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<document id="1FEF9D1D5BFD44A9472B46EF822AE69B" ID-DOI="10.11646/phytotaxa.135.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="5086131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625843137740" checkinUser="felipe" docAuthor="Henderson, Andrew &amp; Dung, Nguyen Quoc" docDate="2013" docId="CE7B87D9FFB8FFAD03E5FDFDFA83FC04" docLanguage="en" docName="Phytotaxa.135.1.19-26.pdf" docOrigin="Phytotaxa 135 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.135.1.3" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Calamus phuocbinhensis Henderson &amp; N. Q. Dung 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="24" masterDocId="3242FFA1FFBCFFA80372FFE7FF9EFFDA" masterDocTitle="Four new species of Calamus (Arecaceae) from Vietnam" masterLastPageNumber="26" masterPageNumber="19" pageNumber="23" updateTime="1698997002057" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="D9FD51CF64A65ACA583F1155F83D0F82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="971B17A6B5C5D4766D483AB1A9BFCC06">
<mods:title id="76979F1E828081D50D1F3BEB64BA33F7">Four new species of Calamus (Arecaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="BC5F57075A9FDE38A5A2F9C2729E9A94" type="personal">
<mods:role id="007A833DE1010B24F762F84CD0C58763">
<mods:roleTerm id="7ED871285D2130B587AD288E1BD024FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A31AB367F11509C41BCEEE47AF81ABCE">Henderson, Andrew</mods:namePart>
<mods:affiliation id="E1A73D3D6F5D7D42D97AFC8911FD2D4C">Institute of Systematic Botany, The New York Botanical Garden, Bronx, NY 10458 - 5126, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="0E9A047A1D133E427B8B1A0DE6CD8027" type="personal">
<mods:role id="2A4826DBE66B4C88057A585F32F7B2A1">
<mods:roleTerm id="0A776E35AC88D7C1833ECB3FB109540F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E7F74E05D067798AE7957DE50F21B13">Dung, Nguyen Quoc</mods:namePart>
<mods:affiliation id="4BED9A1C74DFE355C524C5DFC4F9EA2F">Forest Inventory and Planning Institute, Thanh Tri, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource id="486D6C8E696F53CA7913D6322A46F8AC">text</mods:typeOfResource>
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<mods:titleInfo id="F5E1A1C0E3B1CA574F904F9422D1325B">
<mods:title id="E15D4A06B0A242A3CD95E85B949BE256">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="E7DB05467930D4BFA540BF554BE299F3">
<mods:date id="E5AFD6C8015F2C02BC9EB52FAB9A5CDF">2013</mods:date>
<mods:detail id="674082688A9516F07E435B5087C1382A" type="pubDate">
<mods:number id="E6F7B66F90AC654F87ACD48A4D2D84E4">2013-10-01</mods:number>
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<mods:number id="4E57E7F6C19980C3C186A6C6D6B5F0FC">135</mods:number>
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<mods:number id="1EB521DD4E39CE303D0A460C28CB2182">1</mods:number>
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<mods:start id="F1068E82BE0F4A2FDFF2B078A5ABA481">19</mods:start>
<mods:end id="C65DBA5F7C3ADD8DE57418D39BE1D3AD">26</mods:end>
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<mods:url id="96373575715C2D564675049CFC814681">http://dx.doi.org/10.11646/phytotaxa.135.1.3</mods:url>
</mods:location>
<mods:classification id="9E4811A3FDE810644B50B67396DE3CBD">journal article</mods:classification>
<mods:identifier id="9238FD2C868B823DF65B20F0857F36AF" type="DOI">10.11646/phytotaxa.135.1.3</mods:identifier>
<mods:identifier id="99EBD88B5B886A570DA44E391C9F6714" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="6D92F9D4B60C0BAF5953749A68DC9865" type="Zenodo-Dep">5086131</mods:identifier>
</mods:mods>
<treatment id="CE7B87D9FFB8FFAD03E5FDFDFA83FC04" ID-DOI="http://doi.org/10.5281/zenodo.5085955" ID-Zenodo-Dep="5085955" LSID="urn:lsid:plazi:treatment:CE7B87D9FFB8FFAD03E5FDFDFA83FC04" httpUri="http://treatment.plazi.org/id/CE7B87D9FFB8FFAD03E5FDFDFA83FC04" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
<subSubSection id="0EC86544FFB8FFAC03E5FDFDFAC5FDA2" pageId="4" pageNumber="23" type="nomenclature">
<paragraph id="466D36CFFFB8FFAC03E5FDFDFC31FDEE" blockId="4.[151,943,538,564]" box="[151,943,538,564]" pageId="4" pageNumber="23">
<heading id="1D2581A3FFB8FFAC03E5FDFDFC31FDEE" box="[151,943,538,564]" fontSize="11" level="2" pageId="4" pageNumber="23" reason="3">
<taxonomicName id="81D24D4CFFB8FFAC03E5FDFDFD6CFDEE" authority="Henderson &amp; N. Q. Dung" authorityName="Henderson &amp; N. Q. Dung" authorityYear="2013" box="[151,754,538,564]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="phuocbinhensis" status="sp. nov.">
<emphasis id="74A6EADDFFB8FFAC03E5FDFDFE5DFDEE" bold="true" box="[151,451,538,564]" italics="true" pageId="4" pageNumber="23">Calamus phuocbinhensis</emphasis>
Henderson &amp; N. Q. Dung
</taxonomicName>
<emphasis id="74A6EADDFFB8FFAC018BFDFDFCCCFDE9" box="[761,850,538,563]" italics="true" pageId="4" pageNumber="23">
<taxonomicNameLabel id="6F9557A6FFB8FFAC018BFDFDFCCCFDE9" box="[761,850,538,563]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="DEE92A4AFFB8FFAC0013FDFDFC3AFDEE" box="[865,932,538,564]" captionStart="FIGURE 3" captionStartId="4.[151,246,1774,1796]" captionTargetBox="[151,1436,782,1749]" captionTargetId="figure-366@4.[151,1436,782,1749]" captionTargetPageId="4" captionText="FIGURE 3. Calamus phuocbinhensis. A. Leaf sheath with lack of tomentum, few spines, and small ocrea. B. Pinnae, irregularly arranged and spreading in different planes. (A–B from Henderson &amp; Nguyen Quoc Dung 3853)." figureDoi="http://doi.org/10.5281/zenodo.5086137" httpUri="https://zenodo.org/record/5086137/files/figure.png" pageId="4" pageNumber="23">Fig. 3</figureCitation>
)
</heading>
</paragraph>
<paragraph id="466D36CFFFB8FFAC03E5FDA6FAC5FDA2" blockId="4.[151,1436,577,728]" pageId="4" pageNumber="23">It differs from related species by its leaf sheaths without tomentum and with few spines, its rachises with recurved spines abaxially, and its short, membranous ocreas consisting of two lateral flanges, not developed above the petiole.</paragraph>
</subSubSection>
<subSubSection id="0EC86544FFB8FFAD03E5FD66FB0BFEFC" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23" type="materials_examined">
<paragraph id="466D36CFFFB8FFAC03E5FD66FE34FD02" blockId="4.[151,1436,577,728]" pageId="4" pageNumber="23">
<materialsCitation id="F6BA3C92FFB8FFAC03E5FD66FE38FD02" collectingDate="2013-04-18" collectionCode="VFM" collectorName="A. Henderson &amp; Nguyen Quoc Dung" country="Vietnam" elevation="323" latitude="11.986" location="Phuoc Binh National Park" longLatPrecision="77" longitude="108.747" municipality="Type" pageId="4" pageNumber="23" specimenCount="1" stateProvince="Ninh Thuan" typeStatus="holotype">
<collectingMunicipality id="A609ACB5FFB8FFAC03E5FD66FF50FD42" box="[151,206,641,664]" pageId="4" pageNumber="23">Type</collectingMunicipality>
:—
<collectingCountry id="3EC5765FFFB8FFAC039FFD66FEF2FD42" box="[237,364,641,664]" name="Vietnam" pageId="4" pageNumber="23">VIETNAM</collectingCountry>
.
<collectingRegion id="8416F82DFFB8FFAC0208FD66FD9FFD42" box="[378,513,641,664]" country="Vietnam" name="Ninh Thuan" pageId="4" pageNumber="23">Ninh Thuan</collectingRegion>
:
<location id="430D6014FFB8FFAC017DFD66FCAAFD42" LSID="urn:lsid:plazi:treatment:CE7B87D9FFB8FFAD03E5FDFDFA83FC04:430D6014FFB8FFAC017DFD66FCAAFD42" box="[527,820,641,664]" country="Vietnam" latitude="11.986" longLatPrecision="77" longitude="108.747" municipality="Type" name="Phuoc Binh National Park" pageId="4" pageNumber="23" stateProvince="Ninh Thuan">Phuoc Binh National Park</location>
, near park headquarters, disturbed forest by ecotourism trail,
<geoCoordinate id="23E65008FFB8FFAC0270FD46FEFEFD62" box="[258,352,673,696]" degrees="11.986" direction="north" orientation="latitude" pageId="4" pageNumber="23" precision="55" value="11.986">11.986N</geoCoordinate>
,
<geoCoordinate id="23E65008FFB8FFAC021FFD46FE49FD62" box="[365,471,673,696]" degrees="108.747" direction="east" orientation="longitude" pageId="4" pageNumber="23" precision="55" value="108.747">108.747E</geoCoordinate>
,
<quantity id="812A9B2AFFB8FFAC0296FD46FDB5FD62" box="[484,555,673,696]" metricMagnitude="2" metricUnit="m" metricValue="3.2299999999999995" pageId="4" pageNumber="23" unit="m" value="323.0">
<elevation id="CDFFD1FCFFB8FFAC0296FD46FDB5FD62" box="[484,555,673,696]" metricMagnitude="2" metricUnit="m" metricValue="3.2299999999999995" pageId="4" pageNumber="23" unit="m" value="323.0">323 m</elevation>
</quantity>
elevation,
<date id="326C100FFFB8FFAC01D4FD46FCDAFD62" box="[678,836,673,696]" pageId="4" pageNumber="23" value="2013-04-18">
<collectingDate id="2228E9E7FFB8FFAC01D4FD46FCDAFD62" box="[678,836,673,696]" pageId="4" pageNumber="23" value="2013-04-18">18 April 2013</collectingDate>
</date>
,
<emphasis id="74A6EADDFFB8FFAC0023FD46FABBFD62" box="[849,1317,673,696]" italics="true" pageId="4" pageNumber="23">
<collectorName id="EB275319FFB8FFAC0023FD46FC78FD62" box="[849,998,673,696]" pageId="4" pageNumber="23">A. Henderson</collectorName>
&amp;
<collectorName id="EB275319FFB8FFAC077FFD46FB79FD62" box="[1037,1255,673,696]" pageId="4" pageNumber="23">Nguyen Quoc Dung</collectorName>
3857
</emphasis>
(
<typeStatus id="9969886DFFB8FFAC0644FD46FA06FD62" box="[1334,1432,673,696]" pageId="4" pageNumber="23" type="holotype">holotype</typeStatus>
,
<collectionCode id="20C3AE0AFFB8FFAC03B5FD26FE98FD02" box="[199,262,705,728]" pageId="4" pageNumber="23">VFM</collectionCode>
! isotype, NY!)
</materialsCitation>
.
</paragraph>
<caption id="12AD6647FFB8FFAC03E5F909FBAFF8FD" ID-DOI="http://doi.org/10.5281/zenodo.5086137" ID-Zenodo-Dep="5086137" httpUri="https://zenodo.org/record/5086137/files/figure.png" pageId="4" pageNumber="23" startId="4.[151,246,1774,1796]" targetBox="[151,1436,782,1749]" targetPageId="4">
<paragraph id="466D36CFFFB8FFAC03E5F909FBAFF8FD" blockId="4.[151,1435,1774,1831]" pageId="4" pageNumber="23">
<emphasis id="74A6EADDFFB8FFAC03E5F909FE97F8DE" bold="true" box="[151,265,1774,1796]" pageId="4" pageNumber="23">FIGURE 3</emphasis>
.
<taxonomicName id="81D24D4CFFB8FFAC0264F909FD91F8D9" authorityName="Henderson &amp; N. Q. Dung" authorityYear="2013" box="[278,527,1774,1795]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="phuocbinhensis">
<emphasis id="74A6EADDFFB8FFAC0264F909FD91F8D9" box="[278,527,1774,1795]" italics="true" pageId="4" pageNumber="23">Calamus phuocbinhensis</emphasis>
</taxonomicName>
.
<emphasis id="74A6EADDFFB8FFAC016EF909FDB3F8DE" bold="true" box="[540,557,1774,1796]" pageId="4" pageNumber="23">A</emphasis>
. Leaf sheath with lack of tomentum, few spines, and small ocrea.
<emphasis id="74A6EADDFFB8FFAC07B7F909FB4BF8DE" bold="true" box="[1221,1237,1774,1796]" pageId="4" pageNumber="23">B</emphasis>
. Pinnae, irregularly arranged and spreading in different planes. (A–B from
<emphasis id="74A6EADDFFB8FFAC01D5F8F5FBBAF8FD" box="[679,1060,1810,1831]" italics="true" pageId="4" pageNumber="23">Henderson &amp; Nguyen Quoc Dung 3853</emphasis>
).
</paragraph>
</caption>
<paragraph id="466D36CFFFB8FFAD03E5F8BCFB0BFEFC" blockId="4.[151,1436,1883,2025]" lastBlockId="5.[151,1436,152,990]" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
Stems clustered,
<quantity id="812A9B2AFFB8FFAC0219F8BCFE59F8AF" box="[363,455,1883,1909]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="4" pageNumber="23" unit="m" value="11.5" valueMax="15.0" valueMin="8.0">8–15 m</quantity>
long,
<quantity id="812A9B2AFFB8FFAC016AF8BCFD3FF8AF" box="[536,673,1883,1909]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.1" metricValueMin="1.3" pageId="4" pageNumber="23" unit="cm" value="1.7" valueMax="2.1" valueMin="1.3">1.3–2.1 cm</quantity>
diameter with sheaths,
<quantity id="812A9B2AFFB8FFAC00B9F8BCFBCAF8AF" box="[971,1108,1883,1909]" metricMagnitude="-2" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="4" pageNumber="23" unit="cm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 cm</quantity>
diameter without sheaths. Sheaths completely green, with very few, flat, brown spines to
<quantity id="812A9B2AFFB8FFAC00D9F865FC60F846" box="[939,1022,1922,1948]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="23" unit="cm" value="1.5">1.5 cm</quantity>
long; ocreas short, membranous, consisting of two lateral flanges, not developed above petiole, soon falling; flagella
<quantity id="812A9B2AFFB8FFAC07D9F84FFA9BF818" box="[1195,1285,1960,1986]" metricMagnitude="0" metricUnit="m" metricValue="1.65" pageId="4" pageNumber="23" unit="cm" value="165.0">165 cm</quantity>
long; knees present; cirri absent; petioles
<quantity id="812A9B2AFFB8FFAC029FF828FDDCF833" box="[493,578,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="4" pageNumber="23" unit="cm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 cm</quantity>
long on upper leaves, to
<quantity id="812A9B2AFFB8FFAC0014F828FC30F833" box="[870,942,1999,2025]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="23" unit="cm" value="24.0">24 cm</quantity>
long on lower leaves; rachises
<quantity id="812A9B2AFFB8FFAC0668F828FA02F833" box="[1306,1436,1999,2025]" metricMagnitude="0" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.25" metricValueMin="0.97" pageId="4" pageNumber="23" unit="cm" value="111.0" valueMax="125.0" valueMin="97.0">97–125 cm</quantity>
long with 44–45 pinnae per side, these irregularly arranged in clusters and spreading in different planes, linear, minutely spiny along the margins and mid-vein adaxially; middle pinnae
<quantity id="812A9B2AFFB9FFAD077CFF59FB2FFF02" box="[1038,1201,190,216]" metricMagnitude="-1" metricUnit="m" metricValue="2.44" metricValueMax="2.95" metricValueMin="1.93" pageId="5" pageNumber="24" unit="cm" value="24.4" valueMax="29.5" valueMin="19.3">19.3–29.5 cm</quantity>
long,
<quantity id="812A9B2AFFB9FFAD078DFF59FACFFF02" box="[1279,1361,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="24" unit="cm" value="1.2">1.2 cm</quantity>
wide; apical pair of pinnae joined for
<quantity id="812A9B2AFFB9FFAD017DFF02FD0BFF25" box="[527,661,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="5" pageNumber="24" unit="cm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 cm</quantity>
. Staminate inflorescences not seen; pistillate inflorescence rachis bracts tubular, terminating below the partial inflorescences; flowers and fruits not seen.
