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<document id="181191B449D10F82B186E6D5E29CF4E3" ID-CLB-Dataset="304167" ID-DOI="10.3897/zse.100.128717" ID-GBIF-Dataset="a7f81ef5-5178-48c8-8d40-80b29116daad" ID-ZooBank="FD4C02D2-01F9-4A4C-817B-F81B27F22A41" ID-publisher-id="128717" URI-arpha="A58A01B3-C620-5671-AB59-ABE7916A4ADE" URI-zoobank="http://zoobank.org/FD4C02D2-01F9-4A4C-817B-F81B27F22A41" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1728062842085" checkinUser="pensoft" docAuthor="Klongklaew, Kanyaporn, Poeaim, Supattra &amp; Dumrongrojwattana, Pongrat" docDate="2024" docId="D62C173166ED58C696F84F0A08A5ED14" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1347-1360" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/128717/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Georissa sagitta Klongkaew, Poeaim &amp; Dumrongrojwattana 2024, sp. nov." docType="treatment" docUuid="965F36E8-E123-496F-A43D-9C2D1B80294F" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="1360" masterDocId="A58A01B3C6205671AB59ABE7916A4ADE" masterDocTitle="Four new species of Georissa W. Blanford, 1864 (Gastropoda, Hydrocenidae) from Thailand" masterLastPageNumber="1360" masterPageNumber="1347" pageNumber="1347" updateTime="1728065275675" updateUser="admin">
<mods:mods id="388971584A2E6E1425125586CF45936C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B28E495E972561F7260A63D6C76EFD4">
<mods:title id="479F4A95654618CDB4EAC9566563A1EB">Four new species of Georissa W. Blanford, 1864 (Gastropoda, Hydrocenidae) from Thailand</mods:title>
</mods:titleInfo>
<mods:name id="971186ADD7F5805730B97D7A2EE1DC3C" type="personal">
<mods:role id="C5CF5BDC2566CC49BE9B3698FE175ED9">
<mods:roleTerm id="F0478F178FBA1E93384B8875B7B34E99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E2EF4EA6B5600915692D714860F84D3D">Klongklaew, Kanyaporn</mods:namePart>
<mods:affiliation id="A1F9AED4BD67E76124AEDE36307430F8">Department of Biology, School of Science, King Mongkut's Institute of Technology Ladkrabang, Bangkok 10520, Thailand</mods:affiliation>
</mods:name>
<mods:name id="080D19649AFE9FCD818764D0846B5CF9" type="personal">
<mods:role id="ED4E0287BD4CFE151F58B5E6F55C87E9">
<mods:roleTerm id="526913543516191ACA74BF4A7317B07E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D773EF155E21C87C7CECB61DB4D1B95">Poeaim, Supattra</mods:namePart>
<mods:nameIdentifier id="10E1E4ED1C975E69914B2E96C0A17A79" type="ORCID">0000-0003-1293-3820</mods:nameIdentifier>
<mods:affiliation id="C6A9388731B4DFFFD1033C11298EC33F">Department of Biology, School of Science, King Mongkut's Institute of Technology Ladkrabang, Bangkok 10520, Thailand</mods:affiliation>
</mods:name>
<mods:name id="A4B4A1D531A73850F8324C40E6B2D86C" type="personal">
<mods:role id="F0F8B31101E36E54E4CE9920C054D83A">
<mods:roleTerm id="66B499755DBA77AEA71595402336CF42">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F012A66D721BE84B9E8C2C0125019556">Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation id="71D3AB1953B3DE76D4310A88B0E6CED4">Department of Biology, Faculty of Science, Burapha University, Bangsaen, Chonburi 20131, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E8E66552571B5E18ADE5AA4D6D3667BF">text</mods:typeOfResource>
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<mods:titleInfo id="462B533CC0B83B2172B3D5EB419B764E">
<mods:title id="E701A30352ED7A61AA969AD36DC9EBFD">Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part id="9CC7165BD8645DF0A30DC69A7178E456">
<mods:date id="503002B8CA75F80C34173053A2FAA4F0">2024</mods:date>
<mods:detail id="C125729EB542515ACC7AD9C1D00494D7" type="pubDate">
<mods:number id="6B9B00A46ACC0DC402E116022E41250F">2024-10-04</mods:number>
</mods:detail>
<mods:detail id="0E603AD58EB05BD8C14385A254877337" type="volume">
<mods:number id="6008C4E348684AC1F6D0CC34B7A83901">100</mods:number>
</mods:detail>
<mods:detail id="206004DEF6635A1342EB8CD0B635865C" type="issue">
<mods:number id="5EFC2373103E75FC19F2CC0096CFD780">4</mods:number>
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<mods:start id="5103F560B60000E66A98F19E3F516188">1347</mods:start>
<mods:end id="0F0FFBE19428292C11D83459BF0C542B">1360</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="6F635D8A685AE1DB85087798CF69917D">journal article</mods:classification>
<mods:identifier id="0ECD68BD19435236F8796F9507C588CF" type="CLB-Dataset">304167</mods:identifier>
<mods:identifier id="EC3305FD54392B435DD15FD0D9E1BE44" type="DOI">10.3897/zse.100.128717</mods:identifier>
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</mods:mods>
<treatment id="D62C173166ED58C696F84F0A08A5ED14" ID-DOI="http://doi.org/10.5281/zenodo.13891563" ID-Zenodo-Dep="13891563" ID-arpha="D62C1731-66ED-58C6-96F8-4F0A08A5ED14" ID-zoobank="https://zoobank.org/965F36E8-E123-496F-A43D-9C2D1B80294F" LSID="urn:lsid:zoobank.org:act:965F36E8-E123-496F-A43D-9C2D1B80294F" LSID-ZBK="urn:lsid:zoobank.org:act:965F36E8-E123-496F-A43D-9C2D1B80294F" httpUri="http://treatment.plazi.org/id/D62C173166ED58C696F84F0A08A5ED14">
<subSubSection id="5F9989685EC59713ED99FF0F4876CB61" type="nomenclature">
<paragraph id="9A4BD120B01813EC83AFD0D96576B217">
<taxonomicName id="3FE5AD6D06E1DF91EA6C298EB277EC86" authority="Klongkaew, Poeaim &amp; Dumrongrojwattana" authorityName="Klongkaew, Poeaim &amp; Dumrongrojwattana" authorityYear="2024" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="sagitta" status="sp. nov.">
<emphasis id="D0B46DAEEA54656CC8A96E2FE8D66265" italics="true">Georissa sagitta</emphasis>
Klongkaew, Poeaim &amp; Dumrongrojwattana
</taxonomicName>
<taxonomicNameLabel id="8D31166AFF3888D83994196573E81BC2" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B0A9579C63F067CA2376DE4C5E713D37" type="description">
<paragraph id="FED0BF42CA90C1957111CA3762D77261">
<figureCitation id="74656F3286DC9E149A630D54765DB973" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">Figs 2 C</figureCitation>
,
<figureCitation id="D18AF01E310476ED241FF208EBD8800D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 I – L</figureCitation>
,
