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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="446F97FF1E36ABA5707E784C57A0F4BE" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus arquus Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="226" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="224" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152041015" LSID="urn:lsid:zoobank.org:act:DCDF486F-9574-43D1-B391-B1EDFAE5D241" httpUri="http://treatment.plazi.org/id/446F97FF1E36ABA5707E784C57A0F4BE" lastPageId="225" lastPageNumber="226" pageId="223" pageNumber="224">
<subSubSection lastPageId="224" lastPageNumber="225" pageId="223" pageNumber="224" type="nomenclature">
<paragraph lastPageId="224" lastPageNumber="225" pageId="223" pageNumber="224">
<taxonomicName LSID="urn:lsid:zoobank.org:act:DCDF486F-9574-43D1-B391-B1EDFAE5D241" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arquus" order="Coleoptera" pageId="223" pageNumber="224" phylum="Arthropoda" rank="species" species="arquus">Operclipygus arquus</taxonomicName>
<taxonomicNameLabel pageId="224" pageNumber="225">
<pageBreakToken pageId="224" pageNumber="225" start="start">sp</pageBreakToken>
. n.
</taxonomicNameLabel>
Figs 56A, C57KMap 21
</paragraph>
</subSubSection>
<subSubSection pageId="224" pageNumber="225" type="type locality">
<paragraph pageId="224" pageNumber="225">Type locality.</paragraph>
<paragraph pageId="224" pageNumber="225">
COSTA RICA: Puntarenas: Rincon de Osa [
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.685683">8°41.141'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.518616">83°31.117'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="224" pageNumber="225" type="type material">
<paragraph pageId="224" pageNumber="225">Type material.</paragraph>
<paragraph pageId="224" pageNumber="225">
Holotype male: &quot;COSTA RICA: Punta. Prov, Rincon de Osa, 50m
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.685683">8°41.141'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.518616">83°31.117'W</geoCoordinate>
, 23-26-VI-2001,S.&amp;J.Peck, 01-13, ex FIT, CR1P01 005&quot; / SM0520653 KUNHM-ENT&quot; (INBIO). Paratypes (16): COSTA RICA: Puntarenas: 1: Rincon de Osa,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.685683">8°41.141'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.518616">83°31.117'W</geoCoordinate>
, 40m, 23-26.vi.2001, FIT, S. &amp; J. Peck (SEMC), 2: 150m, 23-26.vi.2001, FIT, S. &amp; J. Peck (CMNC); 4: Rancho Quemado, Peninsula de Osa, 200m, ix.1992, F. Quesada, (INBIO, FMNH, MSCC, AKTC), 1: 292500, 511000, 200m, v.1991, F. Quesada, (INBIO), 2: v.1992, F. Quesada y G. Varela, (INBIO), 1: x.1992, F. Quesada, (INBIO), 1: xi.1992, M. Segura, (INBIO); 2: Reserva Forestal Golfo Dulce, Est. Agujas, Golfito, Sendero Zamia, 250-350m, 9-12.viii.1998, FIT, A. Azofeifa (INBIO), 2: 250-350m, 11-15.xi.1999, FIT, A. Azofeifa (INBIO).
</paragraph>
</subSubSection>
<subSubSection pageId="224" pageNumber="225" type="other material">
<paragraph pageId="224" pageNumber="225">Other material.</paragraph>
<paragraph pageId="224" pageNumber="225">
COSTA RICA: Puntarenas: 6: Est. Biol. Las Cruces, San Vito, 17.
<normalizedToken originalValue="viii">viii-</normalizedToken>
12.ix.1982, FIT, B.D. Gill (BDGC, MSCC, AKTC), 3: 1200m, 1-30.vii.1982, FIT, B.D. Gill (BDGC).
</paragraph>
</subSubSection>
<subSubSection lastPageId="225" lastPageNumber="226" pageId="224" pageNumber="225" type="description">
<paragraph pageId="224" pageNumber="225">Diagnostic description.</paragraph>
<paragraph lastPageId="225" lastPageNumber="226" pageId="224" pageNumber="225">
Length: 2.03-2.12 mm, width: 1.62-1.75 mm; body rufescent, ovoid, sides rounded; frons depressed at middle, prominent over antennae; frontal stria rounded at sides, broadly interrupted in most, with occasional median fragments present; labrum about 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long, very weakly emarginate apically; pronotum with ground punctation fine and sparse with few, ~6-8 coarse lateral punctures, prescutellar region slightly depressed, but lacking distinct fovea; marginal pronotal stria not descending to hypomeron; pronotum with complete lateral submarginal stria close to margin, lateral margin slightly sinuate toward base; anterior submarginal stria detached, forming more or less continuous arc across front of pronotum, reaching back one-fourth to one-third pronotal length; elytra with outer subhumeral stria present in apical half or less, inner subhumeral stria absent, striae 1-4 very fine but complete, 4th often fragmented toward base, 5th stria present in apical third, sutural stria present in apical half; venter with faint microsculpture almost throughout, effaced on swelling of metaventrite; prosternum, mesoventrite, and anterior half of metaventrite depressed (female with depression of metaventrite less distinct); prosternal keel weakly emarginate at base, with complete carinal striae rather widely separated to near apex; prosternal lobe with weak swelling at center of base; mesoventrite very weakly projecting at middle, with marginal stria broadly interrupted; mesometaventral stria strongly arched forward to near mesoventral margin; metaventrite markedly convex in middle of basal half (in both sexes); lateral metaventral stria extending toward outer corner of metacoxa, but generally abbreviated just behind middle; 1st abdominal ventrite with two strongly abbreviated lateral striae; propygidium and pygidium both with conspicuous transverse microsculpture; propygidium with sparse, moderately large punctures mainly in basal half; male especially with anterior half of pygidium simply, finely sparsely punctate, females with coarser pygidial punctures; fine apical marginal stria present only in apical half. Male genitalia very similar to those of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="224" pageNumber="225" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
(see Figs 57
<normalizedToken originalValue="AD">A-D</normalizedToken>
), differing as follows: T8
<pageBreakToken pageId="225" pageNumber="226" start="start">with</pageBreakToken>
sides subparallel; S8 shorter, with apices bluntly subacuminate, ventral creases conspicuous; tegmen (Fig. 57K) shorter, widest distinctly basad middle, with sides rounded, not as strongly narrowed to apex; indistinguishable from that of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pichinchensis" order="Coleoptera" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="pichinchensis">Operclipygus pichinchensis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="225" pageNumber="226" type="remarks">
<paragraph pageId="225" pageNumber="226">Remarks.</paragraph>
<paragraph pageId="225" pageNumber="226">This species is fairly easily recognized in this complex by its broadly arcuate anterior submarginal pronotal stria (Fig. 56C).</paragraph>
</subSubSection>
<subSubSection pageId="225" pageNumber="226" type="etymology">
<paragraph pageId="225" pageNumber="226">Etymology.</paragraph>
<paragraph pageId="225" pageNumber="226">The name of this species means arc, referring to its curved pronotal stria. It is a noun in apposition.</paragraph>
</subSubSection>
</treatment>
</document>