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<document id="736E67AFB7E79AA9090E4F8D58AF4107" ID-DOI="10.11646/zootaxa.4020.2.4" ID-GBIF-Dataset="b7fba5bc-8f23-41a5-8cb7-9e368a419017" ID-ISSN="1175-5326" ID-Zenodo-Dep="236475" ID-ZooBank="7FBBE9CA-E98F-4638-8A5E-7E51816B6B79" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269671186" checkinUser="plazi" docAuthor="Matthews, Eric G. &amp; Lawrence, John F." docDate="2015" docId="4F43853E8416FFF3FF5D21C041E2D7D9" docLanguage="en" docName="zt04020p312.pdf" docOrigin="Zootaxa 4020 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leaus" docType="key" docVersion="8" lastPageNumber="304" masterDocId="B37AFD468419FFFCFFCA22144474D256" masterDocTitle="Trachelostenini sensu novo: redescriptions of Trachelostenus Solier, Myrmecodema Gebien and Leaus Matthews &amp; Lawrence, based on adults and larvae, and descriptions of three new species of Leaus (Coleoptera: Tenebrionidae)" masterLastPageNumber="312" masterPageNumber="289" pageNumber="304" updateTime="1698611360402" updateUser="plazi">
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<mods:title id="D2252CBF359B6603B79E6F1453D9C93F">Trachelostenini sensu novo: redescriptions of Trachelostenus Solier, Myrmecodema Gebien and Leaus Matthews &amp; Lawrence, based on adults and larvae, and descriptions of three new species of Leaus (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="8AAC42022F6D5443980D8AE2A2242CB9">Matthews, Eric G.</mods:namePart>
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<mods:namePart id="B1DA01FAA749F2A13A6557C67BC8B36E">Lawrence, John F.</mods:namePart>
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<mods:date id="883506A94FC3306411FED1AE3F5DBD8C">2015</mods:date>
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<treatment id="4F43853E8416FFF3FF5D21C041E2D7D9" ID-DOI="http://doi.org/10.5281/zenodo.6097784" ID-GBIF-Taxon="119638593" ID-Zenodo-Dep="6097784" LSID="urn:lsid:plazi:treatment:4F43853E8416FFF3FF5D21C041E2D7D9" httpUri="http://treatment.plazi.org/id/4F43853E8416FFF3FF5D21C041E2D7D9" lastPageNumber="304" pageId="15" pageNumber="304">
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Key to the species of
<taxonomicName id="00EA4FAB8416FFF3FE5621C04591D1B8" box="[412,485,980,1006]" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="F59EE83A8416FFF3FE5621C04591D1B8" bold="true" box="[412,485,980,1006]" italics="true" pageId="15" pageNumber="304">Leaus</emphasis>
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<paragraph id="C75534288416FFF3FF5D260E41E8D61A" blockId="15.[151,1436,1050,1424]" pageId="15" pageNumber="304">
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1. Prothorax distinctly narrower than elytral bases, with lateral carinae entire, disc with a pair of low median tumosities (
<figureCitation id="5FD128AD8416FFF3FA87260E4488D61A" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Figs 27, 30</figureCitation>
). Elytra with basal tumosities and a transverse depression behind them (
<figureCitation id="5FD128AD8416FFF3FC642623406AD61A" box="[942,1054,1079,1100]" captionStart="FIGURES 33 36" captionStartId="17.[151,264,1953,1976]" captionTargetBox="[232,1358,201,1919]" captionTargetId="figure@17.[198,1389,193,1932]" captionTargetPageId="17" captionText="FIGURES 33 36. Leaus habitus, lateral: 33, L. tasmanicus holotype ♂ (SAMA); 34, L. monteithi sex not determined, paratype (QMBA); 35, L. tropicalis holotype ♀ (QMBA); 36, L. elusus holotype ♂ (QMBA). Scale bars 1 mm." httpUri="https://zenodo.org/record/236487/files/figure.png" pageId="15" pageNumber="304">Figs 33, 34</figureCitation>
). Total length&gt;
<quantity id="001299CD8416FFF3FB7C26234098D61A" box="[1206,1260,1079,1100]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="304" unit="mm" value="5.0">5 mm</quantity>
............ 2
</keyLead>
</paragraph>
<paragraph id="C75534288416FFF3FF5D264141EFD6D1" blockId="15.[151,1436,1050,1424]" pageId="15" pageNumber="304">
<keyLead id="7C1B971D8416FFF3FF5D264141EFD6D1" pageId="15" pageNumber="304">
