600 lines
69 KiB
XML
600 lines
69 KiB
XML
<document id="50FED4EFB56C644217D30F4D16AA8DC1" ID-ISSN="1175-5326" ID-ZooBank="1A7BCEC8-39FA-43E4-9303-08FFA241270E" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629832097455" checkinUser="felipe" docAuthor="Shibukawa, Koichi, Phousavanh, Phouvin, Phongsa, Koneuma & Iwata, Akihisa" docDate="2012" docId="8E36878DFFADFF8D34DDFA3C7CDCFBA7" docLanguage="en" docName="zt03586p271.pdf" docOrigin="Zootaxa 3586" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Metzia bounthobi Shibukawa & Phousavanh & Phongsa & Iwata 2012, new species" docType="treatment" docVersion="2" lastPageNumber="269" masterDocId="720FFFF5FFACFF88344AFFF97F42FF93" masterDocTitle="A new species of Metzia (Cypriniformes: Cyprinidae) from Northern Laos" masterLastPageNumber="271" masterPageNumber="264" pageNumber="265" updateTime="1699118568820" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="A5B677CDDCC4A33B910D969DAC064B28" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="768511D9FD105F4A83A1D3FD31B9F985">
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<mods:title id="0E2F2496EDD81D0A79617E22A6BD1372">A new species of Metzia (Cypriniformes: Cyprinidae) from Northern Laos</mods:title>
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</mods:titleInfo>
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<mods:name id="98FDD213E9518C4DF1FEA9608CCC5E60" type="personal">
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<mods:role id="6E477B631FD65570512D286FE579616F">
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<mods:roleTerm id="E47C9F60CF176D6393FCD7F2FE670891">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="17E5DD48875154AB30502E71C2BF8B6E">Shibukawa, Koichi</mods:namePart>
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<mods:affiliation id="0BBF2CEB7A1F1DEA34A23C9F4D006BB4">Nagao Natural Environment Foundation, 3 - 10 - 10 Shitaya, Taito-ku, Tokyo 110 - 0004, Japan.</mods:affiliation>
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</mods:name>
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<mods:name id="3461146EC7CD58DB997B240BE1B462CE" type="personal">
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<mods:role id="C88DE1E7C14106AEB944ECAC50C1632A">
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<mods:roleTerm id="2F24B559FDCC8D320E668BD3F84A6588">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7DD275C2F58D3659AAD4E18149609700">Phousavanh, Phouvin</mods:namePart>
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<mods:affiliation id="4479A2558A2F29E057776B2BBBFEA0B5">Faculty of Agriculture, National University of Laos, P. O. Box 7322, Nabong, Vientiane, Lao PDR. & Laboratory of Ecology and Environment, Division of Southeast Asian Area Studies, Graduate School of Asian and African Area Studies, Kyoto University Yoshida, Shimoadachi-cho 46, Sakyo-ku, Kyoto 606 - 8501, Japan. E-mail: aiwata @ asafas. kyoto-u. ac. jp Department of Biology, Faculty of Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Lao PDR.</mods:affiliation>
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</mods:name>
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<mods:name id="D0674F06892FF72FED1F6D5995042E5E" type="personal">
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<mods:roleTerm id="4B9995DA8B3A7AFD090479D635463879">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="02D83A1D95236321E720578D44FF6EE8">Phongsa, Koneuma</mods:namePart>
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</mods:name>
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<mods:name id="66572B28D535B65CAEA1D172CF427506" type="personal">
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<mods:role id="1D26BF2C286BBF6199224BE56DD900F4">
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<mods:roleTerm id="7E1CF5690786C4305468F4E513BE6536">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1FA1CAA74B9230953D99A15D267A75B4">Iwata, Akihisa</mods:namePart>
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<mods:affiliation id="9C8E3410B1B36A6A183A3D58BC61CE34">Laboratory of Ecology and Environment, Division of Southeast Asian Area Studies, Graduate School of Asian and African Area Studies, Kyoto University Yoshida, Shimoadachi-cho 46, Sakyo-ku, Kyoto 606 - 8501, Japan. E-mail: aiwata @ asafas. kyoto-u. ac. jp Department of Biology, Faculty of Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Lao PDR.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="AA7348049CA3F7FB97FB0CC1C99415AC">text</mods:typeOfResource>
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<mods:titleInfo id="97DD89F9163CB398B0D91E8D6C0A051E">
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<mods:title id="856EB45BA374FC9F329D0BF6F81624AB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="547098ECB03F978591B7D5C927BAF9B0">
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<mods:date id="2C4C11A68959FC6B32F2BB797D06AFF8">2012</mods:date>
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<mods:detail id="03862B79C9D5D7FFDC51DB40F0DAE918" type="pubDate">
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<mods:number id="78D67FFFF701BEA504A9D167A7DA1F4F">2012-12-14</mods:number>
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</mods:detail>
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<mods:detail id="E4E31A2659C5FD4402554629C2F3F9CC" type="volume">
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<mods:number id="DA46ADA36F0B0A5CDD13AC1B64C86C1F">3586</mods:number>
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<mods:start id="C66759D3C50220DDE05856952AE7AA93">264</mods:start>
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<mods:end id="DDEFF39764361E72D0AD76B0AF7006E0">271</mods:end>
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</mods:extent>
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<mods:classification id="562832BDA468E754DC33E0F4F8F522B1">journal article</mods:classification>
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<mods:identifier id="0AF41A18D1FFBE094C0F53E20814CA4D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="54D1420B2B92F9B16CA30DDA4F3322F2" type="ZooBank">1A7BCEC8-39FA-43E4-9303-08FFA241270E</mods:identifier>
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</mods:mods>
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<treatment id="8E36878DFFADFF8D34DDFA3C7CDCFBA7" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7" httpUri="http://treatment.plazi.org/id/8E36878DFFADFF8D34DDFA3C7CDCFBA7" lastPageId="5" lastPageNumber="269" pageId="1" pageNumber="265">
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<subSubSection id="4E856510FFADFF8934DDFA3C7EBBFA6C" pageId="1" pageNumber="265" type="nomenclature">
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<paragraph id="0620369BFFADFF8934DDFA3C7D48FA4C" blockId="1.[151,522,1477,1572]" box="[151,522,1477,1503]" pageId="1" pageNumber="265">
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<heading id="5D6881F7FFADFF8934DDFA3C7D48FA4C" bold="true" box="[151,522,1477,1503]" fontSize="11" level="1" pageId="1" pageNumber="265" reason="1">
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<emphasis id="34EBEA89FFADFF8934DDFA3C7D48FA4C" bold="true" box="[151,522,1477,1503]" pageId="1" pageNumber="265">
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<taxonomicName id="C19F4D18FFADFF8934DDFA3C7E29FA4C" ID-CoL="42KGC" authority="Shibukawa & Phousavanh & Phongsa & Iwata, 2012" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[151,363,1477,1503]" class="Actinopterygii" family="Cyprinidae" genus="Metzia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="265" phylum="Chordata" rank="species" species="bounthobi" status="sp. nov.">
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<emphasis id="34EBEA89FFADFF8934DDFA3C7E29FA4C" bold="true" box="[151,363,1477,1503]" italics="true" pageId="1" pageNumber="265">Metzia bounthobi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="2FD857F2FFADFF893530FA3C7D48FA4C" box="[378,522,1477,1503]" pageId="1" pageNumber="265" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="0620369BFFADFF8934DDFA117EBBFA6C" blockId="1.[151,522,1477,1572]" box="[151,505,1511,1535]" pageId="1" pageNumber="265">New Laotian name: Pa tep thob</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4E856510FFADFF8934DDF9F57E53F9B7" box="[151,273,1548,1572]" pageId="1" pageNumber="265" type="description">
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||
<paragraph id="0620369BFFADFF8934DDF9F57E53F9B7" blockId="1.[151,522,1477,1572]" box="[151,273,1548,1572]" pageId="1" pageNumber="265">
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||
(
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<figureCitation id="9EA42A1EFFADFF8934D5F9F57E4BF9B7" box="[159,265,1548,1572]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1380,1403]" captionStartId-1="3.[151,250,979,1002]" captionStartId-2="4.[151,250,1706,1729]" captionTargetBox-0="[151,1436,467,1364]" captionTargetBox-1="[151,1436,193,958]" captionTargetBox-2="[258,1316,177,1650]" captionTargetId-0="figure-458@2.[151,1436,451,1364]" captionTargetId-1="figure-530@3.[151,1436,193,958]" captionTargetId-2="figure-243@4.[258,1329,177,1689]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Freshly collected specimen of Meztia bounthobi (holotype, 57.0 mm SL, NIFI 4680), Mekong Basin in Sopvan, Nyoi District, Luang Prabang Province, Lao PDR (20°36.5’N, 102°39.1’E), 23 June 2010. Same specimen above and below but on different backgrounds to provide full view of characters of the species. Photographed by T. Phommavong." captionText-1="FIGURE 2. Preserved specimen of Meztia bounthobi, new species (holotype, 57.0 mm SL, NIFI 4680). (A) lateral, (B) dorsal and (C) ventral views. Photographed by K. Phongsa." captionText-2="FIGURE 3. Selected osteological features of Metzia bounthobi, new species. Features from one paratype (55.1 mm SL) of NUOL-P 4902: (A) lateral view of jaws, suspensorium and opercular bones; (B) lateral view of infraorbital bones; (C) mesial (a) and anterior (b) views of fifth ceratobranchial; (D) lateral view of anterior part of axial skeletal complex; (E) lateral view of caudal skeletal complex; (F) dorsal fin and its supports; (G) anal fin and its supports. Bars indicate 2 mm. Abbreviations: AN, anguloarticular; C, claustrum; CE, centrum; CIHPU, interhaemal spine cartilage; DEN, dentary; DR, distal radial; ECPT, ectopterygoid; ENPT, endopterygoid; EP, epural; FR, fin ray; H, hypural; HY, hyomandibular; IO, infraorbital; IOP, interopercle; IOS, inner arm of os suspensorium; L, lateral process of vertebral centrum; MPT, metapterygoid; MR, medial radial; MX, maxilla; NS, neural spine; OP, opercle; OPC, opisthural cartilage; OSS, outer arm of os suspensorium; PAL, palatine; PH, parhypural; PLS, pleurostyle; PMX, premaxilla; POP, preopercle; PR, proximal radial; Q, quadrate; R, rib; RA, retroarticular; S, scaphium; SN, supraneural; SOP, subopercle; ST, stay; SY, symplectic; T, tripus; UN, uroneural. Photographed and retouched by K. Phongsa and K. Shibukawa, respectively." pageId="1" pageNumber="265">Figs. 1–3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="4E856510FFADFF8A34DDF9B67E01FE3F" lastPageId="2" lastPageNumber="266" pageId="1" pageNumber="265" type="materials_examined">
