plazi:treatment:8E3D87DAFFC0FFA466A3FF7CFD06F992" httpUri="http://treatment.plazi.org/id/8E3D87DAFFC0FFA466A3FF7CFD06F992" lastPageNumber="391" pageId="12" pageNumber="391">
<subSubSection id="4E8E6547FFC0FFA466A3FF7CFF65FF67" pageId="12" pageNumber="391" type="nomenclature">
<paragraph id="062B36CCFFC0FFA466A3FF7CFD26FF06" blockId="12.[151,700,151,211]" box="[151,700,151,178]" pageId="12" pageNumber="391">
<heading id="5D6381A0FFC0FFA466A3FF7CFD26FF06" bold="true" box="[151,700,151,178]" fontSize="11" level="1" pageId="12" pageNumber="391" reason="1">
<taxonomicName id="C1944D4FFFC0FFA466A3FF7CFD26FF06" ID-CoL="725YM" authority="Bogutskaya &amp; Iliadou, 2006" authorityName="Bogutskaya &amp; Iliadou" authorityYear="2006" box="[151,700,151,178]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA466A3FF7CFD26FF06" bold="true" box="[151,700,151,178]" pageId="12" pageNumber="391">
<emphasis id="34E0EADEFFC0FFA466A3FF7CFEDEFF05" bold="true" box="[151,324,151,177]" italics="true" pageId="12" pageNumber="391">Leucos panosi</emphasis>
(
<bibRefCitation id="62054B3DFFC0FFA46767FF73FD29FF06" author="Bogutskaya" box="[339,691,152,178]" pageId="12" pageNumber="391" refString="Bogutskaya, N. G. &amp; Iliadou, K. (2006) Rutilus panosi, a new roach from western Greece (Teleostei: Cyprinidae). Zoosystematica Rossica, 14, 293 - 298." type="journal article" year="2006">Bogutskaya &amp; Iliadou, 2006</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="062B36CCFFC0FFA466A3FF51FF65FF67" blockId="12.[151,700,151,211]" box="[151,255,186,211]" pageId="12" pageNumber="391">
(
<figureCitation id="9EAF2A49FFC0FFA466ABFF51FF78FF67" box="[159,226,186,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="12" pageNumber="391">Fig. 2</figureCitation>
D)
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFC0FFA466A3FEE9FB72FEAC" box="[151,1256,258,280]" pageId="12" pageNumber="391" type="reference_group">
<paragraph id="062B36CCFFC0FFA466A3FEE9FB72FEAC" blockId="12.[151,1256,258,280]" box="[151,1256,258,280]" pageId="12" pageNumber="391">
<treatmentCitationGroup id="268411E2FFC0FFA466A3FEE9FB72FEAC" box="[151,1256,258,280]" pageId="12" pageNumber="391">
<taxonomicName id="C1944D4FFFC0FFA466A3FEE9FD1BFEAC" ID-CoL="4TTHL" authority="Bogutskaya &amp; Iliadou, 2006: 283" authorityName="Bogutskaya &amp; Iliadou" authorityPageNumber="283" authorityYear="2006" box="[151,641,258,280]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA466A3FEE9FEB3FEAC" box="[151,297,258,280]" italics="true" pageId="12" pageNumber="391">Rutilus panosi</emphasis>
<treatmentCitation id="873510DDFFC0FFA4671BFEE9FD1BFEAC" author="Bogutskaya" box="[303,641,258,280]" page="283" pageId="12" pageNumber="391" year="2006">
<bibRefCitation id="62054B3DFFC0FFA4671BFEE9FDC8FEAC" author="Bogutskaya" box="[303,594,258,280]" pageId="12" pageNumber="391" refString="Bogutskaya, N. G. &amp; Iliadou, K. (2006) Rutilus panosi, a new roach from western Greece (Teleostei: Cyprinidae). Zoosystematica Rossica, 14, 293 - 298." type="journal article" year="2006">Bogutskaya &amp; Iliadou, 2006</bibRefCitation>
: 283
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="D92F886EFFC0FFA464BAFEE9FD21FEAC" box="[654,699,258,280]" pageId="12" pageNumber="391">type</typeStatus>
localities rivers Achelos and Louros, western
<collectingCountry id="7E83765CFFC0FFA462A5FEE9FB44FEAC" box="[1169,1246,258,280]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFC0FFA466A3FEAEFD17FE7E" pageId="12" pageNumber="391" type="materials_examined">
<paragraph id="062B36CCFFC0FFA466A3FEAEFD17FE7E" blockId="12.[151,1437,325,1574]" pageId="12" pageNumber="391">
<emphasis id="34E0EADEFFC0FFA466A3FEAEFE13FEEA" bold="true" box="[151,393,325,350]" pageId="12" pageNumber="391">Examined materials</emphasis>
.
<collectionCode id="6085AE09FFC0FFA467ACFEADFE5DFEEA" box="[408,455,326,350]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="12" pageNumber="391">IZA</collectionCode>
0 416, 7,
<collectingCountry id="7E83765CFFC0FFA46400FEADFD1CFEEA" box="[564,646,326,350]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
, L. Trichonis; P.G. Bianco,
<date id="722A100CFFC0FFA465F5FEAEFBFAFEEA" box="[961,1120,325,350]" pageId="12" pageNumber="391" value="1987-04-25">25 April 1987</date>
.—
<collectionCode id="6085AE09FFC0FFA462B5FEADFB28FEEA" box="[1153,1202,326,350]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="12" pageNumber="391">IZA</collectionCode>
8766,
<collectingCountry id="7E83765CFFC0FFA46336FEADFACEFEEA" box="[1282,1364,326,350]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
, canal emissary of L. Trichonis, P.G. Bianco,
<date id="722A100CFFC0FFA4646FFE82FD73FE35" box="[603,745,361,386]" pageId="12" pageNumber="391" value="1987-05-01">1 May 1987</date>
.—
<collectionCode id="6085AE09FFC0FFA4653EFE82FCA1FE35" box="[778,827,361,385]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="12" pageNumber="391">IZA</collectionCode>
0 417,
<collectingCountry id="7E83765CFFC0FFA465BBFE82FC7BFE35" box="[911,993,361,385]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
, L. Trichonis, P.G. Bianco,
<date id="722A100CFFC0FFA46311FE82FF54FE12" pageId="12" pageNumber="391" value="1998-08-10">10 August 1998</date>
.—
<collectionCode id="6085AE09FFC0FFA466DBFE65FEBAFE12" box="[239,288,398,422]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="12" pageNumber="391">IZA</collectionCode>
85513, 28,
<collectingCountry id="7E83765CFFC0FFA46793FE65FE63FE12" box="[423,505,398,422]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
, L. Ambrakia, P.G. Bianco,
<date id="722A100CFFC0FFA46507FE65FC4BFE12" box="[819,977,397,422]" pageId="12" pageNumber="391" value="1984-04-30">30 April 1984</date>
.—
<collectionCode id="6085AE09FFC0FFA465C6FE65FBB9FE12" box="[1010,1059,398,422]" httpUri="http://grbio.org/cool/9hrc-zefw" name="Universita di l'Aguila, Instituto di Zoologia" pageId="12" pageNumber="391">IZA</collectionCode>
83108B, 27,
<collectingCountry id="7E83765CFFC0FFA4628FFE65FA97FE12" box="[1211,1293,398,422]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
, L. Joannina (introduced), P.G. Bianco,
<date id="722A100CFFC0FFA4678BFE5AFD13FE7E" box="[447,649,433,458]" pageId="12" pageNumber="391" value="1976-02-23">23 February 1976</date>
.
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFC0FFA466F3FE3EFD07FCBA" pageId="12" pageNumber="391" type="diagnosis">
<paragraph id="062B36CCFFC0FFA466F3FE3EFD07FCBA" blockId="12.[151,1437,325,1574]" pageId="12" pageNumber="391">
<emphasis id="34E0EADEFFC0FFA466F3FE3EFEDBFE5A" bold="true" box="[199,321,469,494]" pageId="12" pageNumber="391">Diagnosis.</emphasis>
A species of
<taxonomicName id="C1944D4FFFC0FFA467D9FE3DFDA6FE59" box="[493,572,470,493]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="34E0EADEFFC0FFA467D9FE3DFDA6FE59" box="[493,572,470,493]" italics="true" pageId="12" pageNumber="391">Leucos</emphasis>
</taxonomicName>
characterized by moderate size, not exceeding
<quantity id="C16C9B29FFC0FFA46246FE3EFA96FE59" box="[1138,1292,469,494]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="12" pageNumber="391" unit="mm" value="165.0" valueMax="170.0" valueMin="160.0">160170 mm</quantity>
SL, usually
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SL; absence of a mid-lateral band; peritoneal membrane blackened by dense melanophores.
<taxonomicName id="C1944D4FFFC0FFA46378FE11FF78FD81" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA46378FE11FF78FD81" italics="true" pageId="12" pageNumber="391">Leucos panosi</emphasis>
</taxonomicName>
is quite similar to
<taxonomicName id="C1944D4FFFC0FFA46781FDF5FDA1FD82" box="[437,571,541,566]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="ylikiensis">
<emphasis id="34E0EADEFFC0FFA46781FDF5FDA1FD82" box="[437,571,541,566]" italics="true" pageId="12" pageNumber="391">L. ylikiensis</emphasis>
</taxonomicName>
from which it differs mostly for the number of total GR, usually
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<taxonomicName id="C1944D4FFFC0FFA463B1FDF5FE9AFDEE" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="ylikiensis">
<emphasis id="34E0EADEFFC0FFA463B1FDF5FE9AFDEE" italics="true" pageId="12" pageNumber="391">L. ylikiensis</emphasis>
</taxonomicName>
and
<quantity id="C16C9B29FFC0FFA46700FDAAFE0CFDEE" box="[308,406,577,602]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="12" pageNumber="391" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">1314 in</quantity>
<taxonomicName id="C1944D4FFFC0FFA467A8FDA9FD9EFDED" box="[412,516,578,601]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA467A8FDA9FD9EFDED" box="[412,516,578,601]" italics="true" pageId="12" pageNumber="391">L. panosi</emphasis>
</taxonomicName>
. It may be distinguished from
<taxonomicName id="C1944D4FFFC0FFA4656CFDA9FC2DFDEE" box="[856,951,577,602]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="basak">
<emphasis id="34E0EADEFFC0FFA4656CFDA9FC2DFDEE" box="[856,951,577,602]" italics="true" pageId="12" pageNumber="391">L. basak</emphasis>
</taxonomicName>
for the number of LL scales, usually 4243 as against
<quantity id="C16C9B29FFC0FFA46739FD8EFEEBFDCA" box="[269,369,613,638]" metricMagnitude="-1" metricUnit="m" metricValue="9.398" metricValueMax="9.652" metricValueMin="9.144" pageId="12" pageNumber="391" unit="in" value="37.0" valueMax="38.0" valueMin="36.0">3638 in</quantity>
<taxonomicName id="C1944D4FFFC0FFA4674CFD8DFE4DFDCA" box="[376,471,613,638]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="basak">
<emphasis id="34E0EADEFFC0FFA4674CFD8DFE4DFDCA" box="[376,471,613,638]" italics="true" pageId="12" pageNumber="391">L. basak</emphasis>
</taxonomicName>
. It may be distinguished from
<taxonomicName id="C1944D4FFFC0FFA46500FD8DFC1FFDCA" box="[820,901,613,638]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="aula">