- Prothorax not or only slightly narrower than elytral bases, with lateral carinae dentate or crenulate (
<figureCitation id="5FD128AD8416FFF3FB6D26414117D63C" box="[1191,1379,1109,1130]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Figs 28, 29, 31, 32</figureCitation>
), disc evenly convex. Elytral disc only slightly uneven. Total length ≤
<quantity id="001299CD8416FFF3FC8D2666470AD6D1" box="[839,894,1138,1159]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="304" unit="mm" value="4.0">4 mm</quantity>
.......................................... 3
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<keyStep id="7C1E2C8D8416FFF3FF5D269B41E8D6AA" pageId="15" pageNumber="304">
<paragraph id="C75534288416FFF3FF5D269B41EFD694" blockId="15.[151,1436,1050,1424]" pageId="15" pageNumber="304">
<keyLead id="7C1B971D8416FFF3FF5D269B41EFD694" pageId="15" pageNumber="304">
2(1). Prothorax widest at middle (
<figureCitation id="5FD128AD8416FFF3FE30269B4637D6F2" box="[506,579,1167,1188]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Fig. 27</figureCitation>
). Basal elytral tumosities not very prominent (
<figureCitation id="5FD128AD8416FFF3FBCF269B4039D6F2" box="[1029,1101,1167,1188]" captionStart="FIGURES 33 36" captionStartId="17.[151,264,1953,1976]" captionTargetBox="[232,1358,201,1919]" captionTargetId="figure@17.[198,1389,193,1932]" captionTargetPageId="17" captionText="FIGURES 33 36. Leaus habitus, lateral: 33, L. tasmanicus holotype ♂ (SAMA); 34, L. monteithi sex not determined, paratype (QMBA); 35, L. tropicalis holotype ♀ (QMBA); 36, L. elusus holotype ♂ (QMBA). Scale bars 1 mm." httpUri="https://zenodo.org/record/236487/files/figure.png" pageId="15" pageNumber="304">Fig. 33</figureCitation>
). Legs uniformly castaneous. Ovi- positor with gonostyli. Western Tas.........................................
<taxonomicName id="00EA4FAB8416FFF3FBFF26B941EFD694" authority="Matthews &amp; Lawrence" authorityName="Matthews &amp; Lawrence" box="[1077,1435,1197,1218]" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="tasmanicus">
<emphasis id="F59EE83A8416FFF3FBFF26B940CED694" box="[1077,1210,1197,1218]" italics="true" pageId="15" pageNumber="304">L. tasmanicus</emphasis>
Matthews &amp; Lawrence
</taxonomicName>
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</paragraph>
<paragraph id="C75534288416FFF3FF5D26DE41E8D6AA" blockId="15.[151,1436,1050,1424]" pageId="15" pageNumber="304">
<keyLead id="7C1B971D8416FFF3FF5D26DE41E8D6AA" pageId="15" pageNumber="304">
- Prothorax widest behind middle (
<figureCitation id="5FD128AD8416FFF3FDEC26DE461BD689" box="[550,623,1226,1247]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Fig. 30</figureCitation>
). Basal elytral tumosities prominent (
<figureCitation id="5FD128AD8416FFF3FC1726DE4052D689" box="[989,1062,1226,1247]" captionStart="FIGURES 33 36" captionStartId="17.[151,264,1953,1976]" captionTargetBox="[232,1358,201,1919]" captionTargetId="figure@17.[198,1389,193,1932]" captionTargetPageId="17" captionText="FIGURES 33 36. Leaus habitus, lateral: 33, L. tasmanicus holotype ♂ (SAMA); 34, L. monteithi sex not determined, paratype (QMBA); 35, L. tropicalis holotype ♀ (QMBA); 36, L. elusus holotype ♂ (QMBA). Scale bars 1 mm." httpUri="https://zenodo.org/record/236487/files/figure.png" pageId="15" pageNumber="304">Fig. 34</figureCitation>
). Legs fuscous with contrasting flavo- castaneous bands. Ovipositor without gonostyli. NE NSW.......................................
<taxonomicName id="00EA4FAB8416FFF3FB2626F34114D6AA" box="[1260,1376,1255,1276]" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="F59EE83A8416FFF3FB2626F34114D6AA" box="[1260,1376,1255,1276]" italics="true" pageId="15" pageNumber="304">L. monteithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EEAD55418416FFF3FAAC26F341E8D6AA" box="[1382,1436,1255,1277]" pageId="15" pageNumber="304" rank="species">
<emphasis id="F59EE83A8416FFF3FAAC26F341E1D6AB" bold="true" box="[1382,1429,1255,1277]" pageId="15" pageNumber="304">sp. n</emphasis>
.