</paragraph>
</subSubSection>
<subSubSection id="0EC86544FFB9FFAD03B4FED5FD39FDD4" pageId="5" pageNumber="24" type="distribution">
<paragraph id="466D36CFFFB9FFAD03B4FED5FEC5FEA9" blockId="5.[151,1436,152,990]" pageId="5" pageNumber="24">
<emphasis id="74A6EADDFFB9FFAD03B4FED5FDBEFE96" bold="true" box="[198,544,306,332]" pageId="5" pageNumber="24">Distribution and habitat:—</emphasis>
Endemic to southern
<collectingCountry id="3EC5765FFFB9FFAD006CFED5FC1FFE96" box="[798,897,306,332]" name="Vietnam" pageId="5" pageNumber="24">Vietnam</collectingCountry>
in
<collectingRegion id="8416F82DFFB9FFAD00DBFED5FB39FE96" box="[937,1191,306,332]" country="Vietnam" name="Ninh Thuan" pageId="5" pageNumber="24">Ninh Thuan Province</collectingRegion>
in seasonal forest at
<quantity id="812A9B2AFFB9FFAD03E5FEBEFF7FFEA9" box="[151,225,345,371]" metricMagnitude="2" metricUnit="m" metricValue="3.2299999999999995" pageId="5" pageNumber="24" unit="m" value="323.0">323 m</quantity>
elevation.
</paragraph>
<paragraph id="466D36CFFFB9FFAD03B4FE67FCAAFE40" blockId="5.[151,1436,152,990]" box="[198,820,384,410]" pageId="5" pageNumber="24">
<emphasis id="74A6EADDFFB9FFAD03B4FE67FE62FE40" bold="true" box="[198,508,384,410]" pageId="5" pageNumber="24">Local names and uses:—</emphasis>
<emphasis id="74A6EADDFFB9FFAD028EFE67FDADFE43" box="[508,563,384,409]" italics="true" pageId="5" pageNumber="24">may.</emphasis>
No uses are reported.
</paragraph>
<paragraph id="466D36CFFFB9FFAD03B4FE41FD39FDD4" blockId="5.[151,1436,152,990]" pageId="5" pageNumber="24">
<emphasis id="74A6EADDFFB9FFAD03B4FE41FD1AFE1A" bold="true" box="[198,644,422,448]" pageId="5" pageNumber="24">Additional specimen examined:—</emphasis>
<materialsCitation id="F6BA3C92FFB9FFAD01F6FE41FD3DFDD4" collectingDate="2013-04-18" collectionCode="AAU, VFM" collectorName="A. Henderson &amp; Nguyen Quoc Dung" country="Vietnam" elevation="323" latitude="11.986" location="Phuoc Binh National Park" longLatPrecision="77" longitude="108.747" pageId="5" pageNumber="24" specimenCount="1" stateProvince="Ninh Thuan">
<collectingCountry id="3EC5765FFFB9FFAD01F6FE41FD75FE1A" box="[644,747,422,448]" name="Vietnam" pageId="5" pageNumber="24">Vietnam</collectingCountry>
.
<collectingRegion id="8416F82DFFB9FFAD0073FE41FC03FE1A" box="[769,925,422,448]" country="Vietnam" name="Ninh Thuan" pageId="5" pageNumber="24">Ninh Thuan</collectingRegion>
:
<location id="430D6014FFB9FFAD00C6FE41FA92FE1A" LSID="urn:lsid:plazi:treatment:CE7B87D9FFB8FFAD03E5FDFDFA83FC04:430D6014FFB9FFAD00C6FE41FA92FE1A" box="[948,1292,422,448]" country="Vietnam" latitude="11.986" longLatPrecision="77" longitude="108.747" name="Phuoc Binh National Park" pageId="5" pageNumber="24" stateProvince="Ninh Thuan">Phuoc Binh National Park</location>
, near park headquarters, disturbed forest by ecotourism trail,
<geoCoordinate id="23E65008FFB9FFAD0191FE2AFCD4FE3D" box="[739,842,461,487]" degrees="11.986" direction="north" orientation="latitude" pageId="5" pageNumber="24" precision="55" value="11.986">11.986N</geoCoordinate>
,
<geoCoordinate id="23E65008FFB9FFAD0027FE2AFC57FE3D" box="[853,969,461,487]" degrees="108.747" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="55" value="108.747">108.747E</geoCoordinate>
,
<quantity id="812A9B2AFFB9FFAD00A6FE2AFBBEFE3D" box="[980,1056,461,487]" metricMagnitude="2" metricUnit="m" metricValue="3.2299999999999995" pageId="5" pageNumber="24" unit="m" value="323.0">