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<paragraph id="0620369BFFADFF8934DDF9B67A37F918" blockId="1.[151,1436,1614,2035]" pageId="1" pageNumber="265">
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<materialsCitation id="B6F73CC6FFADFF8934DDF9B67A37F918" collectingDate="2010-06-23" collectionCode="NIFI" collectorName="B. Praxaysombath" county="Mekong" latitude="20.608334" location="Nyoi District" longLatPrecision="126" longitude="102.651665" municipality="Sopvan" pageId="1" pageNumber="265" specimenCode="NIFI 4680, NUOL-P 4060" specimenCount="1" stateProvince="Louangphabang" typeStatus="holotype">
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<emphasis id="34EBEA89FFADFF8934DDF9B67E49F9F4" bold="true" box="[151,267,1615,1639]" inLineHeading="true" pageId="1" pageNumber="265" reason="1">
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||
<typeStatus id="D9248839FFADFF8934DDF9B67E44F9F4" box="[151,262,1615,1639]" pageId="1" pageNumber="265" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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||
<specimenCode id="56399EE0FFADFF89355EF9B67ECEF9F5" box="[276,396,1614,1638]" collectionCode="NIFI" country="Thailand" httpUri="http://grbio.org/cool/1mv4-i4gm" name="National Inland Fisheries Institute" pageId="1" pageNumber="265">NIFI 4680</specimenCode>
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(previously
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<specimenCode id="56399EE0FFADFF893654F9B77D88F9F5" box="[542,714,1614,1638]" collectionCode="NUOL-P" pageId="1" pageNumber="265">NUOL-P 4060</specimenCode>
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), 57.0 mm SL,
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<collectingCounty id="EF414E17FFADFF8937CAF9B77C9FF9F5" box="[896,989,1614,1638]" pageId="1" pageNumber="265">Mekong</collectingCounty>
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basin in
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<collectingMunicipality id="E644ACE1FFADFF893000F9B77BE3F9F5" box="[1098,1185,1614,1638]" pageId="1" pageNumber="265">Sopvan</collectingMunicipality>
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,
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<location id="03406040FFADFF8930E4F9B77A05F9F5" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF8930E4F9B77A05F9F5" box="[1198,1351,1614,1638]" county="Mekong" latitude="20.608334" longLatPrecision="126" longitude="102.651665" municipality="Sopvan" name="Nyoi District" pageId="1" pageNumber="265" stateProvince="Louangphabang">Nyoi District</location>
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,
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<collectingRegion id="C45BF879FFADFF89311EF9B67E1DF919" country="Laos" name="Louangphabang" pageId="1" pageNumber="265">Luang Prabang Province</collectingRegion>
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,
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<location id="03406040FFADFF893521F98A7ED4F919" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF893521F98A7ED4F919" box="[363,406,1651,1674]" county="Mekong" latitude="20.608334" longLatPrecision="126" longitude="102.651665" municipality="Sopvan" name="Lao" pageId="1" pageNumber="265" stateProvince="Louangphabang">Lao</location>
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PDR (
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<geoCoordinate id="63AB505CFFADFF8935A9F98B7D1FF919" box="[483,605,1650,1674]" degrees="20" direction="north" minutes="36.5" orientation="latitude" pageId="1" pageNumber="265" precision="92" value="20.608334">20°36.5’N</geoCoordinate>
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,
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<geoCoordinate id="63AB505CFFADFF893622F98B7DA9F919" box="[616,747,1650,1674]" degrees="102" direction="east" minutes="39.1" orientation="longitude" pageId="1" pageNumber="265" precision="92" value="102.651665">102°39.1’E</geoCoordinate>
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),
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<date id="7221105BFFADFF89374AF98B7CDAF919" box="[768,920,1650,1674]" pageId="1" pageNumber="265" value="2010-06-23">
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<collectingDate id="6265E9B3FFADFF89374AF98B7CDAF919" box="[768,920,1650,1674]" pageId="1" pageNumber="265" value="2010-06-23">23 June 2010</collectingDate>
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</date>
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, collected by
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<collectorName id="AB6A534DFFADFF893078F98A7BBDF919" box="[1074,1279,1650,1674]" pageId="1" pageNumber="265">B. Praxaysombath</collectorName>
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and party.
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</materialsCitation>
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</paragraph>
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<paragraph id="0620369BFFADFF89348DF96E7EE7F815" blockId="1.[151,1436,1614,2035]" pageId="1" pageNumber="265">
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<materialsCitation id="B6F73CC6FFADFF89348DF96E7E74F941" collectingDate="2010-06-23" collectionCode="NIFI" collectorName="B. Praxaysombath" county="Mekong" latitude="20.608334" location="Nyoi District" longLatPrecision="126" longitude="102.651665" municipality="Sopvan" pageId="1" pageNumber="265" specimenCode="NSMT-P 105977" specimenCount="11" stateProvince="Louangphabang" typeStatus="paratype">
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<emphasis id="34EBEA89FFADFF89348DF96E7E07F93C" bold="true" box="[199,325,1687,1711]" inLineHeading="true" pageId="1" pageNumber="265" reason="1">
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<typeStatus id="D9248839FFADFF89348DF96E7E03F93C" box="[199,321,1687,1711]" pageId="1" pageNumber="265" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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Total
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<specimenCount id="1099FD12FFADFF8935C6F96F7D5AF93D" box="[396,536,1686,1710]" pageId="1" pageNumber="265" type="generic">9 specimens</specimenCount>
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||
, 40.5–61.0 mm SL:
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<specimenCode id="56399EE0FFADFF8936B6F96F7CFEF93D" box="[764,956,1686,1710]" collectionCode="NSMT-P" pageId="1" pageNumber="265">NSMT-P 105977</specimenCode>
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||
,
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<specimenCount id="1099FD12FFADFF893782F96F7B17F93D" box="[968,1109,1686,1710]" pageId="1" pageNumber="265" type="generic">2 specimens</specimenCount>
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||
, 53.4–58.4 mm SL, collected with holotype
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</materialsCitation>
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;
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<materialsCitation id="B6F73CC6FFADFF893509F9437EE7F815" collectingDate="2007-09-20" collectingDateMax="2009-10-19" collectingDateMin="2007-09-20" collectionCode="NIFI" collectorName="P. Phousavanh" county="Nam Pok River" latitude="20.608334" location="Samphan District" longLatPrecision="126" longitude="102.651665" municipality="Ou River" pageId="1" pageNumber="265" specimenCode="NULFA-P 715, NULFA-P 716, NUOL-P 4034, NUOL-P 4074, NUOL-P 4902" specimenCount="7" stateProvince="Phongsali" typeStatus="holotype">
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||
<specimenCode id="56399EE0FFADFF893509F9437EAEF941" box="[323,492,1722,1746]" collectionCode="NULFA-P" pageId="1" pageNumber="265">NULFA-P 715</specimenCode>
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||
,
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<specimenCount id="1099FD12FFADFF8935B3F9437D3BF941" box="[505,633,1722,1746]" pageId="1" pageNumber="265" type="generic">1 specimen</specimenCount>
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(cleared and stained), 41.9 mm SL,
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<collectingCounty id="EF414E17FFADFF893050F9427B89F941" box="[1050,1227,1722,1746]" pageId="1" pageNumber="265">Nam Pok River</collectingCounty>
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(a tributary of
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<collectingMunicipality id="E644ACE1FFADFF893130F9437F95F965" pageId="1" pageNumber="265">Ou River</collectingMunicipality>
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,