</taxonomicNameLabel>
</keyLead>
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</keyStep>
<keyStep id="7C1E2C8D8416FFF3FF5D271141E8D702" pageId="15" pageNumber="304">
<paragraph id="C75534288416FFF3FF5D271141E8D761" blockId="15.[151,1436,1050,1424]" pageId="15" pageNumber="304">
<keyLead id="7C1B971D8416FFF3FF5D271141E8D761" pageId="15" pageNumber="304">
3(1). Lateral pronotal carinae irregularly dentate, prothorax a little narrower than elytral bases (
<figureCitation id="5FD128AD8416FFF3FBF7271140D8D74C" box="[1085,1196,1285,1306]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Figs 28, 29</figureCitation>
). Qld wet tropics...........................................................................................
<taxonomicName id="00EA4FAB8416FFF3FB212736412BD761" box="[1259,1375,1314,1335]" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="tropicalis" status="sp. nov.">
<emphasis id="F59EE83A8416FFF3FB212736412BD761" box="[1259,1375,1314,1335]" italics="true" pageId="15" pageNumber="304">L. tropicalis</emphasis>
</taxonomicName>
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<emphasis id="F59EE83A8416FFF3FAAC273641E1D76E" bold="true" box="[1382,1429,1314,1336]" pageId="15" pageNumber="304">sp. n</emphasis>
.
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</paragraph>
<paragraph id="C75534288416FFF3FF5D272B41E8D702" blockId="15.[151,1436,1050,1424]" box="[151,1436,1343,1364]" pageId="15" pageNumber="304">
<keyLead id="7C1B971D8416FFF3FF5D272B41E8D702" box="[151,1436,1343,1364]" pageId="15" pageNumber="304">
- Lateral pronotal carinae regularly denticulate or crenulate. Prothorax as wide as elytral bases (
<figureCitation id="5FD128AD8416FFF3FBAC272B40A7D702" box="[1126,1235,1343,1364]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Figs 31, 32</figureCitation>
).............. 4
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<paragraph id="C75534288416FFF3FF5D274941E8D724" blockId="15.[151,1436,1050,1424]" box="[151,1436,1372,1394]" pageId="15" pageNumber="304">
<keyLead id="7C1B971D8416FFF3FF5D274941E8D724" box="[151,1436,1372,1394]" pageId="15" pageNumber="304">
4(3). Lateral pronotal carinae crenulate (
<figureCitation id="5FD128AD8416FFF3FDFA2749460CD724" box="[560,632,1373,1394]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Fig. 31</figureCitation>
). NE NSW.............................................
<taxonomicName id="00EA4FAB8416FFF3FAC127494114D724" box="[1291,1376,1373,1394]" class="Insecta" family="Tenebrionidae" genus="Leaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="304" phylum="Arthropoda" rank="species" species="elusus" status="sp. nov.">
<emphasis id="F59EE83A8416FFF3FAC127494114D724" box="[1291,1376,1373,1394]" italics="true" pageId="15" pageNumber="304">L. elusus</emphasis>
</taxonomicName>
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<emphasis id="F59EE83A8416FFF3FAAC274841E8D724" bold="true" box="[1382,1436,1372,1394]" pageId="15" pageNumber="304">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
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<keyLead id="7C1B971D8416FFF3FF5D276E41E2D7D9" box="[151,1430,1402,1424]" pageId="15" pageNumber="304">
- Lateral pronotal carinae evenly denticulate (
<figureCitation id="5FD128AD8416FFF3FD42276E46A4D7D9" box="[648,720,1402,1423]" captionStart="FIGURES 27 32" captionStartId="16.[151,264,1895,1918]" captionTargetBox="[181,1424,202,1851]" captionTargetId="figure@16.[151,1436,193,1874]" captionTargetPageId="16" captionText="FIGURES 27 32. Leaus habitus, dorsal except 29: 27, L. tasmanicus paratype ♀ (SAMA); 28, L. tropicalis holotype ♀ (QMBA); 29, L. tropicalis paratype ♂ (ANIC); 30, L. monteithi holotype ♀ (QMBA); 31, L. elusus holotype ♂ (QMBA); 32, L. sp. ♀ (ANIC). Scale bars 1 mm." httpUri="https://zenodo.org/record/236486/files/figure.png" pageId="15" pageNumber="304">Fig. 32</figureCitation>
). SE Qld...............................................
<emphasis id="F59EE83A8416FFF3FA95276E4107D7D9" box="[1375,1395,1402,1423]" italics="true" pageId="15" pageNumber="304">L.</emphasis>
<emphasis id="F59EE83A8416FFF3FAB3276E41FBD7C6" bold="true" box="[1401,1423,1402,1424]" pageId="15" pageNumber="304">sp</emphasis>
.
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