201 lines
24 KiB
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201 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="475FAEB4E99A5A22A5DF46D3D21DE39B" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole anomala Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="7E2C359F-AA66-43D9-A591-8D5B8D2C788F" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090184" LSID="urn:lsid:zoobank.org:act:7E2C359F-AA66-43D9-A591-8D5B8D2C788F" httpUri="http://treatment.plazi.org/id/475FAEB4E99A5A22A5DF46D3D21DE39B" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="475FAEB4-E99A-5A22-A5DF-46D3D21DE39B" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anomala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anomala" status="sp. nov.">Pheidole anomala</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">Figs 23A-F</figureCitation>
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63E</figureCitation>
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<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Parc National Montagne
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<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
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, Mahasarika;
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<geoCoordinate degrees="12.53176" direction="south" orientation="latitude" precision="1" value="-12.53176">-12.53176</geoCoordinate>
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,
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<geoCoordinate degrees="49.17662" direction="east" orientation="longitude" precision="1" value="49.17662">49.17662</geoCoordinate>
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; alt. 1135 m; 17 Nov 2007; B. L. Fisher et al. leg.; montane rainforest, ex rotten log; BLF18461; CASENT0134498 (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0235124 (CASC).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1" start="Figure 23" startId="F23">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 23.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anomala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anomala">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole anomala</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0235124) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0134498).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval and slightly widening posteriorly, with anterior and posterior sides slightly convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with dense, long, suberect to erect pilosity; medial part of frons with irregular and dense rugae, interspaces distinctly rugulate; occipital lobes, and area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by one-fifth of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed inward; outer hypostomal teeth lobe-like, wider than and approximately as high as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity; mesosoma rugofoveolate; gaster smooth; body ferruginous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate; vertex with a few arcing, sparse, and moderately thick rugae; scape, when laid back, exceeding the posterior head margin by one-third of its length; promesonotum very low and moderately long; promesonotal groove absent; propodeal spines small, triangular; mesosoma foveolate; body yellow.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 1.42; HW: 1.37; SL: 0.87; EL: 0.18; WL: 1.25; PSL: 0.21; MTL: 0.87; PNW: 0.57; PTW: 0.16; PPW: 0.34; CI: 103.0; SI: 63.2; PSLI: 14.8; PPI: 47.8; PNI: 41.6; MTI: 63.2.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face sub-oval, widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with dense, long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Frons with thick, irregular, and dense rugae, interspaces distinctly rugulate. Occipital lobes with slightly thinner, dense, and irregular rugae and rugulate interspaces. Area posterolateral from eyes with dense rugoreticulae. Gena with relatively sparse and thick, longitudinal rugae with distinctly rugofoveolate interspaces. Centre of clypeus with sparse foveolae and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23B, D</figureCitation>
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). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed inward; outer hypostomal teeth lobe-like, wider than and approximately as high as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderate, with wide base and acute apex; humeral area produced (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23D</figureCitation>
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). Surface shiny and rugofoveolate. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity moderately dense, long, and erect (Fig. D, F).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Ferruginous with yellowish legs (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23D, F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 0.66; HW: 0.52; SL: 0.82; EL: 0.15; WL: 0.89; PSL: 0.12; MTL: 0.7; PNW: 0.36; PTW: 0.1; PPW: 0.16; CI: 128.2; SI: 158.8; PSLI: 17.4; PPI: 62.7; PNI: 69.5; MTI: 136.7.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Cephalic margin slightly convex (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23A</figureCitation>
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). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and foveolate; vertex with a few arcing, sparse, and moderately thick rugae; antennal sockets with few indistinct, curved outward rugae and foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-third of its length; pilosity dense, subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23A, C</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum very low and moderately long, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines small, triangular (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23C</figureCitation>
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). Sculpture shiny and foveolate. Pilosity very sparse, moderately long, and erect (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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With sparse, erect pilosity (Fig.
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Yellow, legs brighter (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pheidole anomala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235124) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134498)." figureDoi="10.3897/zookeys.949.51269.figure23" httpUri="https://binary.pensoft.net/fig/432192" pageId="0" pageNumber="1">23C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Latin for irregular in reference to strongly irregular rugae on frons of major workers.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected at 1135 m in elevation, in montane rainforest. Nest was located in a rotten log.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole anomala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anomala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole anomala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName lsidName="P. veteratrix" pageId="0" pageNumber="1" rank="species" species="veteratrix">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. veteratrix</emphasis>
|
||
</taxonomicName>
|
||
Forel, a widespread species known from Antananarivo north to Taolagnaro, and sympatric
|
||
<taxonomicName lsidName="P. trichotos" pageId="0" pageNumber="1" rank="species" species="trichotos">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. trichotos</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Major workers of
|
||
<taxonomicName lsidName="P. anomala" pageId="0" pageNumber="1" rank="species" species="anomala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. anomala</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from both taxa in ferruginous body colouration and distinctly rugulate interspaces on frons; minor workers are easy to distinguish based on their bright yellow body colouration and entirely foveolate katepisternum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |