treatments-xml/data/4F/43/85/4F43853E841CFFFBFF5D238445D5D4C8.xml

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<mods:title id="058B8595B22790861D55FD8EC638E2B1">Trachelostenini sensu novo: redescriptions of Trachelostenus Solier, Myrmecodema Gebien and Leaus Matthews &amp; Lawrence, based on adults and larvae, and descriptions of three new species of Leaus (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="171245F047292C40A595C1AC1973E510">Matthews, Eric G.</mods:namePart>
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<mods:namePart id="CBA7A0F5DE3DAAAFBD306B3FEE5854F4">Lawrence, John F.</mods:namePart>
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<paragraph id="C7553428841CFFF9FF5D23844593D3FC" blockId="5.[151,700,400,459]" box="[151,487,400,426]" pageId="5" pageNumber="294">
<heading id="9C1D8344841CFFF9FF5D23844593D3FC" bold="true" box="[151,487,400,426]" fontSize="11" level="1" pageId="5" pageNumber="294" reason="1">
<taxonomicName id="00EA4FAB841CFFF9FF5D23844593D3FC" authority="Solier, 1851" authorityName="Solier" authorityYear="1851" box="[151,487,400,426]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841CFFF9FF5D23844593D3FC" bold="true" box="[151,487,400,426]" pageId="5" pageNumber="294">
<emphasis id="F59EE83A841CFFF9FF5D23844524D3FC" bold="true" box="[151,336,400,426]" italics="true" pageId="5" pageNumber="294">Trachelostenus</emphasis>
<bibRefCitation id="A37B49D9841CFFF9FE9C23844593D3FC" author="Solier" box="[342,487,400,426]" pageId="5" pageNumber="294" refString="Solier, A. J. J. (1851) IV. Traquelosteno- Trachelostenus. In: Gay, C. (Ed.), Historia fisica y politica de Chile, 5 (Zoologia). pp. 255 - 256. [Author, Paris]" type="journal article" year="1851">Solier, 1851</bibRefCitation>
</emphasis>
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<paragraph id="C7553428841CFFF9FF5D23A646C8D39D" blockId="5.[151,700,400,459]" box="[151,700,434,459]" pageId="5" pageNumber="294">
(
<figureCitation id="5FD128AD841CFFF9FF5523A64491D39D" box="[159,229,434,459]" captionStart="FIGURES 1, 2" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[215,1369,205,1911]" captionTargetId="figure@6.[194,1393,193,1932]" captionTargetPageId="6" captionText="FIGURES 1, 2. Female tracts: 1, Trachelostenus inaequalis; 2, Myrmecodema michelbacheri. sp—spermatheca; vs—vaginal sclerite. Scale bars 1 mm." httpUri="https://zenodo.org/record/236476/files/figure.png" pageId="5" pageNumber="294">Figs 1</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FF3923A74577D39D" box="[243,259,435,459]" captionStart="FIGURES 3 – 5" captionStartId="7.[151,264,1000,1023]" captionTargetBox="[163,1422,196,964]" captionTargetId="figure@7.[151,1436,193,979]" captionTargetPageId="7" captionText="FIGURES 3 – 5. Aedeagi: 3, Trachelostenus inaequalis with penis extracted; 4, Myrmecodema michelbacheri; 5, Leaus monteithi. a—ala of paramere. Scale bars 1 mm." httpUri="https://zenodo.org/record/236477/files/figure.png" pageId="5" pageNumber="294">3</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FEC523A6456BD39D" box="[271,287,434,459]" captionStart="FIGURES 6, 7" captionStartId="8.[151,264,1772,1795]" captionTargetBox="[166,1415,233,1730]" captionTargetId="figure@8.[151,1436,193,1751]" captionTargetPageId="8" captionText="FIGURES 6, 7. Ovipositors: 6, Trachelostenus inaequalis; 7, Myrmecodema michelbacheri. cb—baculus of coxite 1; pbbaculus of paraproct. Scale bars 1 mm." httpUri="https://zenodo.org/record/236478/files/figure.png" pageId="5" pageNumber="294">6</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FEE123A7454FD39D" box="[299,315,435,459]" captionStart="FIGURES 8 – 10" captionStartId="9.[151,264,882,905]" captionTargetBox="[166,1421,201,849]" captionTargetId="figure@9.[151,1436,193,861]" captionTargetPageId="9" captionText="FIGURES 8 – 10. Inside of abdominal ventrites. 8, Trachelostenus inaequalis; 9, Myrmecodema michelbacheri; 10, Leaus tropicalis. Scale bars 1 mm." httpUri="https://zenodo.org/record/236479/files/figure.png" pageId="5" pageNumber="294">8</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FE8D23A74510D39D" box="[327,356,435,459]" captionStart="FIGURE 11" captionStartId="9.[151,250,1607,1629]" captionTargetBox="[163,1410,1002,1566]" captionTargetId="figure@9.[151,1435,985,1586]" captionTargetPageId="9" captionText="FIGURE 11. Traschelostenus inaequalis, tergite VII showing striated patches (s) in both sexes. Scale bar 1 mm." httpUri="https://zenodo.org/record/236480/files/figure.png" pageId="5" pageNumber="294">11</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FEBA23A645CDD39D" box="[368,441,434,459]" captionStart-0="FIGURES 12 – 14" captionStart-1="FIGURES 15, 16" captionStartId-0="10.[151,264,1470,1493]" captionStartId-1="11.[151,264,1650,1673]" captionTargetBox-0="[169,1419,337,1430]" captionTargetBox-1="[170,1400,222,1614]" captionTargetId-0="figure@10.[151,1436,315,1450]" captionTargetId-1="figure@11.[151,1436,193,1629]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 12 – 14. Trachelostenus inaequalis ♂♂: 12, dorsal; 13, ¾ ventral; 14, ventral. Scale bars 1 mm." captionText-1="FIGURES 15, 16. Trachelostenus fasciculiferus ♂: 15, dorsal; 16, lateral. Scale bars 1 mm." httpUri-0="https://zenodo.org/record/236481/files/figure.png" httpUri-1="https://zenodo.org/record/236482/files/figure.png" pageId="5" pageNumber="294">12–16</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FE0E23A74678D39D" box="[452,524,435,459]" captionStart="FIGURES 37 – 41" captionStartId="18.[151,264,1953,1976]" captionTargetBox="[209,1374,227,1913]" captionTargetId="figure@18.[187,1400,193,1932]" captionTargetPageId="18" captionText="FIGURES 37 – 41. Trachelostenine larvae: 37 – 39, head and prothorax, dorsal; 37, Myrmecodema nycterinoides; 38, Trachelostenus inaequalis; 39, Leaus tasmanicus; 40, L. tasmanicus, larval head, dorsal; 41, T. inaequalis, larval head, lateral." httpUri="https://zenodo.org/record/236488/files/figure.png" pageId="5" pageNumber="294">38, 41</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FDD223A74614D39D" box="[536,608,434,459]" captionStart="FIGURES 42 – 48" captionStartId="19.[151,264,1938,1961]" captionTargetBox="[177,1409,217,1864]" captionTargetId="figure@19.[151,1436,193,1917]" captionTargetPageId="19" captionText="FIGURES 42 – 48. Trachelostenine larvae: 42 – 44: abdominal apex, lateral: 42, Trachelostenus inaequalis; 43, Myrmecodema nycterinoides, 44, Leaus tasmanicus; 45 – 48: abdominal apex, dorsal: 45, Trachelostenus inaequalis; 46, Myrmecodema nycterinoides; 47 – 48, Leaus tasmanicus." httpUri="https://zenodo.org/record/236489/files/figure.png" pageId="5" pageNumber="294">42, 45</figureCitation>
, 
<figureCitation id="5FD128AD841CFFF9FDA623A746C0D39D" box="[620,692,434,459]" captionStart="FIGURES 49 – 56" captionStartId="20.[151,264,1907,1930]" captionTargetBox="[197,1365,225,1842]" captionTargetId="figure@20.[175,1402,193,1865]" captionTargetPageId="20" captionText="FIGURES 49 – 56. Trachelostenine larvae: 49, Trachelostenus inaequalis, ventral mouthparts and hypopharynx, dorsal. 50 – 51, Leaus tasmanicus, mandibles: 50, right, dorsal; 51, left, dorsal. 52 – 54: labrum-epipharynx, ventral: 52, Trachelostenus inaequalis, 53, Myrmecodema nycterinoides, 54, Leaus tasmanicus. 55 – 56, abdominal spiracle: 55, Myrmecodema nycterinoides. 56, Leaus tasmanicus." httpUri="https://zenodo.org/record/236490/files/figure.png" pageId="5" pageNumber="294">49, 52</figureCitation>
)
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<paragraph id="C7553428841CFFF9FF5D23EE47E5D045" blockId="5.[151,1437,506,2007]" box="[151,913,506,531]" pageId="5" pageNumber="294">
<emphasis id="F59EE83A841CFFF9FF5D23EE4540D045" bold="true" box="[151,308,506,531]" pageId="5" pageNumber="294">
<typeStatus id="18518A8A841CFFF9FF5D23EE44A6D045" box="[151,210,506,531]" pageId="5" pageNumber="294">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="00EA4FAB841CFFF9FEF123EE469BD045" authority="Solier, 1851" authorityName="Solier" authorityYear="1851" box="[315,751,506,531]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841CFFF9FEF123EE4615D045" box="[315,609,506,531]" italics="true" pageId="5" pageNumber="294">Trachelostenus inaequalis</emphasis>
<bibRefCitation id="A37B49D9841CFFF9FDAD23EE469BD045" author="Solier" box="[615,751,506,531]" pageId="5" pageNumber="294" refString="Solier, A. J. J. (1851) IV. Traquelosteno- Trachelostenus. In: Gay, C. (Ed.), Historia fisica y politica de Chile, 5 (Zoologia). pp. 255 - 256. [Author, Paris]" type="journal article" year="1851">Solier, 1851</bibRefCitation>
</taxonomicName>
by monotypy.
</paragraph>
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<subSubSection id="8FF067A3841CFFF9FF0D200A4525D751" pageId="5" pageNumber="294" type="description">
<paragraph id="C7553428841CFFF9FF0D200A4525D751" blockId="5.[151,1437,506,2007]" pageId="5" pageNumber="294">
<emphasis id="F59EE83A841CFFF9FF0D200A4519D061" bold="true" box="[199,365,542,567]" pageId="5" pageNumber="294">Redescription</emphasis>
. 
