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<document id="AF5371C1B6DD302D7F45E52EBAC3EC22" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC1FFF88509F3863BEC1FEC9" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus riedeli Bocak 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="516" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="515" updateTime="1698468596363" updateUser="ExternalLinkService">
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<mods:title id="FB1DB26B57DE2CCFEB3B4D0D127DD78E">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
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<mods:namePart id="E04F3D12A49CCE1EEE5302346505768D">Bocak, Ladislav</mods:namePart>
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<mods:date id="0F4D02A8FF69A911B880A69B2B2A8CBF">2017</mods:date>
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<mods:number id="2E4CE6AA847DF79C570293B85A183697">2017-09-18</mods:number>
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<treatment id="A312190FEC1FFF88509F3863BEC1FEC9" ID-DOI="http://doi.org/10.5281/zenodo.6018093" ID-GBIF-Taxon="134090571" ID-Zenodo-Dep="6018093" LSID="urn:lsid:plazi:treatment:A312190FEC1FFF88509F3863BEC1FEC9" httpUri="http://treatment.plazi.org/id/A312190FEC1FFF88509F3863BEC1FEC9" lastPageId="11" lastPageNumber="516" pageId="10" pageNumber="515">
<subSubSection id="63A1FB92EC1FFF89509F3863BAF0FA7A" pageId="10" pageNumber="515" type="nomenclature">
<paragraph id="2B04A819EC1FFF89509F3863BAB0FA59" blockId="10.[151,492,1394,1454]" box="[151,492,1394,1421]" pageId="10" pageNumber="515">
<heading id="704C1F75EC1FFF89509F3863BAB0FA59" bold="true" box="[151,492,1394,1421]" fontSize="11" level="1" pageId="10" pageNumber="515" reason="1">
<emphasis id="19CF740BEC1FFF89509F3863BAB0FA59" bold="true" box="[151,492,1394,1421]" pageId="10" pageNumber="515">
<taxonomicName id="ECBBD39AEC1FFF89509F3863BADBFA58" authority="Bocak, 2017" authorityName="Bocak" authorityYear="2017" box="[151,391,1394,1420]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="19CF740BEC1FFF89509F3863BADBFA58" bold="true" box="[151,391,1394,1420]" italics="true" pageId="10" pageNumber="515">Cladophorus riedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02FCC970EC1FFF8951863862BAB0FA59" box="[398,492,1395,1421]" pageId="10" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2B04A819EC1FFF89509F3884BAF0FA7A" blockId="10.[151,492,1394,1454]" box="[151,428,1429,1454]" pageId="10" pageNumber="515">
(
<figureCitation id="B380B49CEC1FFF8950973884BA7AFA7A" box="[159,294,1429,1454]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="10" pageNumber="515">Figs. 16, 31</figureCitation>
,
<figureCitation id="B380B49CEC1FFF89513A3887BA0DFA7A" box="[306,337,1430,1454]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="10" pageNumber="515">48</figureCitation>
,
<figureCitation id="B380B49CEC1FFF8951553884BAF8FA7A" box="[349,420,1429,1454]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="10" pageNumber="515">7576</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1FFF89509F38CCBECEF9CE" pageId="10" pageNumber="515" type="materials_examined">
<paragraph id="2B04A819EC1FFF89509F38CCBECEF9CE" blockId="10.[151,1436,1501,2030]" pageId="10" pageNumber="515">
<materialsCitation id="9BD3A244EC1FFF89509F38CCBA39F9CE" ID-GBIF-Occurrence="1638540600" collectionCode="LMBC" collectorName="Pt. Moresby" country="Papua New Guinea" county="Berem" elevation="700" latitude="-9.433333" location="Male" longLatPrecision="1299" longitude="147.55" municipality="Kailaki" pageId="10" pageNumber="515" specimenCount="2" typeStatus="holotype">
<emphasis id="19CF740BEC1FFF89509F38CCBA18FA22" bold="true" box="[151,324,1501,1526]" pageId="10" pageNumber="515">
<typeStatus id="F40016BBEC1FFF89509F38CCBB8EFA22" box="[151,210,1501,1526]" pageId="10" pageNumber="515">Type</typeStatus>
material.
</emphasis>
<typeStatus id="F40016BBEC1FFF89514338CCBAEBFA22" box="[331,439,1501,1526]" pageId="10" pageNumber="515" type="holotype">Holotype</typeStatus>
.
<location id="2E64FEC2EC1FFF8951C938CCBAA1FA22" LSID="urn:lsid:plazi:treatment:A312190FEC1FFF88509F3863BEC1FEC9:2E64FEC2EC1FFF8951C938CCBAA1FA22" box="[449,509,1501,1526]" country="Papua New Guinea" county="Berem" latitude="-9.433333" longLatPrecision="1299" longitude="147.55" municipality="Kailaki" name="Male" pageId="10" pageNumber="515">Male</location>
,
<collectingCountry id="53ACE889EC1FFF89520038CFB9BFFA22" box="[520,739,1501,1526]" name="Papua New Guinea" pageId="10" pageNumber="515">Papua New Guinea</collectingCountry>
,
<collectorName id="864ECDCFEC1FFF8952E638CFB82BFA22" box="[750,887,1501,1526]" pageId="10" pageNumber="515">Pt. Moresby</collectorName>
,
<collectingMunicipality id="CB603263EC1FFF89538A38CCB88BFA22" box="[898,983,1501,1526]" pageId="10" pageNumber="515">Kailaki</collectingMunicipality>
,
<collectingCounty id="C265D095EC1FFF8953EA38CFBF6CFA21" box="[994,1072,1502,1525]" pageId="10" pageNumber="515">Berem</collectingCounty>
,
<geoCoordinate id="4E8FCEDEEC1FFF89543338CCBFC5FA22" box="[1083,1177,1501,1526]" direction="south" orientation="latitude" pageId="10" pageNumber="515" precision="925" value="-9.433333">09°26´S</geoCoordinate>
,
<geoCoordinate id="4E8FCEDEEC1FFF8954AC38CFBE4EFA22" box="[1188,1298,1502,1526]" direction="east" orientation="longitude" pageId="10" pageNumber="515" precision="925" value="147.55">147°33´E</geoCoordinate>
,
<elevation id="A0964F2AEC1FFF89551638CCBEC0FA21" box="[1310,1436,1501,1526]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="10" pageNumber="515" unit="m" value="700.0" valueMax="850.0" valueMin="550.0">
<quantity id="EC4305FCEC1FFF89551638CCBEC0FA21" box="[1310,1436,1501,1526]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="10" pageNumber="515" unit="m" value="700.0" valueMax="850.0" valueMin="550.0">550850 m</quantity>
</elevation>
(A00121,
<collectionCode id="4DAA30DCEC1FFF8951003B13BA00F9CE" box="[264,348,1538,1562]" pageId="10" pageNumber="515">LMBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9BD3A244EC1FFF8951673B13BECEF9CE" ID-GBIF-Occurrence="1638540592" box="[367,1426,1537,1562]" collectionCode="LMBC" collectorName="Pt. Moresby &amp; Varirata N. P." country="Papua New Guinea" elevation="750" latitude="-9.433333" location="Papua New Guinea" longLatPrecision="1299" longitude="147.35" pageId="10" pageNumber="515" specimenCount="1" typeStatus="paratype">
<typeStatus id="F40016BBEC1FFF8951673B13BA89F9CE" box="[367,469,1538,1562]" pageId="10" pageNumber="515" type="paratype">Paratype</typeStatus>
.
<collectingCountry id="53ACE889EC1FFF8951D73B13B9E6F9CE" box="[479,698,1537,1562]" name="Papua New Guinea" pageId="10" pageNumber="515">Papua New Guinea</collectingCountry>
,
<collectorName id="864ECDCFEC1FFF8952CD3B13B80CF9CE" box="[709,848,1537,1562]" pageId="10" pageNumber="515">Pt. Moresby</collectorName>
,
<collectorName id="864ECDCFEC1FFF8953523B10B8AEF9CD" box="[858,1010,1537,1562]" pageId="10" pageNumber="515">Varirata N. P.</collectorName>
,
<geoCoordinate id="4E8FCEDEEC1FFF8953F63B10BF06F9CE" box="[1022,1114,1537,1562]" direction="south" orientation="latitude" pageId="10" pageNumber="515" precision="925" value="-9.433333">09°26´S</geoCoordinate>
<geoCoordinate id="4E8FCEDEEC1FFF8954693B13BF93F9CE" box="[1121,1231,1538,1562]" direction="east" orientation="longitude" pageId="10" pageNumber="515" precision="925" value="147.35">147°21´E</geoCoordinate>
,
<elevation id="A0964F2AEC1FFF8954D23B10BE7DF9CD" box="[1242,1313,1537,1562]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="515" unit="m" value="750.0">
<quantity id="EC4305FCEC1FFF8954D23B10BE7DF9CD" box="[1242,1313,1537,1562]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="515" unit="m" value="750.0">750 m</quantity>
</elevation>
(
<collectionCode id="4DAA30DCEC1FFF8955383B13BEDAF9CE" box="[1328,1414,1538,1562]" pageId="10" pageNumber="515">LMBC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1FFF8950CF3B34BA3AF97E" pageId="10" pageNumber="515" type="diagnosis">
<paragraph id="2B04A819EC1FFF8950CF3B34BA3AF97E" blockId="10.[151,1436,1501,2030]" pageId="10" pageNumber="515">
<emphasis id="19CF740BEC1FFF8950CF3B34BA1DF9EA" bold="true" box="[199,321,1573,1598]" pageId="10" pageNumber="515">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC1FFF89515A3B37BA99F9E9" authorityName="Bocak" authorityYear="2017" box="[338,453,1573,1597]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="515" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="19CF740BEC1FFF89515A3B37BA99F9E9" box="[338,453,1573,1597]" italics="true" pageId="10" pageNumber="515">C. riedeli</emphasis>
</taxonomicName>
is the only species with uniformly yellow pronotum and elytra except for an inconspicuous dark patch in the middle of the pronotum and the infuscate area of elytra at the scutellum (
<figureCitation id="B380B49CEC1FFF8955323B58BECBF9B6" box="[1338,1431,1609,1634]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="10" pageNumber="515">Figs. 31</figureCitation>
,
<figureCitation id="B380B49CEC1FFF89509F3B7FBBEBF952" box="[151,183,1646,1670]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="10" pageNumber="515">48</figureCitation>
). This species has a relatively slender, parallel-sided phallus (
<figureCitation id="B380B49CEC1FFF8953753B7CBF5BF952" box="[893,1031,1645,1670]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="10" pageNumber="515">Figs. 7576</figureCitation>
) and extremely long male antennal lamellae (
<figureCitation id="B380B49CEC1FFF89510D3B80BA04F97E" box="[261,344,1681,1706]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="10" pageNumber="515">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1FFF8850CF3BA4B87DFF00" lastPageId="11" lastPageNumber="516" pageId="10" pageNumber="515" type="description">
<paragraph id="2B04A819EC1FFF8950CF3BA4B893F83A" blockId="10.[151,1436,1501,2030]" pageId="10" pageNumber="515">
<emphasis id="19CF740BEC1FFF8950CF3BA4BA05F91A" bold="true" box="[199,345,1717,1742]" pageId="10" pageNumber="515">Description.</emphasis>
Male. Body large-sized,
<quantity id="EC4305FCEC1FFF8952743BA4B9BEF919" box="[636,738,1717,1742]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="10" pageNumber="515" unit="mm" value="15.1">15.1 mm</quantity>
long, dark brown to black, pronotum and elytra except very small dark patch at scutellum brightly yellow (
<figureCitation id="B380B49CEC1FFF8952B93BC8B84CF926" box="[689,784,1753,1778]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="10" pageNumber="515">Figs. 31</figureCitation>
,
<figureCitation id="B380B49CEC1FFF8953163BCBB862F926" box="[798,830,1754,1778]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="10" pageNumber="515">48</figureCitation>
). Head small, with small, hemispherically prominent eyes, frontal interocular distance 2.0 times maximum diameter; antennae flabellate, long, reaching to three quarters of elytral length if inclined, antennomere 3 with lamella 3.9 times longer than stem of antennomere, lamella attached close to middle part of antennomere (
<figureCitation id="B380B49CEC1FFF8952A83A54B9A8F88A" box="[672,756,1861,1886]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="10" pageNumber="515">Fig. 16</figureCitation>
). Pronotum transverse, 1.5 times wider than long at midline, lateral margins elevated, frontal margin projected anteriorly, frontal angles widely rounded, lateral margins slightly concave, posterior angles projected, all carinae well-developed (
<figureCitation id="B380B49CEC1FFF89538F3A9CB883F872" box="[903,991,1933,1958]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="10" pageNumber="515">Fig. 31</figureCitation>
). Elytra flat, parallel-sided, transverse costae well-developed, regular, elytral areoles slightly transverse (
<figureCitation id="B380B49CEC1FFF89539C3AA0B8B7F81E" box="[916,1003,1969,1994]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="10" pageNumber="515">Fig. 48</figureCitation>
). Phallus slender, parallel-sided, with gradually acuminate apical part and projected slender apex (
<figureCitation id="B380B49CEC1FFF8953313AC4B89FF83A" box="[825,963,2005,2030]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="10" pageNumber="515">Figs. 7576</figureCitation>
).
</paragraph>
<paragraph id="2B04A819EC1EFF8850CF3D86B87DFF00" blockId="11.[151,1437,151,285]" pageId="11" pageNumber="516">
<emphasis id="19CF740BEC1EFF8850CF3D86BA21FF64" bold="true" box="[199,381,151,176]" pageId="11" pageNumber="516">Measurements.</emphasis>
BL
<quantity id="EC4305FCEC1EFF8851A73D86B949FF7B" box="[431,533,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="11" pageNumber="516" unit="mm" value="15.1">15.1 mm</quantity>
, PL
<quantity id="EC4305FCEC1EFF8852403D86B9F3FF7B" box="[584,687,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="11" pageNumber="516" unit="mm" value="1.66">1.66 mm</quantity>
, PW
<quantity id="EC4305FCEC1EFF8852E33D89B80EFF7B" box="[747,850,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.48" pageId="11" pageNumber="516" unit="mm" value="2.48">2.48 mm</quantity>
, WH
<quantity id="EC4305FCEC1EFF88539B3D86B8A5FF7B" box="[915,1017,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="11" pageNumber="516" unit="mm" value="1.56">1.56 mm</quantity>
, Edist
<quantity id="EC4305FCEC1EFF88544E3D86BFF0FF7B" box="[1094,1196,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="516" unit="mm" value="0.95">0.95 mm</quantity>
, Ediam
<quantity id="EC4305FCEC1EFF8855003D89BE32FF7B" box="[1288,1390,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="516" unit="mm" value="0.48">0.48 mm</quantity>
, LE 13.0 mm, antennomere 3: stem
<quantity id="EC4305FCEC1EFF8851F13DADB903FF00" box="[505,607,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="516" unit="mm" value="0.85">0.85 mm</quantity>
, lamella
<quantity id="EC4305FCEC1EFF8852CC3DADB841FF00" box="[708,797,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="516" unit="mm" value="3.3">3.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1EFF8850CF3DCEBEC1FEC9" pageId="11" pageNumber="516" type="distribution">
<paragraph id="2B04A819EC1EFF8850CF3DCEBEC1FEC9" blockId="11.[151,1437,151,285]" pageId="11" pageNumber="516">
<emphasis id="19CF740BEC1EFF8850CF3DCEBA3CFF2C" bold="true" box="[199,352,223,248]" pageId="11" pageNumber="516">Distribution.</emphasis>
Eastern New
<collectingCountry id="53ACE889EC1EFF8851F63DCEB90EFF2C" box="[510,594,223,248]" name="Guinea-Bissau" pageId="11" pageNumber="516">Guinea</collectingCountry>
, the southern slopes of the Owen Stanley Range (
<figureCitation id="B380B49CEC1EFF8854753DCEBF9DFF2C" box="[1149,1217,223,248]" captionStart="FIGURES 1 3. 1 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 3. 1 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="11" pageNumber="516">Fig. 3</figureCitation>
).
<emphasis id="19CF740BEC1EFF8850CF3C15BA12FEC9" bold="true" box="[199,334,260,285]" pageId="11" pageNumber="516">Etymology.</emphasis>
The species name is a patronym in honor of Alexander Riedel (Karlsruhe), a specialist in weevils.
</paragraph>
</subSubSection>
</treatment>
</document>