<elevation id="CDFFD1FCFFB9FFAD00A6FE2AFBBEFE3D" box="[980,1056,461,487]" metricMagnitude="2" metricUnit="m" metricValue="3.2299999999999995" pageId="5" pageNumber="24" unit="m" value="323.0">323 m</elevation>
</quantity>
,
<date id="326C100FFFB9FFAD075EFE2AFB4CFE3D" box="[1068,1234,461,487]" pageId="5" pageNumber="24" value="2013-04-18">
<collectingDate id="2228E9E7FFB9FFAD075EFE2AFB4CFE3D" box="[1068,1234,461,487]" pageId="5" pageNumber="24" value="2013-04-18">18 April 2013</collectingDate>
</date>
,
<emphasis id="74A6EADDFFB9FFAD07ACFE2BFE5CFDD7" italics="true" pageId="5" pageNumber="24">
<collectorName id="EB275319FFB9FFAD07ACFE2BFAE1FE3C" box="[1246,1407,460,486]" pageId="5" pageNumber="24">A. Henderson</collectorName>
&amp;
<collectorName id="EB275319FFB9FFAD03E5FE13FEE1FDD7" box="[151,383,499,525]" pageId="5" pageNumber="24">Nguyen Quoc Dung</collectorName>
3853
</emphasis>
(
<collectionCode id="20C3AE0AFFB9FFAD02A3FE13FD8BFDD4" box="[465,533,500,526]" country="Ethiopia" httpUri="http://grbio.org/cool/5dxx-xet2" name="Addis Ababa University, Department of Biology" pageId="5" pageNumber="24">AAU</collectionCode>
, NY,
<collectionCode id="20C3AE0AFFB9FFAD0124FE13FD05FDD4" box="[598,667,500,526]" pageId="5" pageNumber="24">VFM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0EC86544FFB9FFAD03B4FDFDFA83FC04" pageId="5" pageNumber="24" type="discussion">
<paragraph id="466D36CFFFB9FFAD03B4FDFDFA83FC04" blockId="5.[151,1436,152,990]" pageId="5" pageNumber="24">
<emphasis id="74A6EADDFFB9FFAD03B4FDFDFEEEFDEE" bold="true" box="[198,368,538,564]" pageId="5" pageNumber="24">Discussion:—</emphasis>
Although the
<specimenCount id="50D4FD46FFB9FFAD0163FDFDFD23FDEE" box="[529,701,538,564]" pageId="5" pageNumber="24" type="generic">two specimens</specimenCount>
are incomplete and lack both flowers and fruits, and with only an old, decaying inflorescence present, we have no doubt that they represent an undescribed species. In fact, the sheaths and leaves of this species are the most distinctive and easily recognized of all Vietnamese palms. In Evans
<emphasis id="74A6EADDFFB9FFAD0399FD69FEB4FD7D" box="[235,298,653,679]" italics="true" pageId="5" pageNumber="24">et al.</emphasis>
’s (2001a) treatment of
<taxonomicName id="81D24D4CFFB9FFAD0127FD6AFD5CFD7D" authorityName="Linnaeus" authorityYear="1753" box="[597,706,653,679]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="74A6EADDFFB9FFAD0127FD6AFD5CFD7D" box="[597,706,653,679]" italics="true" pageId="5" pageNumber="24">Calamus</emphasis>
</taxonomicName>
in neighboring
<collectingCountry id="3EC5765FFFB9FFAD00FFFD69FC52FD72" box="[909,972,654,680]" name="Laos" pageId="5" pageNumber="24">Laos</collectingCountry>
, this palm would key to
<taxonomicName id="81D24D4CFFB9FFAD0676FD6AFE34FD15" authority="Willdenow (1799: 203)" authorityName="Willdenow" authorityPageNumber="203" authorityYear="1799" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="viminalis">
<emphasis id="74A6EADDFFB9FFAD0676FD6AFA02FD7D" box="[1284,1436,653,679]" italics="true" pageId="5" pageNumber="24">C. viminalis</emphasis>
<bibRefCitation id="22434B3EFFB9FFAD03E5FD52FE34FD15" author="Willdenow, K." box="[151,426,693,719]" pageId="5" pageNumber="24" refId="ref4285" refString="Willdenow, K. (1799) Species plantarum. Volume II. Berolini. Impensis G. C. Nauk, 1340 pp." type="book" year="1799">Willdenow (1799: 203)</bibRefCitation>
</taxonomicName>
, based on its presence of a flagellum and irregularly arranged pinnae lying in different planes. In the same study of