<emphasis id="F59EE83A841CFFF9FEB7200A45B8D061" bold="true" box="[381,460,542,567]" pageId="5" pageNumber="294">Adults</emphasis>
. With the characteristics of the tribe. Length 
<quantity id="001299CD841CFFF9FC1D200A4005D060" box="[983,1137,542,567]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.5" metricValueMin="6.5" pageId="5" pageNumber="294" unit="mm" value="8.5" valueMax="10.5" valueMin="6.5">6.5–10.5 mm</quantity>
. Body parallel-sided, 3.3– 3.6 times as long as wide, with prothorax narrower than elytral bases, fuscous, legs with alternating flavous and fuscous bands. Upper surfaces glabrous or clothed with patches of semierect white hairs, or sparse semirecumbent pilosity. 
<emphasis id="F59EE83A841CFFF9FF3C209E4546D0F5" box="[246,306,650,675]" italics="true" pageId="5" pageNumber="294">Head</emphasis>
: Basal membrane of labrum moderately wide. Eyes shallowly emarginate anteriorly. Frons with deep median pit. Antennae very long, filiform, reaching to about anterior third of elytral length, antennomeres elongate, terminal one oblong, not enlarged. Mentum transverse, about twice as wide as long. 
<emphasis id="F59EE83A841CFFF9FB8520C640B5D0BD" box="[1103,1217,722,747]" italics="true" pageId="5" pageNumber="294">Prothorax</emphasis>
: Pronotum oblong, sides subparallel, almost straight or a little convex, lateral carinae absent. Procoxal cavities internally closed by very slender bar. Concealed propleurotrochantins present. 
<emphasis id="F59EE83A841CFFF9FC84210E47A0D165" box="[846,980,794,819]" italics="true" pageId="5" pageNumber="294">Pterothorax</emphasis>
: Elytral disc with elevations, at least anteriorly. Wings fully developed. 
<emphasis id="F59EE83A841CFFF9FDD5212B4620D100" box="[543,596,831,854]" italics="true" pageId="5" pageNumber="294">Legs</emphasis>
: Pro- and mesocoxae subconical and prominent. Procoxae projecting well below prosternum, approximated so as to cover latter. Mesocoxae prominent and narrowly separated. Femora and tibiae long and slender, tarsomeres linear except penultimate, which is slightly widened, together as long as or longer than tibiae. 
<emphasis id="F59EE83A841CFFF9FEB921BE45A9D195" box="[371,477,938,963]" italics="true" pageId="5" pageNumber="294">Abdomen</emphasis>
: Intercoxal process of first ventrite narrow, subparallel or triangular, with rounded apex. With only first 2 ventrites connate. Tergite VII with a pair of finely rugose patches in both sexes. Defensive glands absent. 
<emphasis id="F59EE83A841CFFF9FEF621E64501D65D" box="[316,373,1010,1035]" italics="true" pageId="5" pageNumber="294">Male</emphasis>
: In 
<taxonomicName id="00EA4FAB841CFFF9FE6921E04640D65D" box="[419,564,1010,1035]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841CFFF9FE6921E04640D65D" box="[419,564,1010,1035]" italics="true" pageId="5" pageNumber="294">T. inaequalis</emphasis>
</taxonomicName>
with a deep longitudinal groove joining frontal pit with fronto-clypeal suture, not present in other species. Tergite VII a little longer than wide. Sternite VIII arcuately excised, in one unnamed species with outer angles of excision prolonged into short spurs. Sternite IX partly or completely divided into 2 sclerites, 
<emphasis id="F59EE83A841CFFF9FECC264A452ED621" box="[262,346,1118,1143]" italics="true" pageId="5" pageNumber="294">Female</emphasis>
: Tergite VII about as long as wide. Ovipositor with paraproct baculi subparallel to axis of ovipositor, baculi of first coxite lobes also parallel, about 1.5 times as long as paraproct baculi. Coxites with only 3 pairs of lobes, last 2 weakly sclerotised, rest of ovipositor membranous except for baculi. Vaginal sclerites (of 
<taxonomicName id="00EA4FAB841CFFF9FA4D26B34578D6B5" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841CFFF9FA4D26B34578D6B5" italics="true" pageId="5" pageNumber="294">T. inaequalis</emphasis>
</taxonomicName>
) in form of 4 coarsely-toothed saw blades. Spermatheca a tightly coiled tube attached to basal part of accessory gland.
</paragraph>
</subSubSection>
<subSubSection id="8FF067A3841CFFF9FF0D2706406FD7ED" pageId="5" pageNumber="294" type="materials_examined">
<paragraph id="C7553428841CFFF9FF0D2706406FD7ED" blockId="5.[151,1437,506,2007]" pageId="5" pageNumber="294">
<emphasis id="F59EE83A841CFFF9FF0D2706458CD77D" bold="true" box="[199,504,1298,1323]" pageId="5" pageNumber="294">Adult material examined</emphasis>
: 
<taxonomicName id="00EA4FAB841CFFF9FDC0270046E9D77D" box="[522,669,1298,1323]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841CFFF9FDC0270046E9D77D" box="[522,669,1298,1323]" italics="true" pageId="5" pageNumber="294">T. inaequalis</emphasis>
</taxonomicName>
. 