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<location id="03406040FFADFF8934A8F9277EEBF965" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF8934A8F9277EEBF965" box="[226,425,1758,1782]" county="Nam Pok River" municipality="Ou River" name="Samphan District" pageId="1" pageNumber="265" stateProvince="Phongsali">Samphan District</location>
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,
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<collectingRegion id="C45BF879FFADFF8935FFF9277DD5F965" box="[437,663,1758,1782]" country="Laos" name="Phongsali" pageId="1" pageNumber="265">Phongsaly Province</collectingRegion>
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,
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<location id="03406040FFADFF8936E9F9267D8CF965" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF8936E9F9267D8CF965" box="[675,718,1759,1782]" county="Nam Pok River" municipality="Ou River" name="Lao" pageId="1" pageNumber="265" stateProvince="Phongsali">Lao</location>
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PDR,
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<date id="7221105BFFADFF893750F9277C99F965" box="[794,987,1758,1782]" pageId="1" pageNumber="265" value="2009-10-19">
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<collectingDate id="6265E9B3FFADFF893750F9277C99F965" box="[794,987,1758,1782]" pageId="1" pageNumber="265" value="2009-10-19">19 October 2009</collectingDate>
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</date>
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, collected by
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<collectorName id="AB6A534DFFADFF89303CF9267A5EF965" box="[1142,1308,1758,1782]" pageId="1" pageNumber="265">P. Phousavanh</collectorName>
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;
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<specimenCode id="56399EE0FFADFF893162F9277F86F889" collectionCode="NULFA-P" pageId="1" pageNumber="265">NULFA-P 716</specimenCode>
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,
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<specimenCount id="1099FD12FFADFF89349BF8FB7E17F889" box="[209,341,1794,1818]" pageId="1" pageNumber="265" type="generic">1 specimen</specimenCount>
|
||
, 57.1 mm SL,
|
||
<location id="03406040FFADFF893649F8FB7D2DF889" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF893649F8FB7D2DF889" box="[515,623,1794,1818]" county="Nam Pok River" municipality="Ou River" name="Ou River" pageId="1" pageNumber="265" stateProvince="Phongsali">Ou River</location>
|
||
, Na,
|
||
<location id="03406040FFADFF8936E7F8FB7C34F889" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF8936E7F8FB7C34F889" box="[685,886,1794,1818]" county="Nam Pok River" municipality="Ou River" name="Samphan District" pageId="1" pageNumber="265" stateProvince="Phongsali">Samphan District</location>
|
||
,
|
||
<collectingRegion id="C45BF879FFADFF8937C9F8FB7CBBF889" box="[899,1017,1794,1818]" country="Laos" name="Phongsali" pageId="1" pageNumber="265">Phongsaly</collectingRegion>
|
||
,
|
||
<date id="7221105BFFADFF89304CF8FB7BA4F889" box="[1030,1254,1794,1818]" pageId="1" pageNumber="265" value="2007-09-20">
|
||
<collectingDate id="6265E9B3FFADFF89304CF8FB7BA4F889" box="[1030,1254,1794,1818]" pageId="1" pageNumber="265" value="2007-09-20">20 September 2007</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="AB6A534DFFADFF8931C2F8FA7E60F8AD" pageId="1" pageNumber="265">P. Phousavanh</collectorName>
|
||
;
|
||
<specimenCode id="56399EE0FFADFF893566F8DF7E97F8AD" box="[300,469,1830,1854]" collectionCode="NUOL-P" pageId="1" pageNumber="265">NUOL-P 4034</specimenCode>
|
||
,
|
||
<specimenCount id="1099FD12FFADFF8935AAF8DF7D29F8AD" box="[480,619,1830,1854]" pageId="1" pageNumber="265" type="generic">2 specimens</specimenCount>
|
||
, 40.5–61.0 mm SL, collected with holotype;
|
||
<specimenCode id="56399EE0FFADFF893029F8DF7A4EF8AD" box="[1123,1292,1830,1854]" collectionCode="NUOL-P" pageId="1" pageNumber="265">NUOL-P 4074</specimenCode>
|
||
,
|
||
<specimenCount id="1099FD12FFADFF89315CF8DF7AD5F8AD" box="[1302,1431,1830,1854]" pageId="1" pageNumber="265" type="generic">1 specimen</specimenCount>
|
||
, 58.0 mm SL, collected with
|
||
<typeStatus id="D9248839FFADFF8935A7F8B37D10F8F1" box="[493,594,1866,1890]" pageId="1" pageNumber="265" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCode id="56399EE0FFADFF893629F8B37C52F8F1" box="[611,784,1866,1890]" collectionCode="NUOL-P" pageId="1" pageNumber="265">NUOL-P 4902</specimenCode>
|
||
,
|
||
<specimenCount id="1099FD12FFADFF89376BF8B37CEDF8F1" box="[801,943,1866,1890]" pageId="1" pageNumber="265" type="generic">2 specimens</specimenCount>
|
||
(cleared and stained), 55.1–55.9 mm SL, collected with
|
||
<typeStatus id="D9248839FFADFF893576F8977EE0F815" box="[316,418,1902,1926]" pageId="1" pageNumber="265" type="holotype">holotype</typeStatus>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="0620369BFFADFF8A348DF86A7E01FE3F" blockId="1.[151,1436,1614,2035]" lastBlockId="2.[151,1436,151,428]" lastPageId="2" lastPageNumber="266" pageId="1" pageNumber="265">
|
||
<emphasis id="34EBEA89FFADFF89348DF86A7E82F838" bold="true" box="[199,448,1938,1963]" pageId="1" pageNumber="265">Non-type specimens.</emphasis>
|
||
Total
|
||
<materialsCitation id="B6F73CC6FFADFF893646F86B7B74F839" box="[524,1078,1938,1962]" pageId="1" pageNumber="265" specimenCode="NULFA-P 717" specimenCount="8">
|
||
<specimenCount id="1099FD12FFADFF893646F86B7DD9F839" box="[524,667,1938,1962]" pageId="1" pageNumber="265" type="generic">8 specimens</specimenCount>
|
||
, 33.0–40.1 mm SL:
|
||
<specimenCode id="56399EE0FFADFF8937C6F86B7B74F839" box="[908,1078,1938,1962]" collectionCode="NULFA-P" pageId="1" pageNumber="265">NULFA-P 717</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFADFF89300FF86B7E03F861" collectingDate="2007-08-25" collectorName="P. Phousavanh" county="Ou River" location="Lao" municipality="Tay" pageId="1" pageNumber="265" specimenCode="NULFA-P 718" specimenCount="1" stateProvince="Gnot Ou District, Phongsaly Province">
|
||
<specimenCount id="1099FD12FFADFF89300FF86B7B88F839" box="[1093,1226,1938,1962]" pageId="1" pageNumber="265" type="generic">1 specimen</specimenCount>
|
||
, 34.2 mm SL,
|
||
<collectingCounty id="EF414E17FFADFF893130F86B7F95F85D" pageId="1" pageNumber="265">Ou River</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="E644ACE1FFADFF8934A2F84E7E51F85C" box="[232,275,1975,1999]" pageId="1" pageNumber="265">Tay</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="C45BF879FFADFF89356EF84F7DABF85D" box="[292,745,1974,1998]" isEnumeration="true" pageId="1" pageNumber="265">Gnot Ou District, Phongsaly Province</collectingRegion>
|
||
,
|
||
<location id="03406040FFADFF8936B0F84E7C67F85D" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF8936B0F84E7C67F85D" box="[762,805,1975,1998]" county="Ou River" municipality="Tay" name="Lao" pageId="1" pageNumber="265" stateProvince="Gnot Ou District, Phongsaly Province">Lao</location>
|
||
PDR,
|
||
<date id="7221105BFFADFF893737F84F7B02F85D" box="[893,1088,1974,1998]" pageId="1" pageNumber="265" value="2007-08-25">
|
||
<collectingDate id="6265E9B3FFADFF893737F84F7B02F85D" box="[893,1088,1974,1998]" pageId="1" pageNumber="265" value="2007-08-25">25 August 2007</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="AB6A534DFFADFF8930A6F84E7ADAF85D" box="[1260,1432,1974,1998]" pageId="1" pageNumber="265">P. Phousavanh</collectorName>
|
||
;
|
||
<specimenCode id="56399EE0FFADFF8934DDF8237E03F861" box="[151,321,2010,2034]" collectionCode="NULFA-P" pageId="1" pageNumber="265">NULFA-P 718</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFADFF8A351BF8237B96FF3C" collectingDate="2008-08-15" collectorName="P. Phousavanh" county="Huay Tom River" lastPageId="2" lastPageNumber="266" location="Pak Bak" municipality="Ou River" pageId="1" pageNumber="265" specimenCode="NULFA-P 719" specimenCount="1" stateProvince="Ngoy District, Luang Prabang Province">
|
||
<specimenCount id="1099FD12FFADFF89351BF8237E94F861" box="[337,470,2010,2034]" pageId="1" pageNumber="265" type="generic">1 specimen</specimenCount>
|
||
, 33.0 mm SL,
|
||
<collectingCounty id="EF414E17FFADFF8936CDF8227C0BF861" box="[647,841,2010,2035]" pageId="1" pageNumber="265">Huay Tom River</collectingCounty>
|
||
(a tributary of
|
||
<collectingMunicipality id="E644ACE1FFADFF89304AF8237B2DF861" box="[1024,1135,2010,2034]" pageId="1" pageNumber="265">Ou River</collectingMunicipality>
|
||
),
|
||
<location id="03406040FFADFF8930CCF8237BA8F861" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFADFF8930CCF8237BA8F861" box="[1158,1258,2010,2034]" county="Huay Tom River" municipality="Ou River" name="Pak Bak" pageId="1" pageNumber="265" stateProvince="Ngoy District, Luang Prabang Province">Pak Bak</location>
|
||
,
|
||
<collectingRegion id="C45BF879FFADFF8A30B2F8227EEFFF3C" isEnumeration="true" lastPageId="2" lastPageNumber="266" pageId="1" pageNumber="265">Ngoy District, Luang Prabang Province</collectingRegion>
|
||
,
|
||
<location id="03406040FFAEFF8A35F2FF617EA1FF3C" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFAEFF8A35F2FF617EA1FF3C" box="[440,483,152,175]" county="Huay Tom River" municipality="Ou River" name="Lao" pageId="2" pageNumber="266" stateProvince="Ngoy District, Luang Prabang Province">Lao</location>
|
||
PDR,
|
||
<date id="7221105BFFAEFF8A3665FF6E7DA1FF3C" box="[559,739,151,175]" pageId="2" pageNumber="266" value="2008-08-15">
|
||
<collectingDate id="6265E9B3FFAEFF8A3665FF6E7DA1FF3C" box="[559,739,151,175]" pageId="2" pageNumber="266" value="2008-08-15">15 August 2008</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="AB6A534DFFAEFF8A3737FF617B60FF3C" box="[893,1058,151,175]" pageId="2" pageNumber="266">P. Phousavanh</collectorName>
|
||
;
|
||
<specimenCode id="56399EE0FFAEFF8A3066FF6E7B96FF3C" box="[1068,1236,151,175]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 719</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFAEFF8A3095FF6E7DC1FF64" collectingDate="2008-08-16" collectorName="A. Iwata & P. Phousavanh" county="Ou River" location="Lao" municipality="Pak Bak" pageId="2" pageNumber="266" specimenCode="NULFA-P 720" specimenCount="1" stateProvince="Ngoy District, Luang Prabang Province">