<taxonomicName id="81D24D4CFFB9FFAD0282FD3CFDC7FD2F" authorityName="Linnaeus" authorityYear="1753" box="[496,601,731,757]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="74A6EADDFFB9FFAD0282FD3CFDC7FD2F" box="[496,601,731,757]" italics="true" pageId="5" pageNumber="24">Calamus</emphasis>
</taxonomicName>
cited above (Henderson, in prep.),
<taxonomicName id="81D24D4CFFB9FFAD0777FD3CFB7CFD2F" authorityName="Henderson &amp; N. Q. Dung" authorityYear="2013" box="[1029,1250,731,757]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="phuocbinhensis">
<emphasis id="74A6EADDFFB9FFAD0777FD3CFB7CFD2F" box="[1029,1250,731,757]" italics="true" pageId="5" pageNumber="24">C. phuocbinhensis</emphasis>
</taxonomicName>
(named for the National Park in which it occurs) is placed in a clade comprising two other species;
<taxonomicName id="81D24D4CFFB9FFAD07DFFCE6FADCFCC1" authorityName="Willdenow" authorityYear="1799" box="[1197,1346,769,795]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="viminalis">
<emphasis id="74A6EADDFFB9FFAD07DFFCE6FADCFCC1" box="[1197,1346,769,795]" italics="true" pageId="5" pageNumber="24">C. viminalis</emphasis>
</taxonomicName>
and
<taxonomicName id="81D24D4CFFB9FFAD06F3FCE6FD9CFC99" authority="Beccari (1902: 203)" authorityName="Beccari" authorityPageNumber="203" authorityYear="1902" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="74A6EADDFFB9FFAD06F3FCE6FE95FC98" italics="true" pageId="5" pageNumber="24">C. siamensis</emphasis>
<bibRefCitation id="22434B3EFFB9FFAD0261FCCEFD9CFC99" author="Beccari, O." box="[275,514,809,835]" pageId="5" pageNumber="24" pagination="197 - 230" refId="ref3642" refString="Beccari, O. (1902) Systematic enumeration of the species of Calamus and Daemonorops, with diagnoses of the new ones. Records of the Botanical Survey of India 2: 197 - 230." type="journal article" year="1902">Beccari (1902: 203)</bibRefCitation>
</taxonomicName>
, characterized by the rachis with straight spines abaxially and the very short staminate rachillae with the flowers arranged in glomerules.
<taxonomicName id="81D24D4CFFB9FFAD0004FCA8FB3DFCB3" authorityName="Henderson &amp; N. Q. Dung" authorityYear="2013" box="[886,1187,847,873]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="phuocbinhensis">
<emphasis id="74A6EADDFFB9FFAD0004FCA8FB3DFCB3" box="[886,1187,847,873]" italics="true" pageId="5" pageNumber="24">Calamus phuocbinhensis</emphasis>
</taxonomicName>
differs from both of these in its leaf sheaths without tomentum and with few spines, its rachises with recurved spines abaxially, and in particular its short, membranous ocreas consisting of two lateral flanges, not developed above the petiole. In both
<taxonomicName id="81D24D4CFFB9FFAD03A0FC24FEC1FC07" authorityName="Willdenow" authorityYear="1799" box="[210,351,963,989]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="viminalis">
<emphasis id="74A6EADDFFB9FFAD03A0FC24FEC1FC07" box="[210,351,963,989]" italics="true" pageId="5" pageNumber="24">C. viminalis</emphasis>
</taxonomicName>
and
<taxonomicName id="81D24D4CFFB9FFAD02E5FC24FDB4FC07" authorityName="Beccari" authorityYear="1902" box="[407,554,963,989]" class="Actinopterygii" family="Sparidae" genus="Calamus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="74A6EADDFFB9FFAD02E5FC24FDB4FC07" box="[407,554,963,989]" italics="true" pageId="5" pageNumber="24">C. siamensis</emphasis>
</taxonomicName>
the ocrea is completely tubular and developed above the petiole.
</paragraph>
</subSubSection>
</treatment>
</document>
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