<emphasis id="34E0EADEFFC0FFA46500FD8DFC1FFDCA" box="[820,901,613,638]" italics="true" pageId="12" pageNumber="391">L. aula</emphasis>
</taxonomicName>
for the absence of lateral band and the color of the peritoneal membrane, black in
<taxonomicName id="C1944D4FFFC0FFA4642DFD61FD1BFD15" box="[537,641,650,673]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA4642DFD61FD1BFD15" box="[537,641,650,673]" italics="true" pageId="12" pageNumber="391">L. panosi</emphasis>
</taxonomicName>
and silvery in
<taxonomicName id="C1944D4FFFC0FFA46511FD61FCEEFD16" box="[805,884,649,674]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="aula">
<emphasis id="34E0EADEFFC0FFA46511FD61FCEEFD16" box="[805,884,649,674]" italics="true" pageId="12" pageNumber="391">L. aula</emphasis>
</taxonomicName>
. It differs from all other species of
<taxonomicName id="C1944D4FFFC0FFA462C9FD61FAD6FD15" box="[1277,1356,650,673]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="34E0EADEFFC0FFA462C9FD61FAD6FD15" box="[1277,1356,650,673]" italics="true" pageId="12" pageNumber="391">Leucos</emphasis>
</taxonomicName>
for the high number of total GR, usually
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<taxonomicName id="C1944D4FFFC0FFA4644FFD45FD79FD71" box="[635,739,686,709]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA4644FFD45FD79FD71" box="[635,739,686,709]" italics="true" pageId="12" pageNumber="391">L. panosi</emphasis>
</taxonomicName>
and
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all others, excepted
<taxonomicName id="C1944D4FFFC0FFA46262FD45FB47FD72" box="[1110,1245,685,710]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="ylikiensis">
<emphasis id="34E0EADEFFC0FFA46262FD45FB47FD72" box="[1110,1245,685,710]" italics="true" pageId="12" pageNumber="391">L. ylikiensis</emphasis>
</taxonomicName>
that has a higher number of total GR, usually 1820. The closest species, both on geographical distribution and molecular grounds (
<figureCitation id="9EAF2A49FFC0FFA466ABFD1EFF45FCBA" box="[159,223,757,782]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1610,1632]" captionTargetBox="[182,1420,597,1530]" captionTargetId="figure@2.[151,1436,591,1588]" captionTargetPageId="2" captionText="FIGURE 1. A) Collecting localities (L. = lakes, R. = rivers) of Rutilus complex species. 1, R. Thames; 2, R. Danube (Slovakia); 3, R. Danube (Romania); 4, L. Como; 5, L. Maggiore; 6, R. Arno and R. Serchio; 7, R. Albegna and R. Cecina; 8, R. Ombrone and R. Fiora; 9, R. Tiber and R. Mignone; 10, L. Trasimeno; 11, R. Liri and R. Garigliano; 12, R. Volturno and R. Sele; 13, R. Pescara and R. Tavo; 14, R. Trigno; 15, R. Tronto; 16, R. Livenza; 17, L. Pernica; 18, R. Sava; 19, L. Drusino; 20, R. Krupa (R. Neretva basin); 21, L. Besanka (R. Neretva basin); 22, R. Moraca and R. Zeta; 23, L. Skadar; 24, L. Dojran; 25, L. Horid; 26, L. Prespa and L. Micraprespa; 27, L. Vegoritis; 28, L. Volvi; 29, L. Joannina; 30, L. Yliki and L. Paralimni; 31, L. Ambrakia; 32, L. Trichonis. Bold numbers are the different peri-Mediterranean ichthyogeographic districts described in Bianco (1990). B) A schematic temporal reconstruction of the diversification of the genus in southern Mediterranean Europe based on cyt- b is shown (modified from Ketmaier et al., 2008). For each taxon the corresponding number of the district of origin is shown; triangles indicate that multiple populations were analyzed. Numbers at nodes are divergence times in million years; all labeled nodes were robustly supported in the phylogenetic reconstructions (see Ketmaier et al., 2008 for further details)." httpUri="https://zenodo.org/record/225779/files/figure.png" pageId="12" pageNumber="391">Fig. 1</figureCitation>