|
||
<specimenCount id="1099FD12FFAEFF8A3095FF6E7A23FF3C" box="[1247,1377,151,175]" pageId="2" pageNumber="266" type="generic">1 specimen</specimenCount>
|
||
, 33.1 mm SL,
|
||
<collectingCounty id="EF414E17FFAEFF8A34B0FF457E26FF47" box="[250,356,188,212]" pageId="2" pageNumber="266">Ou River</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="E644ACE1FFAEFF8A353AFF457E90FF47" box="[368,466,188,212]" pageId="2" pageNumber="266">Pak Bak</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="C45BF879FFAEFF8A3594FF457CE2FF47" box="[478,928,188,212]" isEnumeration="true" pageId="2" pageNumber="266">Ngoy District, Luang Prabang Province</collectingRegion>
|
||
,
|
||
<location id="03406040FFAEFF8A37E7FF457C9AFF40" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFAEFF8A37E7FF457C9AFF40" box="[941,984,188,211]" county="Ou River" municipality="Pak Bak" name="Lao" pageId="2" pageNumber="266" stateProvince="Ngoy District, Luang Prabang Province">Lao</location>
|
||
PDR.,
|
||
<date id="7221105BFFAEFF8A3066FF457BA6FF47" box="[1068,1252,188,212]" pageId="2" pageNumber="266" value="2008-08-16">
|
||
<collectingDate id="6265E9B3FFAEFF8A3066FF457BA6FF47" box="[1068,1252,188,212]" pageId="2" pageNumber="266" value="2008-08-16">16 August 2008</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="AB6A534DFFAEFF8A31CBFF457F97FF64" pageId="2" pageNumber="266">A. Iwata</collectorName>
|
||
and
|
||
<collectorName id="AB6A534DFFAEFF8A3552FF197E81FF64" box="[280,451,223,247]" pageId="2" pageNumber="266">P. Phousavanh</collectorName>
|
||
;
|
||
<specimenCode id="56399EE0FFAEFF8A359EFF267DC1FF64" box="[468,643,223,247]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 720</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFAEFF8A36DEFF267B25FE8F" collectingDate="2007-09-19" collectorName="P. Phousavanh" location="Hat Sa" municipality="Ou River" pageId="2" pageNumber="266" specimenCode="NULFA-P 721" specimenCount="1" stateProvince="Phongsaly District, Phongsaly Province">
|
||
<specimenCount id="1099FD12FFAEFF8A36DEFF267C5EFF64" box="[660,796,223,247]" pageId="2" pageNumber="266" type="generic">1 specimen</specimenCount>
|
||
, 40.1 mm SL,
|
||
<collectingMunicipality id="E644ACE1FFAEFF8A3793FF267B0AFF64" box="[985,1096,223,247]" pageId="2" pageNumber="266">Ou River</collectingMunicipality>
|
||
,
|
||
<location id="03406040FFAEFF8A3013FF197BEFFF64" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFAEFF8A3013FF197BEFFF64" box="[1113,1197,223,247]" municipality="Ou River" name="Hat Sa" pageId="2" pageNumber="266" stateProvince="Phongsaly District, Phongsaly Province">Hat Sa</location>
|
||
,
|
||
<collectingRegion id="C45BF879FFAEFF8A30F5FF267E39FE8F" isEnumeration="true" pageId="2" pageNumber="266">Phongsaly District, Phongsaly Province</collectingRegion>
|
||
,
|
||
<date id="7221105BFFAEFF8A35C2FEFD7D2BFE8F" box="[392,617,260,284]" pageId="2" pageNumber="266" value="2007-09-19">
|
||
<collectingDate id="6265E9B3FFAEFF8A35C2FEFD7D2BFE8F" box="[392,617,260,284]" pageId="2" pageNumber="266" value="2007-09-19">19 September 2007</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="AB6A534DFFAEFF8A3743FEFD7CF2FE8F" box="[777,944,260,284]" pageId="2" pageNumber="266">P. Phousavanh</collectorName>
|
||
;
|
||
<specimenCode id="56399EE0FFAEFF8A37F7FEFD7B25FE8F" box="[957,1127,260,284]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 721</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFAEFF8A303FFEFD7EA7FEAC" pageId="2" pageNumber="266" specimenCode="NULFA-P 720" specimenCount="1" stateProvince="Phongsaly District, Phongsaly Province">
|
||
<specimenCount id="1099FD12FFAEFF8A303FFEFD7BBBFE8F" box="[1141,1273,260,284]" pageId="2" pageNumber="266" type="generic">1 specimen</specimenCount>
|
||
, 38.7 mm SL, collected with
|
||
<specimenCode id="56399EE0FFAEFF8A3576FEDE7EA7FEAC" box="[316,485,295,319]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 720</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B6F73CC6FFAEFF8A35BAFEDE7DDBFEAC" box="[496,665,295,319]" isTailEnumeration="true" pageId="2" pageNumber="266" specimenCode="NULFA-P 722" specimenCount="1" stateProvince="Phongsaly District, Phongsaly Province">
|
||
<specimenCode id="56399EE0FFAEFF8A35BAFEDE7DDBFEAC" box="[496,665,295,319]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 722</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFAEFF8A36EEFEDE7AC0FEF7" collectingDate="2009-10-19" collectorName="P. Phousavanh" county="Nam Pok River" location="Lao" municipality="Ou River" pageId="2" pageNumber="266" specimenCode="NULFA-P 723" specimenCount="1" stateProvince="Samphan District, Phongsaly Province">
|
||
<specimenCount id="1099FD12FFAEFF8A36EEFEDE7C64FEAC" box="[676,806,295,319]" pageId="2" pageNumber="266" type="generic">1 specimen</specimenCount>
|
||
, 38.3 mm SL,
|
||
<collectingCounty id="EF414E17FFAEFF8A3781FED17B38FEAC" box="[971,1146,295,319]" pageId="2" pageNumber="266">Nam Pok River</collectingCounty>
|
||
(a tributary of
|
||
<collectingMunicipality id="E644ACE1FFAEFF8A316EFEDE7ACDFEAC" box="[1316,1423,295,319]" pageId="2" pageNumber="266">Ou River</collectingMunicipality>
|
||
),
|
||
<collectingRegion id="C45BF879FFAEFF8A34DDFEB57D09FEF7" box="[151,587,332,356]" isEnumeration="true" pageId="2" pageNumber="266">Samphan District, Phongsaly Province</collectingRegion>
|
||
,
|
||
<location id="03406040FFAEFF8A361DFEB57DC0FEF0" LSID="urn:lsid:plazi:treatment:8E36878DFFADFF8D34DDFA3C7CDCFBA7:03406040FFAEFF8A361DFEB57DC0FEF0" box="[599,642,332,355]" county="Nam Pok River" municipality="Ou River" name="Lao" pageId="2" pageNumber="266" stateProvince="Samphan District, Phongsaly Province">Lao</location>
|
||
PDR,
|
||
<date id="7221105BFFAEFF8A3684FEB57CCCFEF7" box="[718,910,332,356]" pageId="2" pageNumber="266" value="2009-10-19">
|
||
<collectingDate id="6265E9B3FFAEFF8A3684FEB57CCCFEF7" box="[718,910,332,356]" pageId="2" pageNumber="266" value="2009-10-19">19 October 2009</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="AB6A534DFFAEFF8A3063FEB57B8DFEF7" box="[1065,1231,332,356]" pageId="2" pageNumber="266">P. Phousavanh</collectorName>
|
||
;
|
||
<specimenCode id="56399EE0FFAEFF8A3090FEB57AC0FEF7" box="[1242,1410,332,356]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 723</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFAEFF8A31C4FEB57C47FE14" pageId="2" pageNumber="266" specimenCode="NULFA-P 722" specimenCount="1" stateProvince="Samphan District, Phongsaly Province">
|
||
<specimenCount id="1099FD12FFAEFF8A31C4FEB57E46FE14" pageId="2" pageNumber="266" type="generic">1 specimen</specimenCount>
|
||
, 36.4 mm SL, collected with
|
||
<specimenCode id="56399EE0FFAEFF8A3610FE967C47FE14" box="[602,773,367,391]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 722</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B6F73CC6FFAEFF8A3759FE967CFFFE14" box="[787,957,367,391]" isTailEnumeration="true" pageId="2" pageNumber="266" specimenCode="NULFA-P 724" specimenCount="1" stateProvince="Samphan District, Phongsaly Province">
|
||
<specimenCode id="56399EE0FFAEFF8A3759FE967CFFFE14" box="[787,957,367,391]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 724</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B6F73CC6FFAEFF8A3781FE967E7DFE3F" pageId="2" pageNumber="266" specimenCode="NULFA-P 722" specimenCount="1" stateProvince="Samphan District, Phongsaly Province">
|
||
<specimenCount id="1099FD12FFAEFF8A3781FE967B0DFE14" box="[971,1103,367,391]" pageId="2" pageNumber="266" type="generic">1 specimen</specimenCount>
|
||
, 35.9 mm SL, collected with
|
||
<specimenCode id="56399EE0FFAEFF8A34DDFE6D7E7DFE3F" box="[151,319,404,428]" collectionCode="NULFA-P" pageId="2" pageNumber="266">NULFA-P 722</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="52E06613FFAEFF8A34DDFA9D7A53FA2B" pageId="2" pageNumber="266" startId="2.[151,250,1380,1403]" targetBox="[151,1436,467,1364]" targetPageId="2">
|
||
<paragraph id="0620369BFFAEFF8A34DDFA9D7A53FA2B" blockId="2.[151,1436,1380,1464]" pageId="2" pageNumber="266">
|
||
<emphasis id="34EBEA89FFAEFF8A34DDFA9D7E56FAE8" bold="true" box="[151,276,1380,1403]" pageId="2" pageNumber="266">FIGURE 1.</emphasis>
|
||
Freshly collected specimen of
|
||
<taxonomicName id="C19F4D18FFAEFF8A361CFA9C7C48FAE9" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[598,778,1380,1402]" class="Actinopterygii" family="Cyprinidae" genus="Meztia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="species" species="bounthobi">
|
||
<emphasis id="34EBEA89FFAEFF8A361CFA9C7C48FAE9" box="[598,778,1380,1402]" italics="true" pageId="2" pageNumber="266">Meztia bounthobi</emphasis>
|
||
</taxonomicName>
|
||
(holotype, 57.0 mm SL, NIFI 4680), Mekong Basin in Sopvan, Nyoi District, Luang Prabang Province, Lao PDR (20°36.5’N, 102°39.1’E), 23 June 2010. Same specimen above and below but on different backgrounds to provide full view of characters of the species. Photographed by T. Phommavong.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4E856510FFAEFF8A348DFA1D7BBAF867" pageId="2" pageNumber="266" type="diagnosis">
|
||
<paragraph id="0620369BFFAEFF8A348DFA1D7BBAF867" blockId="2.[151,1436,1508,2037]" pageId="2" pageNumber="266">
|
||
<emphasis id="34EBEA89FFAEFF8A348DFA1D7E03FA6E" bold="true" box="[199,321,1508,1533]" inLineHeading="true" pageId="2" pageNumber="266" reason="1">Diagnosis.</emphasis>
|
||
<taxonomicName id="C19F4D18FFAEFF8A3501FA1C7D50FA6E" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[331,530,1509,1533]" class="Actinopterygii" family="Cyprinidae" genus="Metzia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="species" species="bounthobi">
|
||
<emphasis id="34EBEA89FFAEFF8A3501FA1C7D50FA6E" box="[331,530,1509,1533]" italics="true" pageId="2" pageNumber="266">Metzia bounthobi</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from its congeners in having the following characters: 18–20 branched anal-fin rays (vs.