<collectingCountry id="BFFD74B8841CFFF9FD6527074772D77D" box="[687,774,1299,1323]" name="Chile" pageId="5" pageNumber="294">CHILE</collectingCountry>
: Cudico Villarrica, 
<date id="B35412E8841CFFF9FC202707402ED77D" box="[1002,1114,1299,1323]" pageId="5" pageNumber="294" value="1978-01-27">27-I-1978</date>
(
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) (1); Osorno Prov. Parque nac. Puyehue, 
<quantity id="001299CD841CFFF9FE5027234598D719" box="[410,492,1334,1359]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="294" unit="km" value="4.1">4.1 km</quantity>
E Anticura, 
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Site 662, 
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, Valdivian 
<collectingCountry id="BFFD74B8841CFFF9FB5D272340CED718" box="[1175,1210,1335,1358]" name="Russia" pageId="5" pageNumber="294">RF</collectingCountry>
, Newton &amp; Thayer (
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) (1); 
<quantity id="001299CD841CFFF9FEFB274F450CD725" box="[305,376,1370,1395]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="294" unit="km" value="30.0">30 km</quantity>
E of Puyehue, Osorno (
<collectionCode id="A1FBACED841CFFF9FD4D274F46B4D725" LSID="urn:lsid:biocol.org:col:34700" box="[647,704,1371,1395]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="5" pageNumber="294">CAS</collectionCode>
) (1); Arauco Caramavida (
<collectionCode id="A1FBACED841CFFF9FC25274F4030D725" LSID="urn:lsid:biocol.org:col:34795" box="[1007,1092,1371,1395]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="294">FMNH</collectionCode>
) (1); Chaitén Aysén, 
<date id="B35412E8841CFFF9FAFE274E41E8D725" box="[1332,1436,1370,1395]" pageId="5" pageNumber="294" value="1985-12">XII-1985</date>
(
<collectionCode id="A1FBACED841CFFF9FF54276B4486D7C1" LSID="urn:lsid:biocol.org:col:34795" box="[158,242,1407,1431]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="294">FMNH</collectionCode>
) (3); Cherquenco [abdomen only] (
<collectionCode id="A1FBACED841CFFF9FD4A276B46A3D7C1" LSID="urn:lsid:biocol.org:col:34795" box="[640,727,1407,1431]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="294">FMNH</collectionCode>
). 
<taxonomicName id="00EA4FAB841CFFF9FD23276B47EFD7C1" box="[745,923,1406,1431]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="fasciculiferus">
<emphasis id="F59EE83A841CFFF9FD23276B47EFD7C1" box="[745,923,1406,1431]" italics="true" pageId="5" pageNumber="294">T. fasciculiferus</emphasis>
</taxonomicName>
. La Selva 
<quantity id="001299CD841CFFF9FBD9276B4027D7C1" box="[1043,1107,1407,1431]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="294" unit="m" value="700.0">700m</quantity>
Nueva Imperial, 11-81, Coll. L. E. Peña (
<collectionCode id="A1FBACED841CFFF9FED027B7451AD7ED" LSID="urn:lsid:biocol.org:col:34795" box="[282,366,1443,1467]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="294">FMNH</collectionCode>
) (3). One unnamed species. Tepuhuelco Chiloe (
<collectionCode id="A1FBACED841CFFF9FC5A27B74790D7ED" LSID="urn:lsid:biocol.org:col:34795" box="[912,996,1443,1467]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="294">FMNH</collectionCode>
) (1).
</paragraph>
</subSubSection>
<subSubSection id="8FF067A3841CFFF9FF0D27D245D9D511" pageId="5" pageNumber="294" type="description">
<paragraph id="C7553428841CFFF9FF0D27D245D9D511" blockId="5.[151,1437,506,2007]" pageId="5" pageNumber="294">
<emphasis id="F59EE83A841CFFF9FF0D27D24519D789" bold="true" box="[199,365,1478,1503]" pageId="5" pageNumber="294">Redescription</emphasis>
. 
<emphasis id="F59EE83A841CFFF9FEB127D245A4D789" bold="true" box="[379,464,1478,1503]" pageId="5" pageNumber="294">Larvae</emphasis>
. With the characteristics of the tribe. Dorsal surfaces bicolored with a complex pattern of dark brown markings on a yellow background. 
<emphasis id="F59EE83A841CFFF9FD2227FE4751D455" box="[744,805,1514,1539]" italics="true" pageId="5" pageNumber="294">Head</emphasis>
distinctly rounded; frontal arms somewhat irregularly curved and angulate; larger anterior epipharyngeal sensilla with anterior 2 moderately widely separated and close to the posterior 4, which form a sharply angulate row (
<figureCitation id="5FD128AD841CFFF9FCD124264706D41D" box="[795,882,1586,1611]" captionStart="FIGURES 49 – 56" captionStartId="20.[151,264,1907,1930]" captionTargetBox="[197,1365,225,1842]" captionTargetId="figure@20.[175,1402,193,1865]" captionTargetPageId="20" captionText="FIGURES 49 – 56. Trachelostenine larvae: 49, Trachelostenus inaequalis, ventral mouthparts and hypopharynx, dorsal. 50 – 51, Leaus tasmanicus, mandibles: 50, right, dorsal; 51, left, dorsal. 52 – 54: labrum-epipharynx, ventral: 52, Trachelostenus inaequalis, 53, Myrmecodema nycterinoides, 54, Leaus tasmanicus. 55 – 56, abdominal spiracle: 55, Myrmecodema nycterinoides. 56, Leaus tasmanicus." httpUri="https://zenodo.org/record/236490/files/figure.png" pageId="5" pageNumber="294">Fig. 52</figureCitation>
); posterior sensilla forming an irregular cluster; mandibles bidentate with short subapical tooth on incisor edge; left mola with several weak transverse ridges; gula slightly shorter than wide; hypopharyngeal sclerome anteriorly trilobed. 
<emphasis id="F59EE83A841CFFF9FC1E246E4033D4C5" box="[980,1095,1658,1683]" italics="true" pageId="5" pageNumber="294">Prothorax</emphasis>
almost as long as wide. Legs slightly longer than thoracic width and clothed with fine hairs. 