B) is
<taxonomicName id="C1944D4FFFC0FFA4672AFD1DFE3EFCBA" box="[286,420,757,782]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="ylikiensis">
<emphasis id="34E0EADEFFC0FFA4672AFD1DFE3EFCBA" box="[286,420,757,782]" italics="true" pageId="12" pageNumber="391">L. ylikiensis</emphasis>
</taxonomicName>
from western
<collectingCountry id="7E83765CFFC0FFA46472FD1DFD03FCBA" box="[582,665,758,782]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFC0FFA466F3FCF2FBCDFAB2" pageId="12" pageNumber="391" type="description">
<paragraph id="062B36CCFFC0FFA466F3FCF2FBCDFAB2" blockId="12.[151,1437,325,1574]" pageId="12" pageNumber="391">
<emphasis id="34E0EADEFFC0FFA466F3FCF2FEC3FC86" bold="true" box="[199,345,793,818]" pageId="12" pageNumber="391">Description.</emphasis>
Body uniformly silvery without longitudinal band; color of eye in living fish, yellowish; fins yellowish or pale grayish in preserved specimens; a triangular pale, crescent spots at insertion of each scale of the body; snout pointed; pre-orbital distance near equal to the horizontal diameter of the eye; lips smooth; mouth terminal, or slightly inferior, its opening oblique; the corner of maxillae at the same level or slightly overpass the vertical crossing the anterior border of the orbit; profile of dorsum convex, slightly humped; paired and unpaired fins yellowish with few scattered melanophores; free margin of dorsal and anal fin concave; caudal fin moderately forked; peritoneal membrane blackened by several dense and fused melanophores; head length about 4.24.6 times in SL; body deep about 2.83.2 times in the SL; origin of the D placed slightly above the vertical crossing the insertion of the P2; LL complete, and extending from the margin of opercular membrane, to the end of caudal peduncle; 3945 pored scales, usually 4243; 7.5 above and 3.5 below the LL; 14 circum-peduncular scales; constantly 3 un-branched rays followed by 9 branched rays in the D; constantly 3 un-branched and usually 8 branched rays in the anal fin; P1 with 1 un-branched and 1416 branched rays; P2 with 1 un-branched ray and constantly 8 branched rays; 1216 total GR; fine granular tubercles on head and checks has been observed in adult reproductive males. For additional description and shape, see
<figureCitation id="9EAF2A49FFC0FFA4657FFB06FC16FAB2" box="[843,908,1261,1286]" captionStart="FIGURE 2" captionStartId="4.[151,250,1514,1536]" captionTargetBox="[173,1424,552,1475]" captionTargetId="figure@4.[151,1436,519,1492]" captionTargetPageId="4" captionText="FIGURE 2. Native species of the Rutilus complex in southern Mediterranean Europe: A) Leucos aula, 115 mm SL, R. Bacchiglione, northern Italy, April 1996 (IZA 0425); B) Leucos basak, 95 mm SL, Lake Ohrid, FYROM, August 1987, (IZA 0429); C) Leucos ylikiensis, 120 mm SL, Lake Yliki, April 1987 (IZA 8733); D) Leucos panosi, 130 mm SL, canal to Lake Trichonis, May 1987 (IZA 8766); E) Leucos albus, 98 mm SL, River Moraca, Montenegro, November 1999 (IZA 0421); F) Sarmarutilus rubilio, 100 mm SL, River Ofanto, southern Italy, February 2002 (IZA 02183); G) Rutilus pigus, 230 mm SL, Lake Major, northern Italy, (IZA 0474); H) Holotype of Rutilus stoumboudae new species, 148 mm SL, October 1998, Lake Volvi, Greece (IZA 02107)." httpUri="https://zenodo.org/record/225780/files/figure.png" pageId="12" pageNumber="391">Fig. 2</figureCitation>
D and Tables 1, 2.