|
||
<quantity id="C1679B7EFFAEFF8A3585F9F17D71F9B3" box="[463,563,1544,1568]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.572" metricValueMin="2.54" pageId="2" pageNumber="266" unit="in" value="14.0" valueMax="18.0" valueMin="10.0">10–18 in</quantity>
|
||
the others); 49–55 lateral-line scale rows (vs. 35–48); 33–36 predorsal scale rows (vs. 15–20); 20–22 circumpeduncular scale rows (vs. 14–18); 8–10 gill rakers on outer surface of first gill arch (vs. 12–18). The new species also resembles species of
|
||
<taxonomicName id="C19F4D18FFAEFF8A3720F9A97B53F9FB" box="[874,1041,1616,1640]" class="Actinopterygii" family="Cyprinidae" genus="Hemiculterella" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A3720F9A97B53F9FB" box="[874,1041,1616,1640]" italics="true" pageId="2" pageNumber="266">Hemiculterella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C19F4D18FFAEFF8A3069F9A97BD5F9FB" authorityName="Jordan & Snyder" authorityYear="1900" box="[1059,1175,1616,1640]" class="Actinopterygii" family="Cyprinidae" genus="Ischikauia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A3069F9A97BD5F9FB" box="[1059,1175,1616,1640]" italics="true" pageId="2" pageNumber="266">Ischikauia</emphasis>
|
||
</taxonomicName>
|
||
and at least a part of
|
||
<taxonomicName id="C19F4D18FFAEFF8A34DDF98C7E59F91E" authorityName="Cockerell. These" authorityYear="1923" box="[151,283,1653,1677]" class="Actinopterygii" family="Cyprinidae" genus="Anabarilius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A34DDF98C7E59F91E" box="[151,283,1653,1677]" italics="true" pageId="2" pageNumber="266">Anabarilius</emphasis>
|
||
</taxonomicName>
|
||
in sharing a sharp keel restricted in development to the area between pelvic-fin base and anus, last unbranched dorsal-fin ray being soft, and an air bladder composed of two chambers. However,
|
||
<taxonomicName id="C19F4D18FFAEFF8A314AF9607ADEF923" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[1280,1436,1688,1712]" class="Actinopterygii" family="Cyprinidae" genus="Metzia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="species" species="bounthobi">
|
||
<emphasis id="34EBEA89FFAEFF8A314AF9607ADEF923" box="[1280,1436,1688,1712]" italics="true" pageId="2" pageNumber="266">M. bounthobi</emphasis>
|
||
</taxonomicName>
|
||
differs in having a notably rounded snout (vs. pointed in
|
||
<taxonomicName id="C19F4D18FFAEFF8A370CF9447CAFF946" box="[838,1005,1725,1749]" class="Actinopterygii" family="Cyprinidae" genus="Hemiculterella" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A370CF9447CAFF946" box="[838,1005,1725,1749]" italics="true" pageId="2" pageNumber="266">Hemiculterella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C19F4D18FFAEFF8A304AF9447B37F946" authorityName="Jordan & Snyder" authorityYear="1900" box="[1024,1141,1725,1749]" class="Actinopterygii" family="Cyprinidae" genus="Ischikauia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A304AF9447B37F946" box="[1024,1141,1725,1749]" italics="true" pageId="2" pageNumber="266">Ischikauia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C19F4D18FFAEFF8A30FFF9447A7BF946" authorityName="Cockerell. These" authorityYear="1923" box="[1205,1337,1725,1749]" class="Actinopterygii" family="Cyprinidae" genus="Anabarilius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A30FFF9447A7BF946" box="[1205,1337,1725,1749]" italics="true" pageId="2" pageNumber="266">Anabarilius</emphasis>
|
||
</taxonomicName>
|
||
), 18–20 branched anal-fin rays (vs.
|
||
<quantity id="C1679B7EFFAEFF8A358DF9197D5EF96B" box="[455,540,1760,1784]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="4.318" metricValueMin="2.032" pageId="2" pageNumber="266" unit="in" value="12.5" valueMax="17.0" valueMin="8.0">8–17 in</quantity>
|
||
<taxonomicName id="C19F4D18FFAEFF8A3668F9197C0EF96B" authority=", Ischiakuia" authorityName="Ischiakuia" box="[546,844,1760,1784]" class="Actinopterygii" family="Cyprinidae" genus="Hemiculterella" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A3668F9197D8BF96B" box="[546,713,1760,1784]" italics="true" pageId="2" pageNumber="266">Hemiculterella</emphasis>
|
||
,
|
||
<emphasis id="34EBEA89FFAEFF8A369DF9197C0EF96B" box="[727,844,1760,1784]" italics="true" pageId="2" pageNumber="266">Ischiakuia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C19F4D18FFAEFF8A37CBF9197B47F96B" authorityName="Cockerell. These" authorityYear="1923" box="[897,1029,1760,1784]" class="Actinopterygii" family="Cyprinidae" genus="Anabarilius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A37CBF9197B47F96B" box="[897,1029,1760,1784]" italics="true" pageId="2" pageNumber="266">Anabarilius</emphasis>
|
||
</taxonomicName>
|
||
), 49–55 lateral-line scales (vs. more than
|
||
<quantity id="C1679B7EFFAEFF8A3484F8FD7E45F88F" box="[206,263,1796,1820]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" pageId="2" pageNumber="266" unit="in" value="58.0">58 in</quantity>
|
||
<taxonomicName id="C19F4D18FFAEFF8A3544F8FC7EC0F88E" authorityName="Jordan & Snyder" authorityYear="1900" box="[270,386,1797,1821]" class="Actinopterygii" family="Cyprinidae" genus="Ischikauia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A3544F8FC7EC0F88E" box="[270,386,1797,1821]" italics="true" pageId="2" pageNumber="266">Ischikauia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C19F4D18FFAEFF8A35F0F8FC7D7CF88E" authorityName="Cockerell. These" authorityYear="1923" box="[442,574,1797,1821]" class="Actinopterygii" family="Cyprinidae" genus="Anabarilius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A35F0F8FC7D7CF88E" box="[442,574,1797,1821]" italics="true" pageId="2" pageNumber="266">Anabarilius</emphasis>
|
||
</taxonomicName>
|
||
, except for
|
||
<taxonomicName id="C19F4D18FFAEFF8A368EF8FC7C3DF88F" baseAuthorityName="Nichols" baseAuthorityYear="1925" box="[708,895,1797,1821]" class="Actinopterygii" family="Cyprinidae" genus="Anabarilius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="species" species="transmontana">
|
||
<emphasis id="34EBEA89FFAEFF8A368EF8FC7C3DF88F" box="[708,895,1797,1821]" italics="true" pageId="2" pageNumber="266">A. transmontana</emphasis>
|
||
</taxonomicName>
|
||
with 54–57), air bladder with rounded posterior end (vs. posterior end with a small lobe at least in
|
||
<taxonomicName id="C19F4D18FFAEFF8A36A1F8D17CD0F8D3" box="[747,914,1832,1856]" class="Actinopterygii" family="Cyprinidae" genus="Hemiculterella" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A36A1F8D17CD0F8D3" box="[747,914,1832,1856]" italics="true" pageId="2" pageNumber="266">Hemiculterella</emphasis>
|
||
</taxonomicName>
|
||
) and 39–40 vertebrae (vs. 40–43, 42–43 and
|
||
<quantity id="C1679B7EFFAEFF8A34DDF8B57E42F8F7" box="[151,256,1868,1892]" metricMagnitude="0" metricUnit="m" metricValue="1.1557" metricValueMax="1.2192" metricValueMin="1.0922" pageId="2" pageNumber="266" unit="in" value="45.5" valueMax="48.0" valueMin="43.0">43–48 in</quantity>
|
||
<taxonomicName id="C19F4D18FFAEFF8A3547F8B47EF6F8F6" box="[269,436,1869,1893]" class="Actinopterygii" family="Cyprinidae" genus="Hemiculterella" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A3547F8B47EF6F8F6" box="[269,436,1869,1893]" italics="true" pageId="2" pageNumber="266">Hemiculterella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C19F4D18FFAEFF8A3583F8B47D7CF8F6" authorityName="Jordan & Snyder" authorityYear="1900" box="[457,574,1869,1893]" class="Actinopterygii" family="Cyprinidae" genus="Ischikauia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A3583F8B47D7CF8F6" box="[457,574,1869,1893]" italics="true" pageId="2" pageNumber="266">Ischikauia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C19F4D18FFAEFF8A36CAF8B47C46F8F6" authorityName="Cockerell. These" authorityYear="1923" box="[640,772,1869,1893]" class="Actinopterygii" family="Cyprinidae" genus="Anabarilius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="genus">
|
||
<emphasis id="34EBEA89FFAEFF8A36CAF8B47C46F8F6" box="[640,772,1869,1893]" italics="true" pageId="2" pageNumber="266">Anabarilius</emphasis>
|
||
</taxonomicName>
|
||
, respectively). The new species is fairly similar to the sympatric
|
||
<taxonomicName id="C19F4D18FFAEFF8A3545F8897D42F81B" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[271,512,1904,1928]" class="Actinopterygii" family="Cyprinidae" genus="Paralaubuca" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="species" species="barroni">
|
||
<emphasis id="34EBEA89FFAEFF8A3545F8897D42F81B" box="[271,512,1904,1928]" italics="true" pageId="2" pageNumber="266">Paralaubuca barroni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="9EA42A1EFFAEFF8A3659F8897D1BF81B" box="[531,601,1904,1928]" captionStart="FIGURE 4" captionStartId="5.[151,250,1677,1700]" captionTargetBox="[151,1436,1129,1656]" captionTargetId="figure-500@5.[151,1436,1129,1656]" captionTargetPageId="5" captionText="FIGURE 4. Paralaubuca barroni, NUOL-P 4038, 75.4 mm SL, Nyoi District, Luang Prabang Province, Lao PDR. Photographed by T. Phommavong." pageId="2" pageNumber="266">Fig. 4</figureCitation>
|
||
) in counts of scales and the other general features, and can be possibly confused; however,
|
||
<taxonomicName id="C19F4D18FFAEFF8A353DF86C7DCDF83F" authority="(Fowler 1934)" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[375,655,1940,1965]" class="Actinopterygii" family="Cyprinidae" genus="Paralaubuca" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="266" phylum="Chordata" rank="species" species="barroni">
|
||
<emphasis id="34EBEA89FFAEFF8A353DF86C7EAAF83E" box="[375,488,1941,1965]" italics="true" pageId="2" pageNumber="266">P. barroni</emphasis>
|
||
(Fowler 1934)
|
||
</taxonomicName>
|
||
is readily distinguished from the former by having more anal-fin rays (24–29 vs.