<emphasis id="F59EE83A841CFFF9FC91248A47B1D4E1" box="[859,965,1694,1719]" italics="true" pageId="5" pageNumber="294">Abdomen</emphasis>
about 3.5 times as long as thorax; tergum IX distinctly longer and slightly wider than VIII, subcircular carina simple (
<figureCitation id="5FD128AD841CFFF9FBD024D64007D48D" box="[1050,1139,1730,1755]" captionStart="FIGURES 42 – 48" captionStartId="19.[151,264,1938,1961]" captionTargetBox="[177,1409,217,1864]" captionTargetId="figure@19.[151,1436,193,1917]" captionTargetPageId="19" captionText="FIGURES 42 – 48. Trachelostenine larvae: 42 – 44: abdominal apex, lateral: 42, Trachelostenus inaequalis; 43, Myrmecodema nycterinoides, 44, Leaus tasmanicus; 45 – 48: abdominal apex, dorsal: 45, Trachelostenus inaequalis; 46, Myrmecodema nycterinoides; 47 – 48, Leaus tasmanicus." httpUri="https://zenodo.org/record/236489/files/figure.png" pageId="5" pageNumber="294">Fig. 45</figureCitation>
); urogomphi each with 4 setiferous tubercles at about middle; surface of concave disc lightly pigmented; segment X with a pair of tubular pygopods. 
<emphasis id="F59EE83A841CFFF9FED3251E45F5D575" box="[281,385,1802,1827]" italics="true" pageId="5" pageNumber="294">Spiracles</emphasis>
annular-multiforous with a variable number of accessory chambers located around the peritreme (as in 
<figureCitation id="5FD128AD841CFFF9FE87253A45D4D511" box="[333,416,1838,1863]" captionStart="FIGURES 49 – 56" captionStartId="20.[151,264,1907,1930]" captionTargetBox="[197,1365,225,1842]" captionTargetId="figure@20.[175,1402,193,1865]" captionTargetPageId="20" captionText="FIGURES 49 – 56. Trachelostenine larvae: 49, Trachelostenus inaequalis, ventral mouthparts and hypopharynx, dorsal. 50 – 51, Leaus tasmanicus, mandibles: 50, right, dorsal; 51, left, dorsal. 52 – 54: labrum-epipharynx, ventral: 52, Trachelostenus inaequalis, 53, Myrmecodema nycterinoides, 54, Leaus tasmanicus. 55 – 56, abdominal spiracle: 55, Myrmecodema nycterinoides. 56, Leaus tasmanicus." httpUri="https://zenodo.org/record/236490/files/figure.png" pageId="5" pageNumber="294">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8FF067A3841CFFF9FF0D254640EFD581" pageId="5" pageNumber="294" type="materials_examined">
<paragraph id="C7553428841CFFF9FF0D254640EFD581" blockId="5.[151,1437,506,2007]" pageId="5" pageNumber="294">
<emphasis id="F59EE83A841CFFF9FF0D25464675D53D" bold="true" box="[199,513,1874,1899]" pageId="5" pageNumber="294">Larval material examined</emphasis>
: 
<taxonomicName id="00EA4FAB841CFFF9FDD8254046D7D53D" box="[530,675,1874,1899]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841CFFF9FDD8254046D7D53D" box="[530,675,1874,1899]" italics="true" pageId="5" pageNumber="294">T. inaequalis</emphasis>
</taxonomicName>
. 
<collectingCountry id="BFFD74B8841CFFF9FD7E2547477FD53D" box="[692,779,1875,1899]" name="Chile" pageId="5" pageNumber="294">CHILE</collectingCountry>
: VIII: Parque Nac. Nahuelbuta, Pichinahuel Exit, 
<quantity id="001299CD841CFFF9FA8D254741E3D53D" box="[1351,1431,1875,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="5" pageNumber="294" unit="m" value="1219.0">1219m</quantity>
, 
<date id="B35412E8841CFFF9FF5D2562456DD5D9" box="[151,281,1910,1935]" pageId="5" pageNumber="294" value="2001-12-08">08.xii.2001</date>
, fogging 
<taxonomicName id="00EA4FAB841CFFF9FE4E25624036D5D9" authority="E. Arias et al. U.C. Berkeley Expedition" authorityName="E. Arias et al. U.C. Berkeley Expedition" box="[388,1090,1910,1935]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="294" phylum="Tracheophyta" rank="species" species="dombeyi">
<emphasis id="F59EE83A841CFFF9FE4E2562461FD5D9" box="[388,619,1910,1935]" italics="true" pageId="5" pageNumber="294">Nothofagus dombeyi</emphasis>
, E. Arias 
<emphasis id="F59EE83A841CFFF9FD1725634762D5D9" box="[733,790,1910,1935]" italics="true" pageId="5" pageNumber="294">et al.</emphasis>
U.C. Berkeley Expedition
</taxonomicName>
(
<collectionCode id="A1FBACED841CFFF9FB9B256340EFD5D9" LSID="urn:lsid:biocol.org:col:32981" box="[1105,1179,1911,1935]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="294">ANIC</collectionCode>
); X: Osorno: 
<quantity id="001299CD841CFFF9FAFF256341F7D5D9" box="[1333,1411,1910,1935]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="294" unit="km" value="4.1">4.1 km</quantity>
E Anticura, 
<quantity id="001299CD841CFFF9FEC6258F453BD5E5" box="[268,335,1947,1971]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="294" unit="m" value="430.0">430m</quantity>
, 
<date id="B35412E8841CFFF9FE95258E467FD5E5" box="[351,523,1946,1971]" pageId="5" pageNumber="294" value="1982-12-19" valueMax="1982-12-26" valueMin="1982-12-19">19–26.xii.1982</date>
, A. Newton &amp; M. Thayer colls (
<collectionCode id="A1FBACED841CFFF9FC53258F4790D5E5" LSID="urn:lsid:biocol.org:col:32981" box="[921,996,1947,1971]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="294">ANIC</collectionCode>
). 