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFC0FFA466F3FAFAFAD3FAC6" pageId="12" pageNumber="391" type="distribution">
<paragraph id="062B36CCFFC0FFA466F3FAFAFAD3FAC6" blockId="12.[151,1437,325,1574]" pageId="12" pageNumber="391">
<emphasis id="34E0EADEFFC0FFA466F3FAFAFEC3FA9E" bold="true" box="[199,345,1297,1322]" pageId="12" pageNumber="391">Distribution</emphasis>
. The species was reported for the Acheloos, and Louros river-systems in western
<collectingCountry id="7E83765CFFC0FFA46379FAF9FA06FA9E" box="[1357,1436,1298,1322]" name="Greece" pageId="12" pageNumber="391">Greece</collectingCountry>
(
<bibRefCitation id="62054B3DFFC0FFA466AAFADEFE72FAFA" author="Bogutskaya" box="[158,488,1333,1358]" pageId="12" pageNumber="391" refString="Bogutskaya, N. G. &amp; Iliadou, K. (2006) Rutilus panosi, a new roach from western Greece (Teleostei: Cyprinidae). Zoosystematica Rossica, 14, 293 - 298." type="journal article" year="2006">Bogutskaya &amp; Iliadou, 2006</bibRefCitation>
). The species was found also in lakes Trichonis and Ambrakia.
<taxonomicName id="C1944D4FFFC0FFA462F0FADDFAF2FAF9" box="[1220,1384,1334,1357]" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA462F0FADDFAF2FAF9" box="[1220,1384,1334,1357]" italics="true" pageId="12" pageNumber="391">Leucos panosi</emphasis>
</taxonomicName>
was introduced in Lake Joannina, (
<bibRefCitation id="62054B3DFFC0FFA467DEFAB2FD71FAC6" author="Leonardos" box="[490,747,1369,1394]" pageId="12" pageNumber="391" refString="Leonardos, D., Kagalou, I. I., Tsoumani, M. &amp; Economidis, P. S. (2008) Fish fauna in a protected Greek lake: biodiversity, introduced fish species over a 80 - year period and their impacts on the ecosystem. Ecology of Freshwater Fish, 17, 165 - 173. http: // dx. doi. org / 10.1111 / j. 1600 - 0633.2007.00268. x" type="journal article" year="2008">
Leonardos
<emphasis id="34E0EADEFFC0FFA4645DFAB1FD32FAC6" box="[617,680,1369,1394]" italics="true" pageId="12" pageNumber="391">et al.,</emphasis>
2008
</bibRefCitation>
), where it coexists with the introduced
<emphasis id="34E0EADEFFC0FFA46297FAB2FAD3FAC6" box="[1187,1353,1369,1394]" italics="true" pageId="12" pageNumber="391">
<taxonomicName id="C1944D4FFFC0FFA46297FAB2FADFFAC6" box="[1187,1349,1369,1394]" class="Actinopterygii" family="Cyprinidae" genus="Rutilus" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="rutilus">Rutilus rutilus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4E8E6547FFC0FFA466F3FA96FD06F992" pageId="12" pageNumber="391" type="discussion">
<paragraph id="062B36CCFFC0FFA466F3FA96FD06F992" blockId="12.[151,1437,325,1574]" pageId="12" pageNumber="391">
<emphasis id="34E0EADEFFC0FFA466F3FA96FD67FA22" bold="true" box="[199,765,1405,1430]" pageId="12" pageNumber="391">Remarks on ecology, biology and conservation.</emphasis>
A still-water adapted species. In fish communities,
<taxonomicName id="C1944D4FFFC0FFA46378FA95FF78FA0D" family="Cyprinidae" genus="Leucos" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="391" phylum="Chordata" rank="species" species="panosi">
<emphasis id="34E0EADEFFC0FFA46378FA95FF78FA0D" italics="true" pageId="12" pageNumber="391">Leucos panosi</emphasis>
</taxonomicName>
tends to become dominant and invasive in lakes while it is quite rare in the flowing part of rivers Louros and Acheloos, (P.G. Bianco, pers. obs.). Spawners have been collected in February suggesting that the reproductive season may start at the end of January and extend possibly until MarchApril. The species is classified as of “Low Concern” among the IUCN (2013) categories.
</paragraph>
</subSubSection>
</treatment>
</document>