<figureCitation id="D91C76C75E292B517D29782BD5A3C696" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of radula morphology of Georissa. A – D. G. quinquelirata sp. nov.; E – H. G. kohsichangensis sp. nov.; A, E. Radula overview; B, F. Radula segment; C, G. Lateral teeth (Lt); D, H. Marginal teeth (Mt); I – L. G. sagitta sp. nov. M – P; G. digitinota sp. nov.; I, M. Radula segment; J, M. Radula segment close up; K, O. Lateral teeth (Lt); L, P. Marginal teeth (Mt). Color highlights show radula position; blue: small central teeth (Ct), purple: lateral teeth, and orange: marginal teeth." figureDoi="10.3897/zse.100.128717.figure5" httpUri="https://binary.pensoft.net/fig/1148344">5 I – L</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMTBG" type="Type examined">
<paragraph id="80C2D63CE4BE10C3F231FA81E99F8E63">
<heading id="D6DD355536D430559E66E9AA1EEE5182" reason="title">Type examined.</heading>
</paragraph>
<paragraph id="22D01526F5B11582CC4F713A1916A509">
<materialsCitation id="E394CCD8A6CC3FC782D76276CB64DD9B" collectingDate="2019-07-20" collectionCode="ZRCBUU" collectorName="P. Dumrongrojwattana" country="Thailand" county="Mueang Sakaeo district" latitude="13.787945" location="Khao Maka Cave Temple" longLatPrecision="1" longitude="101.94825" specimenCode="ZRCBUU 0904" specimenCount="1" stateProvince="Sakaeo Province" typeStatus="holotype">
<emphasis id="A1759A00E0A26350384F0A21B44EC6B7" bold="true" italics="true">
<typeStatus id="706565EF70041F2837BCBA78804C231A" type="holotype">Holotype</typeStatus>
</emphasis>
•
<specimenCode id="729523852CECA54ED71203B311BEC887">
<abbrev id="ABBRID0EXTBG" xlink_title="Zoological Research Collection of Burapha University">ZRCBUU</abbrev>
0904
</specimenCode>
(Fig.
<figureCitation id="018D1EF81BAC4B51AB6EF17E3FA029E4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 C</figureCitation>
);
<abbrev id="ABBRID0E6TBG" xlink_title="Shell height">SH</abbrev>
=
<quantity id="2BAA9DDC2462D82734989130F193A81F" metricMagnitude="-3" metricUnit="m" metricValue="2.61" unit="mm" value="2.61">2.61 mm</quantity>
,
<abbrev id="ABBRID0EDUBG" xlink_title="shell width">SW</abbrev>
=
<quantity id="0BD14534A76E1CE9FFD1C0D91A5BAE2D" metricMagnitude="-3" metricUnit="m" metricValue="1.63" unit="mm" value="1.63">1.63 mm</quantity>
,
<abbrev id="ABBRID0EHUBG" xlink_title="aperture height">AH</abbrev>
=
<quantity id="EF2B08FEE0AAB5BC9D6949950E304082" metricMagnitude="-4" metricUnit="m" metricValue="7.9" unit="mm" value="0.79">0.79 mm</quantity>
,
<abbrev id="ABBRID0ELUBG" xlink_title="aperture width">AW</abbrev>
=
<quantity id="C377A7293D9F1948B66E4ADA2AFAEC31" metricMagnitude="-4" metricUnit="m" metricValue="9.8" unit="mm" value="0.98">0.98 mm</quantity>
, W = 4.25–4.5
</materialsCitation>
.
<materialsCitation id="34D4BAFF26A13178A76D25B2964A5557" collectingDate="2019-07-20" collectionCode="ZRCBUU" collectorName="P. Dumrongrojwattana" country="Thailand" county="Mueang Sakaeo district" latitude="13.787945" location="Khao Maka Cave Temple" longLatPrecision="1" longitude="101.94825" specimenCode="ZRCBUU 0905" specimenCount="1" stateProvince="Sakaeo Province" typeStatus="paratype">
<emphasis id="F22DF73F258FEDE47CBCC27B75EFE788" bold="true" italics="true">
<typeStatus id="51CA9BF9CF7BFC785DEE0525E74A0E50" type="paratype">Paratypes</typeStatus>
</emphasis>
•
<specimenCode id="7364976CE3DCBC51CA120629A42BE72C">
<abbrev id="ABBRID0ESUBG" xlink_title="Zoological Research Collection of Burapha University">ZRCBUU</abbrev>
0905
</specimenCode>
(Fig.
<figureCitation id="12595592BE5992AFF55EB1AEC04EEF0B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 I – L</figureCitation>
) (10 shells); Shell measurements:
<abbrev id="ABBRID0E1UBG" xlink_title="Shell height">SH</abbrev>
=
<quantity id="61AC14D47C226B50AE51B3C7DCEBD588" metricMagnitude="-3" metricUnit="m" metricValue="2.66" metricValueMax="2.88" metricValueMin="2.44" unit="mm" value="2.66" valueMax="2.88" valueMin="2.44">2.44–2.88 mm</quantity>
(2.65 ±
<quantity id="FFF66BA7AF57C8A93E9A687A57D6CAE3" metricMagnitude="-4" metricUnit="m" metricValue="1.4" unit="mm" value="0.14">0.14 mm</quantity>
),
<abbrev id="ABBRID0E5UBG" xlink_title="shell width">SW</abbrev>
=
<quantity id="74520A05E74707AAC9BB628D1A8D86A6" metricMagnitude="-3" metricUnit="m" metricValue="1.7349999999999999" metricValueMax="1.84" metricValueMin="1.63" unit="mm" value="1.7349999999999999" valueMax="1.84" valueMin="1.63">1.63–1.84 mm</quantity>
(1.70 ±
<quantity id="2B0A2C157FD9E2EF9C94F2B41BAE97D6" metricMagnitude="-5" metricUnit="m" metricValue="7" unit="mm" value="0.07">0.07 mm</quantity>
),
<abbrev id="ABBRID0ECVBG" xlink_title="aperture height">AH</abbrev>
=
<quantity id="FBF9FB3F6FA7329B7019C4017C62AE94" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="9.7" metricValueMin="7.9" unit="mm" value="0.88" valueMax="0.97" valueMin="0.79">0.79–0.97 mm</quantity>
(0.87 ±
<quantity id="AA8C00E68DDD221FB4C2624FEDE96FC3" metricMagnitude="-5" metricUnit="m" metricValue="5.0" unit="mm" value="0.05">0.05 mm</quantity>
),
<abbrev id="ABBRID0EGVBG" xlink_title="aperture width">AW</abbrev>
=
<quantity id="CFF8795952C4463981E0D8E69430CADB" metricMagnitude="-3" metricUnit="m" metricValue="1.02" metricValueMax="1.09" metricValueMin="0.95" unit="mm" value="1.02" valueMax="1.09" valueMin="0.95">0.95–1.09 mm</quantity>
(1.00 ±
<quantity id="2D4AE98FF2B5A09BF7B889EDD3CEB9C8" metricMagnitude="-5" metricUnit="m" metricValue="4.0" unit="mm" value="0.04">0.04 mm</quantity>
), W = 4–4.5
</materialsCitation>
(all type material from type locality;
<date id="2151E6688470ACB56786F8F83C516779" value="2019-07-20">
<collectingDate id="CB9866018F2E7A060CDE95C5A63D009A" value="2019-07-20">20 July 2019</collectingDate>
</date>
,
<collectorName id="20CEF7F50CA44C15E8DA5AE871C26A2D">P. Dumrongrojwattana</collectorName>
leg).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKVBG" type="type locality">
<paragraph id="CE27582C085470D44CCA574B247042E8">
<heading id="6CE7C7DF18A29370EFDA53D110A59FC3" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="2D4988B7EC27AAD2DFCBE421ADE77872">
<collectingCountry id="0BCEE4D765F97004B08DF58768A59402" name="Thailand">Thailand</collectingCountry>
,
<location id="943D6538B67D6DDD2270DA2BCB71140E" LSID="urn:lsid:plazi:treatment:D62C173166ED58C696F84F0A08A5ED14:943D6538B67D6DDD2270DA2BCB71140E" country="Thailand" county="Mueang Sakaeo district" latitude="13.787945" longLatPrecision="1" longitude="101.94825" name="Khao Maka Cave Temple" stateProvince="Sakaeo Province">Khao Maka Cave Temple</location>
,
<collectingCounty id="BBB18DF566DE29D2602094048299E26A">Mueang Sakaeo district</collectingCounty>
,
<collectingRegion id="01C2EC2A0D471B2F8AB940FE619BDA8B">Sakaeo Province</collectingRegion>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETVBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.948250,13.787944]}">