|
||
<quantity id="C1679B7EFFAEFF8A3550F8417EC2F843" box="[282,384,1976,2000]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="2" pageNumber="266" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
|
||
the new species) and gill rakers (19–24 vs. 8–10) and a distinct, complete abdominal keel extending from anus to isthmus (
|
||
<bibRefCitation id="620E4B6AFFAEFF8A3642F8257DFDF867" author="Banarescu, P." box="[520,703,2012,2036]" pageId="2" pageNumber="266" pagination="347 - 355" refId="ref5066" refString="Banarescu, P. (1971) Revision of the genus Paralaubuca Bleeker (Pisces, Cypriniformes). Travaux du Museum d'Histoire Naturelle " Grigore Antipa ", 11, 347 - 355." type="journal article" year="1971">Bănărescu 1971</bibRefCitation>
|
||
; Luo & Chen
|
||
<emphasis id="34EBEA89FFAEFF8A3717F8247C31F867" box="[861,883,2013,2036]" italics="true" pageId="2" pageNumber="266">in</emphasis>
|
||
<bibRefCitation id="620E4B6AFFAEFF8A3730F8257B75F867" author="Chen, Y. & Chu, X. & Luo, Y. & Chen Y. & Liu, H. & He, M. & Chen, W. & Yue, P. & He, S & Lin, R." box="[890,1079,2012,2037]" pageId="2" pageNumber="266" refId="ref5172" refString="Chen, Y., Chu, X., Luo, Y., Chen Y., Liu, H., He, M., Chen, W., Yue, P., He, S & Lin, R. (1998) Fauna Sinica. Osteichthyes. Cypriniformes II. Science Press, Beijing, 531 pp." type="book" year="1998">
|
||
Chen
|
||
<emphasis id="34EBEA89FFAEFF8A37F6F8247CACF866" box="[956,1006,2013,2037]" italics="true" pageId="2" pageNumber="266">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="620E4B6AFFAEFF8A3008F8257BAEF867" author="Kottelat, M." box="[1090,1260,2012,2036]" pageId="2" pageNumber="266" refId="ref5347" refString="Kottelat, M. (2001 a) Fishes of Laos. WHT Publications, Sri Lanka, 198 pp." type="book" year="2001">Kottelat 2001a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="52E06613FFAFFF8B34DDFC2A7DEFFB9B" pageId="3" pageNumber="267" startId="3.[151,250,979,1002]" targetBox="[151,1436,193,958]" targetPageId="3">
|
||
<paragraph id="0620369BFFAFFF8B34DDFC2A7DEFFB9B" blockId="3.[151,1436,979,1032]" pageId="3" pageNumber="267">
|
||
<emphasis id="34EBEA89FFAFFF8B34DDFC2A7E51FC79" bold="true" box="[151,275,979,1002]" pageId="3" pageNumber="267">FIGURE 2.</emphasis>
|
||
Preserved specimen of
|
||
<taxonomicName id="C19F4D18FFAFFF8B364CFC2D7DFBFC7A" authority="Shibukawa & Phousavanh & Phongsa & Iwata, 2012" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[518,697,979,1001]" class="Actinopterygii" family="Cyprinidae" genus="Meztia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="267" phylum="Chordata" rank="species" species="bounthobi" status="sp. nov.">
|
||
<emphasis id="34EBEA89FFAFFF8B364CFC2D7DFBFC7A" box="[518,697,979,1001]" italics="true" pageId="3" pageNumber="267">Meztia bounthobi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2FD857F2FFAFFF8B368FFC2D7C7DFC7A" box="[709,831,979,1001]" pageId="3" pageNumber="267" rank="species">new species</taxonomicNameLabel>
|
||
(holotype, 57.0 mm SL, NIFI 4680). (A) lateral, (B) dorsal and (C) ventral views. Photographed by K. Phongsa.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4E856510FFAFFF8D348DFBCD7CE1FCCF" lastPageId="5" lastPageNumber="269" pageId="3" pageNumber="267" type="description">
|
||
<paragraph id="0620369BFFAFFF8B348DFBCD7D79FA03" blockId="3.[151,1437,1076,2037]" pageId="3" pageNumber="267">
|
||
<emphasis id="34EBEA89FFAFFF8B348DFBCD7E1AFBDE" bold="true" box="[199,344,1076,1101]" inLineHeading="true" pageId="3" pageNumber="267" reason="1">Description.</emphasis>
|
||
In the following description, the counts of the
|
||
<typeStatus id="D9248839FFAFFF8B3723FBCD7C89FBDF" box="[873,971,1076,1100]" pageId="3" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
are asterisked, and the frequency or each count is given in parentheses following the relevant count. Dorsal-fin rays 3 unbranched and 7* (10) branched rays, last one split to base; anal-fin rays 3 unbranched and 18* (4), 19 (2) or 20 (3) branched rays, last one split to base; total pectoral-fin rays 12* (7) or 13* (13), including uppermost 1* (20) and lowermost 1 (8) or 2* (12) unbranched rays; total pelvic-fin rays 8* (18), including anteriormost 1* (20) and innermost 1* (20) unbranched rays; branched caudal-fin rays 9+8* (10); lateral-line scales 49 (1), 50 (2), 51* (4), 52 (3), 53 (5), 54 (1) or 55* (2), followed by 2* (8), 3* (9) or 4 (1) pored scales on caudal fin; transverse scale rows above lateral line 9 (4), 10* (13) o:r 11* (1); transverse scale rows below lateral line 4* (16) or 5 (2); predorsal scale rows 33 (3), 34* (3) or 36 (2); circumpeduncular scale rows 20 (2), 21 (4) or 22* (3); gill rakers on outer surface of first gill arch 0+8 (1), 1+7 (2), 1+8* (3), 1+9 (2), 2+7 (1) or 2+8 (1).