<taxonomicName id="00EA4FAB841CFFF9FC31258E40D1D5E5" box="[1019,1189,1946,1971]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841CFFF9FC31258E40D1D5E5" box="[1019,1189,1946,1971]" italics="true" pageId="5" pageNumber="294">Trachelostenus</emphasis>
</taxonomicName>
sp. IX: Parque Nac. Villarrica, Sector Puesco, 
<quantity id="001299CD841CFFF9FE7025AB4589D581" box="[442,509,1983,2007]" metricMagnitude="2" metricUnit="m" metricValue="8.73" pageId="5" pageNumber="294" unit="m" value="873.0">873m</quantity>
, 
<date id="B35412E8841CFFF9FDC225AA4600D581" box="[520,628,1982,2007]" pageId="5" pageNumber="294" value="2007-01-16">16.i.2007</date>
, under bark of logs, J. F. Lawrence coll. (
<collectionCode id="A1FBACED841CFFF9FB8E25AB40FBD581" LSID="urn:lsid:biocol.org:col:32981" box="[1092,1167,1983,2007]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="294">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="939564A0841FFFFAFF5D25B445E8D583" httpUri="https://zenodo.org/record/236476/files/figure.png" pageId="6" pageNumber="295" targetBox="[215,1369,205,1911]" targetPageId="6">
<paragraph id="C7553428841FFFFAFF5D25B445E8D583" blockId="6.[151,1436,1952,2005]" pageId="6" pageNumber="295">
<emphasis id="F59EE83A841FFFFAFF5D25B44548D5E0" bold="true" box="[151,316,1952,1975]" pageId="6" pageNumber="295">FIGURES 1, 2.</emphasis>
Female tracts: 1, 
<taxonomicName id="00EA4FAB841FFFFAFE3F25B5468BD5E1" box="[501,767,1953,1975]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841FFFFAFE3F25B5468BD5E1" box="[501,767,1953,1975]" italics="true" pageId="6" pageNumber="295">Trachelostenus inaequalis</emphasis>
</taxonomicName>
; 2, 
<taxonomicName id="00EA4FAB841FFFFAFCE225B54023D5E1" box="[808,1111,1953,1975]" class="Insecta" family="Tenebrionidae" genus="Myrmecodema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="michelbacheri">
<emphasis id="F59EE83A841FFFFAFCE225B54023D5E1" box="[808,1111,1953,1975]" italics="true" pageId="6" pageNumber="295">Myrmecodema michelbacheri</emphasis>
</taxonomicName>
. sp—spermatheca; vs—vaginal sclerite. Scale bars 1 mm.
</paragraph>
</caption>
<caption id="939564A0841EFFFBFF5D21FC46F5D64B" httpUri="https://zenodo.org/record/236477/files/figure.png" pageId="7" pageNumber="296" targetBox="[163,1422,196,964]" targetPageId="7">
<paragraph id="C7553428841EFFFBFF5D21FC46F5D64B" blockId="7.[151,1435,1000,1053]" pageId="7" pageNumber="296">
<emphasis id="F59EE83A841EFFFBFF5D21FC454DD1A8" bold="true" box="[151,313,1000,1023]" pageId="7" pageNumber="296">FIGURES 3–5</emphasis>
. Aedeagi: 3, 
<taxonomicName id="00EA4FAB841EFFFBFE1921FD4696D1A9" box="[467,738,1001,1023]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="F59EE83A841EFFFBFE1921FD4696D1A9" box="[467,738,1001,1023]" italics="true" pageId="7" pageNumber="296">Trachelostenus inaequalis</emphasis>
</taxonomicName>
with penis extracted; 4, 
<taxonomicName id="00EA4FAB841EFFFBFC3321FD4159D1A9" box="[1017,1325,1001,1023]" class="Insecta" family="Tenebrionidae" genus="Myrmecodema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="michelbacheri">
<emphasis id="F59EE83A841EFFFBFC3321FD4159D1A9" box="[1017,1325,1001,1023]" italics="true" pageId="7" pageNumber="296">Myrmecodema michelbacheri</emphasis>
</taxonomicName>
; 5, 
<taxonomicName id="00EA4FAB841EFFFBFA9521FE4483D64B" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="F59EE83A841EFFFBFA9521FE4483D64B" italics="true" pageId="7" pageNumber="296">Leaus monteithi</emphasis>
</taxonomicName>
. a—ala of paramere. Scale bars 1 mm.