<geoCoordinate id="5C71EBAD9D132149C65DCBA0EA6E0769" degrees="13" direction="north" minutes="47" orientation="latitude" precision="1" seconds="16.6" value="13.787945">13 ° 47 ' 16.6 &quot; N</geoCoordinate>
,
<geoCoordinate id="E6B6F91ABC5F2FCF5FD849012400806D" degrees="101" direction="east" minutes="56" orientation="longitude" precision="1" seconds="53.7" value="101.94825">101 ° 56 ' 53.7 &quot; E</geoCoordinate>
</named-content>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYVBG" type="etymology">
<paragraph id="57D1342703459D5D413F3A8696641230">
<heading id="63A8F94C334B45BF8C4F2ED6DF5F9DA9" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="60718DF9FB8565E32A2542CED3423451">The specific designation “ sagitta ” for the prominent oval indentation resembling a target in archery.</paragraph>
</subSubSection>
<subSubSection id="SECID0EHWBG" type="diagnosis">
<paragraph id="64F47251C0CA43EDE3769ADED0039425">
<heading id="35CD6A83D35A8191E9241B20C4468DD1" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D7412235E1627274B836DBE1FB13FE91">Shell minute, ovately conical, orangish, protoconch round to slightly ellipsoid and mix of rounded, ellipsoidal to irregular sculpture with oval indentation. Body whorl peripherally rounded with 8–9 spiral lirae. Aperture semi-rounded. Umbilicus closed. Operculum corneous with apophysis, paucispiral, opaque white, and semi-rounded.</paragraph>
</subSubSection>
<subSubSection id="SECID0EMWBG" type="description">
<paragraph id="BAF2F43C22AED8E250832D6E30DA5595">
<heading id="27CA340F859FB922E94D7BE66080E7D5" reason="title">Description.</heading>
</paragraph>
<paragraph id="B152EC158EDF32333F75A2D00BC9E7AB">
Shell minute, dextral, orangish to brownish, ovately conical with 4.25–4.5 whorls (Fig.
<figureCitation id="9F4F105635428C1CC48A5D2E65A52229" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 C</figureCitation>
). Protoconch orange consists of 1 whorl, round to slightly ellipsoid, covered with mix of rounded, ellipsoidal to irregular sculpture and a large oval indentation present (Figs
<figureCitation id="F2D942C73F1E1F6965D571BCB9E55262" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 J</figureCitation>
,
<figureCitation id="0A1BBFCFE4EE463183935D3CC5CDD488" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 K</figureCitation>
). Teleconch orange, consists of 3.25–3.5 whorls. Peripherally rounded, sculptured with 8–9 prominent lirae on the body whorl, crossed with weak oblique growth lines. Basal part of the body whorl, there are more densely spaced spiral cords, approximately 7–8 cords (Figs
<figureCitation id="5C581BEFBD33357F66A2C3B0351C8174" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of the new Georissa species: A. G. quinquelirata sp. nov. (Holotype ZRCBUU 0900); B. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0902); C. G. sagitta sp. nov. (Holotype ZRCBUU 0904), and D. G. digitinota sp. nov. (Holotype ZRCBUU 0906)." figureDoi="10.3897/zse.100.128717.figure2" httpUri="https://binary.pensoft.net/fig/1148341">2 C</figureCitation>
,
<figureCitation id="10828EC49A4A6D98090DC139EDEDA5D5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 I</figureCitation>
). Suture incised. Aperture semi-rounded. Peristome thin, sharp, unexpanded or thickened, not reflected. Umbilicus closed. Operculum paucispiral, nucleus submarginal, corneous, opaque white, semi-rounded, thin, smooth outside surface with apophysis, inside surface with an arched peg arising from base (Fig.
<figureCitation id="30210047CA24687D85C5803F8550FA30" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Georissa new species. A – D. G. quinquelirata sp. nov. (Holotype ZRCBUU 0901); E – H. G. kohsichangensis sp. nov. (Holotype ZRCBUU 0903); A, E. Shell; B, F. Protoconch; C, G. Sculpture of protoconch; D, H. Operculum; I – L. G. sagitta sp. nov. (Holotype ZRCBUU 0905); M – P. G. digitinota sp. nov. (Holotype ZRCBUU 0907); I, M. Shell; J, N. Protoconch; K, O. Sculpture of protoconch; L, P. Operculum." figureDoi="10.3897/zse.100.128717.figure4" httpUri="https://binary.pensoft.net/fig/1148343">4 L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKXBG" type="Radula">
<paragraph id="B2ABE6BF712F1489722A1DD31072E7F8">
<heading id="F9BED8C885AD498B2AE47AC88F50340F" reason="title">Radula.</heading>
</paragraph>
<paragraph id="7E7556A98833CD162A57F186CDA04CA1">
Ribbon-like, slender, longer and delicate towards the outside, rhipidoglossate (Fig.
<figureCitation id="E1F36BFD73411B4C36B8313E15DD0092" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of radula morphology of Georissa. A – D. G. quinquelirata sp. nov.; E – H. G. kohsichangensis sp. nov.; A, E. Radula overview; B, F. Radula segment; C, G. Lateral teeth (Lt); D, H. Marginal teeth (Mt); I – L. G. sagitta sp. nov. M – P; G. digitinota sp. nov.; I, M. Radula segment; J, M. Radula segment close up; K, O. Lateral teeth (Lt); L, P. Marginal teeth (Mt). Color highlights show radula position; blue: small central teeth (Ct), purple: lateral teeth, and orange: marginal teeth." figureDoi="10.3897/zse.100.128717.figure5" httpUri="https://binary.pensoft.net/fig/1148344">5 I – L</figureCitation>
). Central tooth five small teeth, trapezoid shape. Lateral teeth five teeth, claw-like with 7–13 alternately long and short denticles decreasing in size from the first lateral tooth toward the 5
<superScript id="DF02D5AC784CE834CAB8E51FF8D933D0">th</superScript>
lateral tooth. Marginal teeth two teeth, claw-like with 12–14 equal size denticles. Radula formula 2: 5: 1 + 1 + 1 + 1 + 1 + 1: 5: 2.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWXBG" type="Differential diagnosis">
<paragraph id="D50F6FE33DF124DCA91258C6695CCE1A">
<heading id="956723E492DA5BB556A1280785A00900" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="026E562140784D6FBFDBF54E4AC0DF55">
<taxonomicName id="59F2E2ED6B5953A6345E06252707D3CD" authorityName="Klongklaew &amp; Poeaim &amp; Dumrongrojwattana" authorityYear="2024" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="sagitta" status="sp. nov.">
<emphasis id="1E6EA6CD900DE9D6FCBC47EA9C96F894" italics="true">Georissa sagitta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82852D897EDA9B1FF10DC0E5E509CA0D" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="36FB2A4D602241CCEA2CEB5078A50714" authority="Godwin-Austen &amp; Neville, 1879" authorityName="Godwin-Austen &amp; Neville" authorityYear="1879" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="monterosatiana">
<emphasis id="842FCEF3015C81257E7CC768B17B6E67" italics="true">G. monterosatiana</emphasis>
Godwin-Austen &amp; Neville, 1879
</taxonomicName>
.
<taxonomicName id="5CC588234C7879DC76551D04561A6154" authorityName="Godwin-Austen &amp; Neville" authorityYear="1879" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" phylum="Mollusca" rank="species" species="monterosatiana">
<emphasis id="8A920539E9EE908CAF682EF978B3C615" italics="true">Georissa monterosatiana</emphasis>
</taxonomicName>
differs in having more convex body whorl, smooth protoconch, and more lirae (9–11 instead of 8–9 lirae).
</paragraph>
</subSubSection>
</treatment>
</document>
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