|
||
</paragraph>
|
||
<paragraph id="0620369BFFAFFF8B348DFA657EFDF866" blockId="3.[151,1437,1076,2037]" pageId="3" pageNumber="267">Body compressed and moderately deep, with greatest body depth slightly anterior to pelvic-fin insertion. Caudal peduncle moderately deep (its depth 69.4–74.9% of its length), with narrowest part of caudal peduncle located nearer the posterior base of anal fin than to base of caudal fin. Ventral profile of head and body almost as arched as much as, or a slightly more arched than, dorsal profile. Distinct keel between anus and a vertical through posterior end of pelvic-fin base; keel preceded by a very weak midventral, blunt ridge extending anteriorly to a vertical through posterior half of pectoral fin. Snout rounded and short, shorter than eye diameter (snout length 69.4–80.3% of eye diameter). Eye large, midlaterally positioned at slightly anterior to middle of head. Interorbital width subequal or less than (89.5–99.1% of) eye diameter. Adipose eyelid weakly developed, covering half and less than one fourth of iris anteriorly and posteriorly, respectively. Mouth terminal, gape oblique, about 45 degrees relative to longitudinal axis of body. Maxillary extending beyond a vertical through anterior margin of eye, but not to anterior margin of pupil. Symphyseal knob of lower jaw not developed. Barbels absent. Nares closely positioned, with anterior naris as a short tube. Dorsal-fin origin nearer caudal-fin base than to tip of snout; its origin slightly nearer to vertical through pelvic-fin insertion than to anal-fin origin. Last unbranched ray of dorsal fin soft and nonossified. Pectoral fin short, not reaching to a vertical through pelvic-fin insertion. Pelvic fin short, its length 67.0–74.6% of length of pectoral fin; pelvic-fin insertion nearer snout tip than to caudal-fin base. Anal-fin base greater than dorsal-fin base (its length 188.8–220.0% of the latter). Caudal fin moderately forked, with rounded (not pointed) tips of lobes.</paragraph>
|
||
<caption id="52E06613FFA8FF8C34DDF9537C4FF860" pageId="4" pageNumber="268" startId="4.[151,250,1706,1729]" targetBox="[258,1316,177,1650]" targetPageId="4">
|
||
<paragraph id="0620369BFFA8FF8C34DDF9537C4FF860" blockId="4.[151,1436,1706,2035]" pageId="4" pageNumber="268">
|
||
<emphasis id="34EBEA89FFA8FF8C34DDF9537E57F952" bold="true" box="[151,277,1706,1729]" pageId="4" pageNumber="268">FIGURE 3.</emphasis>
|
||
Selected osteological features of
|
||
<taxonomicName id="C19F4D18FFA8FF8C363CF9527C68F953" authority="Shibukawa & Phousavanh & Phongsa & Iwata, 2012" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[630,810,1706,1728]" class="Actinopterygii" family="Cyprinidae" genus="Metzia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="268" phylum="Chordata" rank="species" species="bounthobi" status="sp. nov.">
|
||
<emphasis id="34EBEA89FFA8FF8C363CF9527C68F953" box="[630,810,1706,1728]" italics="true" pageId="4" pageNumber="268">Metzia bounthobi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2FD857F2FFA8FF8C3773F9527CFBF953" box="[825,953,1706,1728]" pageId="4" pageNumber="268" rank="species">new species</taxonomicNameLabel>
|
||
. Features from one paratype (55.1 mm SL) of NUOL-P 4902: (A) lateral view of jaws, suspensorium and opercular bones; (B) lateral view of infraorbital bones; (C) mesial (a) and anterior (b) views of fifth ceratobranchial; (D) lateral view of anterior part of axial skeletal complex; (E) lateral view of caudal skeletal complex; (F) dorsal fin and its supports; (G) anal fin and its supports. Bars indicate 2 mm. Abbreviations:
|
||
<emphasis id="34EBEA89FFA8FF8C313FF8FE7AD7F88E" box="[1397,1429,1799,1821]" italics="true" pageId="4" pageNumber="268">AN</emphasis>
|
||
, anguloarticular;
|
||
<emphasis id="34EBEA89FFA8FF8C3508F8DC7E11F8A8" box="[322,339,1829,1851]" italics="true" pageId="4" pageNumber="268">C</emphasis>
|
||
, claustrum;
|
||
<emphasis id="34EBEA89FFA8FF8C3593F8DC7EBBF8A8" box="[473,505,1829,1851]" italics="true" pageId="4" pageNumber="268">CE</emphasis>
|
||
, centrum;
|
||
<emphasis id="34EBEA89FFA8FF8C3625F8DC7DFEF8A8" box="[623,700,1829,1851]" italics="true" pageId="4" pageNumber="268">CIHPU</emphasis>
|
||
, interhaemal spine cartilage;
|
||
<emphasis id="34EBEA89FFA8FF8C37B3F8DF7B69F8A8" box="[1017,1067,1830,1851]" italics="true" pageId="4" pageNumber="268">DEN</emphasis>
|
||
, dentary;
|
||
<emphasis id="34EBEA89FFA8FF8C30D0F8DF7BFEF8A8" box="[1178,1212,1830,1851]" italics="true" pageId="4" pageNumber="268">DR</emphasis>
|
||
, distal radial;
|
||
<emphasis id="34EBEA89FFA8FF8C3110F8DC7AD4F8A8" box="[1370,1430,1829,1851]" italics="true" pageId="4" pageNumber="268">ECPT</emphasis>
|
||
, ectopterygoid;
|
||
<emphasis id="34EBEA89FFA8FF8C3572F8BD7E36F8CA" box="[312,372,1860,1881]" italics="true" pageId="4" pageNumber="268">ENPT</emphasis>
|
||
, endopterygoid;
|
||
<emphasis id="34EBEA89FFA8FF8C367BF8BD7D0FF8CA" box="[561,589,1860,1881]" italics="true" pageId="4" pageNumber="268">EP</emphasis>
|
||
, epural;
|
||
<emphasis id="34EBEA89FFA8FF8C36FDF8BD7D94F8CA" box="[695,726,1860,1881]" italics="true" pageId="4" pageNumber="268">FR</emphasis>
|
||
, fin ray;
|
||
<emphasis id="34EBEA89FFA8FF8C3706F8BD7C1CF8CA" box="[844,862,1860,1881]" italics="true" pageId="4" pageNumber="268">H</emphasis>
|
||
, hypural;
|
||
<emphasis id="34EBEA89FFA8FF8C379CF8BD7CB1F8CA" box="[982,1011,1860,1881]" italics="true" pageId="4" pageNumber="268">HY</emphasis>
|
||
, hyomandibular;
|
||
<emphasis id="34EBEA89FFA8FF8C30FDF8BD7B90F8C9" box="[1207,1234,1860,1882]" italics="true" pageId="4" pageNumber="268">IO</emphasis>
|
||
, infraorbital;
|
||
<emphasis id="34EBEA89FFA8FF8C3124F8BD7AD7F8C9" box="[1390,1429,1860,1882]" italics="true" pageId="4" pageNumber="268">IOP</emphasis>
|
||
, interopercle;
|
||
<emphasis id="34EBEA89FFA8FF8C356AF89B7E05F8EB" box="[288,327,1890,1912]" italics="true" pageId="4" pageNumber="268">IOS</emphasis>
|
||
, inner arm of os suspensorium;
|
||
<emphasis id="34EBEA89FFA8FF8C36D5F89A7DEFF8EB" box="[671,685,1891,1912]" italics="true" pageId="4" pageNumber="268">L</emphasis>
|
||
, lateral process of vertebral centrum;
|
||
<emphasis id="34EBEA89FFA8FF8C300AF89A7B33F8EB" box="[1088,1137,1891,1912]" italics="true" pageId="4" pageNumber="268">MPT</emphasis>
|
||
, metapterygoid;
|
||
<emphasis id="34EBEA89FFA8FF8C3169F89A7A05F8EB" box="[1315,1351,1891,1912]" italics="true" pageId="4" pageNumber="268">MR</emphasis>
|
||
, medial radial;
|
||
<emphasis id="34EBEA89FFA8FF8C34A8F87B7E44F804" box="[226,262,1922,1943]" italics="true" pageId="4" pageNumber="268">MX</emphasis>
|
||
, maxilla;
|
||
<emphasis id="34EBEA89FFA8FF8C353CF8787ED6F804" box="[374,404,1921,1943]" italics="true" pageId="4" pageNumber="268">NS</emphasis>
|
||
, neural spine;
|
||
<emphasis id="34EBEA89FFA8FF8C367CF8787D17F804" box="[566,597,1921,1943]" italics="true" pageId="4" pageNumber="268">OP</emphasis>
|
||
, opercle;
|
||
<emphasis id="34EBEA89FFA8FF8C3689F8787DB7F804" box="[707,757,1921,1943]" italics="true" pageId="4" pageNumber="268">OPC</emphasis>
|
||
, opisthural cartilage;
|
||
<emphasis id="34EBEA89FFA8FF8C3795F8787B48F804" box="[991,1034,1921,1943]" italics="true" pageId="4" pageNumber="268">OSS</emphasis>
|
||
, outer arm of os suspensorium;
|
||
<emphasis id="34EBEA89FFA8FF8C3126F8787AD4F804" box="[1388,1430,1921,1943]" italics="true" pageId="4" pageNumber="268">PAL</emphasis>
|
||
, palatine;
|
||
<emphasis id="34EBEA89FFA8FF8C34BCF8587E5AF825" box="[246,280,1953,1974]" italics="true" pageId="4" pageNumber="268">PH</emphasis>
|
||
, parhypural;
|
||
<emphasis id="34EBEA89FFA8FF8C35E8F8597E8EF825" box="[418,460,1952,1974]" italics="true" pageId="4" pageNumber="268">PLS</emphasis>
|
||
, pleurostyle;
|
||
<emphasis id="34EBEA89FFA8FF8C3610F8587DCCF825" box="[602,654,1953,1974]" italics="true" pageId="4" pageNumber="268">PMX</emphasis>
|
||
, premaxilla;
|
||
<emphasis id="34EBEA89FFA8FF8C3752F8597C04F825" box="[792,838,1952,1974]" italics="true" pageId="4" pageNumber="268">POP</emphasis>
|
||
, preopercle;
|
||
<emphasis id="34EBEA89FFA8FF8C3784F8587CAFF825" box="[974,1005,1953,1974]" italics="true" pageId="4" pageNumber="268">PR</emphasis>
|
||
, proximal radial;
|
||
<emphasis id="34EBEA89FFA8FF8C30EDF8597BFBF825" box="[1191,1209,1952,1974]" italics="true" pageId="4" pageNumber="268">Q</emphasis>
|
||
, quadrate;
|
||
<emphasis id="34EBEA89FFA8FF8C3164F8587A7FF825" box="[1326,1341,1953,1974]" italics="true" pageId="4" pageNumber="268">R</emphasis>
|
||
, rib;
|
||
<emphasis id="34EBEA89FFA8FF8C313DF8597AD4F825" box="[1399,1430,1952,1974]" italics="true" pageId="4" pageNumber="268">RA</emphasis>
|
||
, retroarticular;
|
||
<emphasis id="34EBEA89FFA8FF8C3563F8467E74F846" box="[297,310,1983,2005]" italics="true" pageId="4" pageNumber="268">S</emphasis>
|
||
, scaphium;
|
||
<emphasis id="34EBEA89FFA8FF8C35FBF8467E8CF846" box="[433,462,1983,2005]" italics="true" pageId="4" pageNumber="268">SN</emphasis>
|
||
, supraneural;
|
||
<emphasis id="34EBEA89FFA8FF8C362AF8467DCEF846" box="[608,652,1983,2005]" italics="true" pageId="4" pageNumber="268">SOP</emphasis>
|
||
, subopercle;
|
||
<emphasis id="34EBEA89FFA8FF8C375EF8467C6FF846" box="[788,813,1983,2005]" italics="true" pageId="4" pageNumber="268">ST</emphasis>
|
||
, stay;
|
||
<emphasis id="34EBEA89FFA8FF8C373AF8467CC5F846" box="[880,903,1983,2005]" italics="true" pageId="4" pageNumber="268">SY</emphasis>
|
||
, symplectic;
|
||
<emphasis id="34EBEA89FFA8FF8C305AF8467B5EF847" box="[1040,1052,1983,2004]" italics="true" pageId="4" pageNumber="268">T</emphasis>
|
||
, tripus;
|
||