</paragraph>
</caption>
<subSubSection id="8FF067A3841EFFFBFF0D2650454AD747" pageId="7" pageNumber="296" type="distribution">
<paragraph id="C7553428841EFFFBFF0D2650454AD747" blockId="7.[151,1437,1092,1694]" pageId="7" pageNumber="296">
<emphasis id="F59EE83A841EFFFBFF0D26504590D60B" bold="true" box="[199,484,1092,1117]" pageId="7" pageNumber="296">Distribution and habits.</emphasis>
The above localities are in three of the biogeographic regions of 
<collectingCountry id="BFFD74B8841EFFFBFB772650408DD60B" box="[1213,1273,1092,1117]" name="Chile" pageId="7" pageNumber="296">Chile</collectingCountry>
recognised by 
<bibRefCitation id="A37B49D9841EFFFBFF5D267C45B9D6D7" author="Vidal" box="[151,461,1128,1153]" pageId="7" pageNumber="296" refString="Vidal Garcia-Huidobro, P. &amp; Guerrero Guerrero, M. (2007) Los tenebrionidos de Chile. Universidad Catolica de Chile, Santiago, 478 pp." type="book" year="2007">Vidal &amp; Guerrero (2007)</bibRefCitation>
: Bosque de Valdivia del Norte, Bosque de Valdivia, and Cordillera de Aysén. 
<emphasis id="F59EE83A841EFFFBFF5D26984558D6F3" box="[151,300,1164,1189]" italics="true" pageId="7" pageNumber="296">Nothophagus</emphasis>
forest is the dominant vegetation 
<typeStatus id="18518A8A841EFFFBFD1D269A477FD6F0" box="[727,779,1166,1190]" pageId="7" pageNumber="296">type</typeStatus>
. 
<bibRefCitation id="A37B49D9841EFFFBFCD72698402FD6F3" author="Philippi" box="[797,1115,1164,1189]" pageId="7" pageNumber="296" refString="Philippi, R. A. &amp; Philippi, A. H. E. (1860) Coleoptera nonnulla nova Chilensis praesertim Valdiviana. Entomologische Zeitung, 21 (7 - 9), 248." type="journal article" year="1860">Philippi &amp; Philippi (1860)</bibRefCitation>
describe the habitat of 
<taxonomicName id="00EA4FAB841EFFFBFA4D269A455AD69F" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="fasciculiferus">
<emphasis id="F59EE83A841EFFFBFA4D269A455AD69F" italics="true" pageId="7" pageNumber="296">T. fasciculiferus</emphasis>
</taxonomicName>
as ‘ 
<emphasis id="F59EE83A841EFFFBFE9626A64744D69F" box="[348,816,1200,1225]" italics="true" pageId="7" pageNumber="296">in Valdivia sub cortice arborum rarissime</emphasis>
’ and 
<bibRefCitation id="A37B49D9841EFFFBFCA526A540C3D69F" author="Lawrence" box="[879,1207,1200,1225]" pageId="7" pageNumber="296" refString="Lawrence, J. F. &amp; Slipinski, A. (2010) 11.13. Trachelostenidae. In: Beutel, R. G. &amp; Leschen, R. A. B. (Eds.), Handbook of Zoology. A Natural History of the Phyla of the Animal Kingdom. Vol. IV. Arthropoda: Insecta. Part 38. Coleoptera, Beetles. Vol. 2. Systematics. Part 2. Walter de Gruyter, Berlin, pp. 571 - 573." type="journal article" year="2010">Lawrence &amp; Slipinski (2010)</bibRefCitation>
also note that adults are found under bark, but the variegated pigmentation of the larvae suggests external feeding, possibly on lichens as in Titaenini.
</paragraph>
</subSubSection>
<subSubSection id="8FF067A3841EFFFBFF0D270945D5D4C8" pageId="7" pageNumber="296" type="discussion">
<paragraph id="C7553428841EFFFBFF0D270945D5D4C8" blockId="7.[151,1437,1092,1694]" pageId="7" pageNumber="296">
<emphasis id="F59EE83A841EFFFBFF0D27094531D760" bold="true" box="[199,325,1309,1334]" pageId="7" pageNumber="296">Discussion</emphasis>
. Evidently 
<taxonomicName id="00EA4FAB841EFFFBFE0027094607D760" box="[458,627,1309,1334]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFE0027094607D760" box="[458,627,1309,1334]" italics="true" pageId="7" pageNumber="296">Trachelostenus</emphasis>
</taxonomicName>
has lost the ancestral defensive glands and with them the connation of the third ventrite (
<figureCitation id="5FD128AD841EFFFBFE95275445D7D70F" box="[351,419,1344,1369]" captionStart="FIGURES 8 – 10" captionStartId="9.[151,264,882,905]" captionTargetBox="[166,1421,201,849]" captionTargetId="figure@9.[151,1436,193,861]" captionTargetPageId="9" captionText="FIGURES 8 – 10. Inside of abdominal ventrites. 8, Trachelostenus inaequalis; 9, Myrmecodema michelbacheri; 10, Leaus tropicalis. Scale bars 1 mm." httpUri="https://zenodo.org/record/236479/files/figure.png" pageId="7" pageNumber="296">Fig. 8</figureCitation>
). There is a striking apomorphy which is shared with 
<taxonomicName id="00EA4FAB841EFFFBFC30275440D4D70F" box="[1018,1184,1344,1369]" class="Insecta" family="Tenebrionidae" genus="Myrmecodema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFC30275440D4D70F" box="[1018,1184,1344,1369]" italics="true" pageId="7" pageNumber="296">Myrmecodema</emphasis>
</taxonomicName>
: the unique ovipositor structure with longitudinal and elongate coxite-1 baculi (
<figureCitation id="5FD128AD841EFFFBFCDD2771470FD72B" box="[791,891,1381,1406]" captionStart="FIGURES 6, 7" captionStartId="8.[151,264,1772,1795]" captionTargetBox="[166,1415,233,1730]" captionTargetId="figure@8.[151,1436,193,1751]" captionTargetPageId="8" captionText="FIGURES 6, 7. Ovipositors: 6, Trachelostenus inaequalis; 7, Myrmecodema michelbacheri. cb—baculus of coxite 1; pbbaculus of paraproct. Scale bars 1 mm." httpUri="https://zenodo.org/record/236478/files/figure.png" pageId="7" pageNumber="296">Figs 6, 7</figureCitation>
cb), first noted by 
<bibRefCitation id="A37B49D9841EFFFBFB99277241E8D72B" author="Lawrence" box="[1107,1436,1381,1406]" pageId="7" pageNumber="296" refString="Lawrence, J. F. &amp; Slipinski, A. (2010) 11.13. Trachelostenidae. In: Beutel, R. G. &amp; Leschen, R. A. B. (Eds.), Handbook of Zoology. A Natural History of the Phyla of the Animal Kingdom. Vol. IV. Arthropoda: Insecta. Part 38. Coleoptera, Beetles. Vol. 2. Systematics. Part 2. Walter de Gruyter, Berlin, pp. 571 - 573." type="journal article" year="2010">Lawrence &amp; Slipinski (2010)</bibRefCitation>
in 
<taxonomicName id="00EA4FAB841EFFFBFF7C279C4514D7F7" box="[182,352,1416,1441]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFF7C279C4514D7F7" box="[182,352,1416,1441]" italics="true" pageId="7" pageNumber="296">Trachelostenus</emphasis>
</taxonomicName>
. Normally in 
<taxonomicName id="00EA4FAB841EFFFBFDCF279C46D1D7F7" box="[517,677,1416,1441]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
coxite-1 baculi and lobes are short and transverse or diagonal, as they are in 
<taxonomicName id="00EA4FAB841EFFFBFEEE27BA4513D793" box="[292,359,1454,1477]" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFEEE27BA4513D793" box="[292,359,1454,1477]" italics="true" pageId="7" pageNumber="296">Leaus</emphasis>
</taxonomicName>
for instance (
<figureCitation id="5FD128AD841EFFFBFDDB27B946E1D790" box="[529,661,1453,1478]" captionStart="FIGURES 24 – 26" captionStartId="14.[151,264,1622,1645]" captionTargetBox="[171,1422,216,1573]" captionTargetId="figure@14.[151,1436,193,1602]" captionTargetPageId="14" captionText="FIGURES 24 – 26. Leaus ovipositors: 24, L. tropicalis; 25, L. monteithi; 26, L. sp. cb—baculus of coxite 1; pb—baculus of paraproct. Scale bars 1 mm." httpUri="https://zenodo.org/record/236485/files/figure.png" pageId="7" pageNumber="296">Figs 24–26</figureCitation>
cb). Otherwise, longitudinal coxite-1 baculi are seen only in the distinctive ovipositor of 
<taxonomicName id="00EA4FAB841EFFFBFE7A27C44634D7BF" box="[432,576,1488,1513]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="subFamily" subFamily="Stenochiinae">Stenochiinae</taxonomicName>
as first described by 
<bibRefCitation id="A37B49D9841EFFFBFCF327C4401BD7BF" author="Tschinkel" box="[825,1135,1488,1513]" pageId="7" pageNumber="296" refString="Tschinkel, W. R. &amp; Doyen, J. T. (1980) Comparative anatomy of the defensive glands, ovipositors and female genital tubes of tenebrionid beetles (Coleoptera). International Journal of Insect Morphology and Embryology, 9, 321 - 368. http: // dx. doi. org / 10.1016 / 0020 - 7322 (80) 90009 - 4" type="journal article" year="1980">Tschinkel &amp; Doyen (1980)</bibRefCitation>
. However, the stenochiine ovipositor has completely transverse paraproctal baculi, whereas these are more or less longitudinal in 
<taxonomicName id="00EA4FAB841EFFFBFF5D240C4535D467" box="[151,321,1560,1585]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFF5D240C4535D467" box="[151,321,1560,1585]" italics="true" pageId="7" pageNumber="296">Trachelostenus</emphasis>
</taxonomicName>
and 
<taxonomicName id="00EA4FAB841EFFFBFEBC240C4668D467" box="[374,540,1560,1585]" class="Insecta" family="Tenebrionidae" genus="Myrmecodema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFEBC240C4668D467" box="[374,540,1560,1585]" italics="true" pageId="7" pageNumber="296">Myrmecodema</emphasis>
</taxonomicName>
. In addition, the stenochiine ovipositor has a pair of newly evolved, extra baculi joining the ends of the paraprocts with the proctigeral baculi. Trachelostenini do not have these extra baculi. Therefore, the particular ovipositor structure of 
<taxonomicName id="00EA4FAB841EFFFBFD12247447F6D42F" box="[728,898,1632,1657]" class="Insecta" family="Tenebrionidae" genus="Trachelostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFD12247447F6D42F" box="[728,898,1632,1657]" italics="true" pageId="7" pageNumber="296">Trachelostenus</emphasis>
</taxonomicName>
and 
<taxonomicName id="00EA4FAB841EFFFBFC0C24744018D42F" box="[966,1132,1632,1657]" class="Insecta" family="Tenebrionidae" genus="Myrmecodema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A841EFFFBFC0C24744018D42F" box="[966,1132,1632,1657]" italics="true" pageId="7" pageNumber="296">Myrmecodema</emphasis>
</taxonomicName>
is unique among known tenebrionid ovipositors.
</paragraph>
</subSubSection>
</treatment>
</document>