treatments-xml/data/4D/A5/07/4DA507BB25DC586EBECC8F221D501CFC.xml
2024-06-21 12:35:58 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1081.75927" ID-GBIF-Dataset="6ebea377-f492-43a0-b87f-eea36d326201" ID-PMC="PMC8782822" ID-Pensoft-Pub="1313-2970-1081-127" ID-Pensoft-UUID="A101072FE2F95283985CF69DC922CED0" ID-PubMed="35087299" ID-ZooBank="AD2D54F556D14DB2B7F8F3B3E0528394" ModsDocID="1313-2970-1081-127" checkinTime="1642207361831" checkinUser="pensoft" docAuthor="Yoshitomi, Hiroyuki &amp; Pham, Thai Hong" docDate="2022" docId="4DA507BB25DC586EBECC8F221D501CFC" docLanguage="en" docName="ZooKeys 1081: 127-136" docOrigin="ZooKeys 1081" docPubDate="2022-01-14" docSource="http://dx.doi.org/10.3897/zookeys.1081.75927" docTitle="Cacodaemon vietnamensis Yoshitomi &amp; Pham 2022, sp. nov." docType="treatment" docUuid="70704EC2-3AE7-4406-9359-47E2A6898E29" docUuidSource="ZooBank" docVersion="4" id="A101072FE2F95283985CF69DC922CED0" lastPageNumber="127" masterDocId="A101072FE2F95283985CF69DC922CED0" masterDocTitle="The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam" masterLastPageNumber="136" masterPageNumber="127" pageNumber="127" updateTime="1668151305564" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshitomi, Hiroyuki</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8416-9249</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan</mods:affiliation>
<mods:nameIdentifier type="email">hymushi@agr.ehime-u.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Thai Hong</mods:namePart>
<mods:affiliation>Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1081</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>127</mods:start>
<mods:end>136</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1081.75927</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1081.75927</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1081-127</mods:identifier>
<mods:identifier type="ZooBank">AD2D54F556D14DB2B7F8F3B3E0528394</mods:identifier>
<mods:identifier type="Pensoft-UUID">A101072FE2F95283985CF69DC922CED0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="192143026" LSID="urn:lsid:zoobank.org:act:70704EC2-3AE7-4406-9359-47E2A6898E29" httpUri="http://treatment.plazi.org/id/4DA507BB25DC586EBECC8F221D501CFC" lastPageNumber="127" pageId="0" pageNumber="127">
<subSubSection pageId="0" pageNumber="127" type="nomenclature">
<paragraph pageId="0" pageNumber="127">
<taxonomicName LSID="http://zoobank.org/70704EC2-3AE7-4406-9359-47E2A6898E29" authority="Yoshitomi &amp; Pham, 2022" authorityName="Yoshitomi &amp; Pham" authorityYear="2022" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon vietnamensis" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">Cacodaemon vietnamensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="127">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">Figs 1E</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">, 2E</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">, 3E</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">, 5C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">, 5D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="materials_examined">
<paragraph pageId="0" pageNumber="127">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="127">
<materialsCitation collectingDate="2007-04-29" country="Vietnam" location="Bao Loc" municipality="Bao Lac" specimenCount="1" stateProvince="Lam Dong Province" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">
<typeStatus>Holotype</typeStatus>
</emphasis>
, female (EUMJ), &quot;Bao Lac
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.7" unit="km" value="27.0">27 km</quantity>
S.
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
<collectingDate value="2007-04-29">29-IV-2007</collectingDate>
Y. YOKOI leg.&quot;, [&quot;
<collectingMunicipality>Bao Lac</collectingMunicipality>
&quot; is probably misspelling of &quot;
<location LSID="urn:lsid:plazi:treatment:4DA507BB25DC586EBECC8F221D501CFC:24BFD3FAEEB26313C73C2C5EC7FB9A68" country="Vietnam" municipality="Bao Lac" name="Bao Loc" stateProvince="Lam Dong Province">Bao Loc</location>
&quot;
<collectingRegion country="Vietnam" name="Lam Dong">Lam Dong Province</collectingRegion>
, S.
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="diagnosis">
<paragraph pageId="0" pageNumber="127">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="127">
This species is similar to
<taxonomicName authority="laotinus" lsidName="C. laotinus subsp. laotinus" pageId="0" pageNumber="127" rank="subSpecies" species="laotinus" subSpecies="laotinus">
<emphasis italics="true" pageId="0" pageNumber="127">C. laotinus laotinus</emphasis>
</taxonomicName>
in having conoidal DA and dull orange elytral appendages but differs from it by the following characteristics: TL smaller, PA consisting of two pairs of small tubercles.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">Description.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Female.</emphasis>
Body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">1E</figureCitation>
) oval, convex dorsally, weakly shiny. Coloration of body black; BA, DA, PA, antennomere 1 and anterolateral corners of pronotum dull orange.
</paragraph>
<paragraph pageId="0" pageNumber="127">
Head finely punctate, closely covered with short setae. Antennae long and slender; antennomere 3 longest, a little shorter than 4 and combined; club (antennomeres 9-11) distinctly wide. Pronotum closely punctate, microreticulate, widest at apical 1/3, basal and lateral grooves deep and distinct; front corners projecting and pointed; lateral margins straight, slightly tapered posteriorly from apical 1/3; posterior corners right-angled; PLM/PLS 0.79; PWM/PWA 1.24; PWM/PLM 1.53; PWM/PLS 1.21. Fore tibia (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">2E</figureCitation>
) straight and long, relatively slender. Elytra (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">3E</figureCitation>
) coarsely and closely covered with shallow punctures, microreticulate; BA tubercle; HA long spine, stout in basal parts, projecting laterally; DA projecting dorsally, conoidal, with a small spine, stout in basal part; PA consisting of two pairs of small tubercles; apex of elytra with short spines; EL/EWH 0.85; EL/PLM 3.18; EWH/PWM 2.43; EWH/EWM 1.62; TL/EWH 1.12.
</paragraph>
<caption doi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 3.</emphasis>
Elytral appendages of
<taxonomicName authorityName="Thomson" authorityYear="1857" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="127">Cacodaemon</emphasis>
</taxonomicName>
spp. in lateral view
<emphasis bold="true" pageId="0" pageNumber="127">A-D</emphasis>
<taxonomicName authority="laotinus" lsidName="C. laotinus subsp. laotinus" pageId="0" pageNumber="127" rank="subSpecies" species="laotinus" subSpecies="laotinus">
<emphasis italics="true" pageId="0" pageNumber="127">C. laotinus laotinus</emphasis>
</taxonomicName>
, Vietnamese specimens (
<emphasis bold="true" pageId="0" pageNumber="127">A, B</emphasis>
) and Laotian specimens (
<emphasis bold="true" pageId="0" pageNumber="127">C, D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="127">E</emphasis>
<taxonomicName lsidName="C. vietnamensis" pageId="0" pageNumber="127" rank="species" species="vietnamensis">
<emphasis italics="true" pageId="0" pageNumber="127">C. vietnamensis</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="127">F</emphasis>
<taxonomicName lsidName="C. proavus" pageId="0" pageNumber="127" rank="species" species="proavus">
<emphasis italics="true" pageId="0" pageNumber="127">C. proavus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="127">A, C</emphasis>
male
<emphasis bold="true" pageId="0" pageNumber="127">B, D-F</emphasis>
female. Scale: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
Sternite VIII (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">5C</figureCitation>
) with tuft of short setae in postero-lateral margins of coxites; posterior margin of fused coxites gently arcuate; styli bearing short setae.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Measurement.</emphasis>
Female (
<emphasis italics="true" pageId="0" pageNumber="127">N</emphasis>
= 1). TL 7.10 mm; PWM 2.60 mm; PWA 2.10 mm; PLM 1.70 mm; PLS 2.15 mm; EL 5.40 mm; EWH 6.33 mm; EWM 3.90 mm.
</paragraph>
<caption doi="10.3897/zookeys.1081.75927.figure4" httpUri="https://binary.pensoft.net/fig/634066" pageId="0" pageNumber="127" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 4.</emphasis>
Aedeagus of
<taxonomicName authority="laotinus" authorityName="laotinus (Arrow" authorityYear="1920" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon laotinus subsp. laotinus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="laotinus" subSpecies="laotinus">
<emphasis italics="true" pageId="0" pageNumber="127">Cacodaemon laotinus laotinus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="127">A-D</emphasis>
vietnamese specimen
<emphasis bold="true" pageId="0" pageNumber="127">E-H</emphasis>
laotian specimen.
<emphasis bold="true" pageId="0" pageNumber="127">A, E</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="127">B, F</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="127">C, G</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="127">D, H</emphasis>
apical. Abbreviations ab: apical branch; sb: subapical branch. Scales: 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="distribution">
<paragraph pageId="0" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="127">Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="etymology">
<paragraph pageId="0" pageNumber="127">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="127">Named after the type locality.</paragraph>
</subSubSection>
</treatment>
</document>