<emphasis id="34EBEA89FFA8FF8C303BF8467BD6F847" box="[1137,1172,1983,2004]" italics="true" pageId="4" pageNumber="268">UN</emphasis>
|
||
, uroneural. Photographed and retouched by K. Phongsa and K. Shibukawa, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0620369BFFA9FF8D348DFF6E7D89FEF7" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="269">Scales cycloid. Lateral line complete, gradually bent ventrally anterior to pelvic-fin insertion, then running parallel with ventral profile and ending midlaterally at basal part of caudal fin; 2–3 lateral-line scales basally at midlateral caudal fin; pectoral and pelvic fin axillary scales long, extending slightly beyond a vertical through base of last ray of each fin. No scales on fins, except for 2–3 and 1–2 scale rows basally on caudal and anal fins, respectively. Air bladder with two chambers; anterior chamber elliptical, posterior chamber elongate (longer than anterior chamber) and with rounded posterior end.</paragraph>
|
||
<paragraph id="0620369BFFA9FF8D348DFE967D01FD10" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="269">
|
||
<emphasis id="34EBEA89FFA9FF8D348DFE967EEBFE1B" bold="true" box="[199,425,367,392]" inLineHeading="true" pageId="5" pageNumber="269" reason="1">Selective osteology.</emphasis>
|
||
(
|
||
<figureCitation id="9EA42A1EFFA9FF8D35F2FE967EBEFE14" box="[440,508,367,391]" captionStart="FIGURE 3" captionStartId="4.[151,250,1706,1729]" captionTargetBox="[258,1316,177,1650]" captionTargetId="figure-243@4.[258,1329,177,1689]" captionTargetPageId="4" captionText="FIGURE 3. Selected osteological features of Metzia bounthobi, new species. Features from one paratype (55.1 mm SL) of NUOL-P 4902: (A) lateral view of jaws, suspensorium and opercular bones; (B) lateral view of infraorbital bones; (C) mesial (a) and anterior (b) views of fifth ceratobranchial; (D) lateral view of anterior part of axial skeletal complex; (E) lateral view of caudal skeletal complex; (F) dorsal fin and its supports; (G) anal fin and its supports. Bars indicate 2 mm. Abbreviations: AN, anguloarticular; C, claustrum; CE, centrum; CIHPU, interhaemal spine cartilage; DEN, dentary; DR, distal radial; ECPT, ectopterygoid; ENPT, endopterygoid; EP, epural; FR, fin ray; H, hypural; HY, hyomandibular; IO, infraorbital; IOP, interopercle; IOS, inner arm of os suspensorium; L, lateral process of vertebral centrum; MPT, metapterygoid; MR, medial radial; MX, maxilla; NS, neural spine; OP, opercle; OPC, opisthural cartilage; OSS, outer arm of os suspensorium; PAL, palatine; PH, parhypural; PLS, pleurostyle; PMX, premaxilla; POP, preopercle; PR, proximal radial; Q, quadrate; R, rib; RA, retroarticular; S, scaphium; SN, supraneural; SOP, subopercle; ST, stay; SY, symplectic; T, tripus; UN, uroneural. Photographed and retouched by K. Phongsa and K. Shibukawa, respectively." pageId="5" pageNumber="269">Fig. 3</figureCitation>
|
||
). Frontal transversely convex; lateral ethmoid bearing a truncate wing with pointed ventral corner laterally; vomer extending anteriorly slightly beyond ethmoid; parasphenoid laterally compressed at interorbital region, followed by a narrow, outwardly curved ascending process; elongate foramen between parasphenoid and basisphenoid; supraorbital reduced and narrow, not attached with infraorbital series; third infraorbital with broad posteroventral lamella; ventral margin of coracoids feebly serrate; two rows of pharyngeal teeth on fifth ceratobranchial, pattern 4, 5–4, 4 (
|
||
<specimenCount id="1099FD12FFA9FF8D36FAFDDD7C70FDAF" box="[688,818,548,572]" pageId="5" pageNumber="269" type="generic">1 specimen</specimenCount>
|
||
), 4, 4–5, 4 (
|
||
<specimenCount id="1099FD12FFA9FF8D37FDFDDD7B74FDAF" box="[951,1078,548,572]" pageId="5" pageNumber="269" type="generic">1 specimen</specimenCount>
|
||
) or 3, 4–5, 4 (
|
||
<specimenCount id="1099FD12FFA9FF8D309CFDDD7A15FDAF" box="[1238,1367,548,572]" pageId="5" pageNumber="269" type="generic">1 specimen</specimenCount>
|
||
), with pointed and slightly curved tips. Vertebrae 18+21 = 39* (8) or 18+22 = 40 (2); anteriormost 4* (8) vertebrae modified into the Weberian apparatus.
|
||
</paragraph>
|
||
<paragraph id="0620369BFFA9FF8D348DFD767B53FD5F" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="269">
|
||
<emphasis id="34EBEA89FFA9FF8D348DFD767EAFFD3B" bold="true" box="[199,493,655,680]" pageId="5" pageNumber="269">Freshly-fixed coloration.</emphasis>
|
||
Coloration of freshly-fixed specimen shown in
|
||
<figureCitation id="9EA42A1EFFA9FF8D3040FD767B28FD34" box="[1034,1130,655,679]" captionStart="FIGURE 1" captionStartId="2.[151,250,1380,1403]" captionTargetBox="[151,1436,467,1364]" captionTargetId="figure-458@2.[151,1436,451,1364]" captionTargetPageId="2" captionText="FIGURE 1. Freshly collected specimen of Meztia bounthobi (holotype, 57.0 mm SL, NIFI 4680), Mekong Basin in Sopvan, Nyoi District, Luang Prabang Province, Lao PDR (20°36.5’N, 102°39.1’E), 23 June 2010. Same specimen above and below but on different backgrounds to provide full view of characters of the species. Photographed by T. Phommavong." pageId="5" pageNumber="269">Figure 1</figureCitation>
|
||
. Dorsum of head and body dull greenish yellow, turning to plain silvery ventrally; iris silvery; fins hyaline.
|
||
</paragraph>
|
||
<paragraph id="0620369BFFA9FF8D348DFD2E7CE1FCCF" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="269">
|
||
<emphasis id="34EBEA89FFA9FF8D348DFD2E7E8EFD63" bold="true" box="[199,460,727,752]" inLineHeading="true" pageId="5" pageNumber="269" reason="1">Coloration in alcohol.</emphasis>
|
||
Coloration after preservation in alcohol shown in
|
||
<figureCitation id="9EA42A1EFFA9FF8D3043FD2E7B29FD7C" box="[1033,1131,727,751]" captionStart="FIGURE 2" captionStartId="3.[151,250,979,1002]" captionTargetBox="[151,1436,193,958]" captionTargetId="figure-530@3.[151,1436,193,958]" captionTargetPageId="3" captionText="FIGURE 2. Preserved specimen of Meztia bounthobi, new species (holotype, 57.0 mm SL, NIFI 4680). (A) lateral, (B) dorsal and (C) ventral views. Photographed by K. Phongsa." pageId="5" pageNumber="269">Figure 2</figureCitation>
|
||
. Ground color of head and body pale brown, with a dusky midlateral stripe from slightly behind posterodorsal end of operculum to caudal-fin base; midlateral stripe usually paler in specimens less than ca.
|
||
<quantity id="C1679B7EFFA9FF8D3713FCE67CEAFCA4" box="[857,936,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="269" unit="mm" value="40.0">40 mm</quantity>
|
||
SL; opercular and infraorbital areas silvery; occipital area blackish brown in some specimens; fins subtranslucent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E856510FFA9FF8D348DFC9E7D0FFC54" pageId="5" pageNumber="269" type="distribution">
|
||
<paragraph id="0620369BFFA9FF8D348DFC9E7D0FFC54" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="269">
|
||
<emphasis id="34EBEA89FFA9FF8D348DFC9E7EB2FC13" bold="true" box="[199,496,871,896]" inLineHeading="true" pageId="5" pageNumber="269" reason="1">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="C19F4D18FFA9FF8D35BCFC917DF8FC13" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[502,698,872,896]" class="Actinopterygii" family="Cyprinidae" genus="Metzia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="269" phylum="Chordata" rank="species" species="bounthobi">
|
||
<emphasis id="34EBEA89FFA9FF8D35BCFC917DF8FC13" box="[502,698,872,896]" italics="true" pageId="5" pageNumber="269">Metzia bounthobi</emphasis>
|
||
</taxonomicName>
|
||
is hitherto known only from the Ou River basin (a tributary of the Mekong) in
|
||
<collectingRegion id="C45BF879FFA9FF8D356DFC757ED9FC37" box="[295,411,908,932]" country="Laos" name="Phongsali" pageId="5" pageNumber="269">Phongsaly</collectingRegion>
|
||
and
|
||
<collectingRegion id="C45BF879FFA9FF8D3592FC757DC7FC37" box="[472,645,908,932]" country="Laos" name="Louangphabang" pageId="5" pageNumber="269">Luang Prabang</collectingRegion>
|
||
provinces of northern
|
||
<collectingCountry id="7E88760BFFA9FF8D37DBFC757C89FC30" box="[913,971,908,931]" name="Laos" pageId="5" pageNumber="269">Laos</collectingCountry>
|
||
. This species was found in upland clear streams with moderate or slow current.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E856510FFA9FF8D348DFC2D7CDCFBA7" pageId="5" pageNumber="269" type="etymology">
|
||
<paragraph id="0620369BFFA9FF8D348DFC2D7CDCFBA7" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="269">
|
||
<emphasis id="34EBEA89FFA9FF8D348DFC2D7E0FFC7F" bold="true" box="[199,333,980,1004]" inLineHeading="true" pageId="5" pageNumber="269" reason="1">Etymology.</emphasis>
|
||
The specific name,
|
||
<taxonomicName id="C19F4D18FFA9FF8D3672FC2D7DEBFC7F" authorityName="Shibukawa & Phousavanh & Phongsa & Iwata" authorityYear="2012" box="[568,681,980,1004]" class="Actinopterygii" family="Cyprinidae" genus="Metzia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="269" phylum="Chordata" rank="species" species="bounthobi">
|
||
<emphasis id="34EBEA89FFA9FF8D3672FC2D7DEBFC7F" box="[568,681,980,1004]" italics="true" pageId="5" pageNumber="269">bounthobi</emphasis>
|
||
</taxonomicName>
|
||
, is in honour of Bounthob Praxaysombath (NUOL), who was the leading researcher performing the field surveys throughout the NUOL-NEF project on 2007–2010. Eight of all
|
||
<specimenCount id="1099FD12FFA9FF8D31CAFC0E7F85FBA7" pageId="5" pageNumber="269" type="generic" typeStatus="type">10 type</specimenCount>
|
||
series of the new species were brought from the project surveys.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |