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<document id="74BBF5F87DA111147C7ADC771A073DD0" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC12FF84509F3CCDBF9AFAA3" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus humeralis Kleine 1926" docType="treatment" docVersion="7" lastPageNumber="512" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="512" updateTime="1698468596363" updateUser="ExternalLinkService">
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<mods:title id="E7049407A1E7DA829555E4FCE68FEBCB">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
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<mods:namePart id="5C0EB053175D75A110F2F365BA592EF2">Bocak, Ladislav</mods:namePart>
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<mods:date id="8D428CB7C4C87564AFFB8FE93F20A98A">2017</mods:date>
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<treatment id="A312190FEC12FF84509F3CCDBF9AFAA3" ID-DOI="http://doi.org/10.5281/zenodo.6018036" ID-GBIF-Taxon="134090580" ID-Zenodo-Dep="6018036" LSID="urn:lsid:plazi:treatment:A312190FEC12FF84509F3CCDBF9AFAA3" httpUri="http://treatment.plazi.org/id/A312190FEC12FF84509F3CCDBF9AFAA3" lastPageNumber="512" pageId="7" pageNumber="512">
<subSubSection id="63A1FB92EC12FF84509F3CCDBAF0FDC3" pageId="7" pageNumber="512" type="nomenclature">
<paragraph id="2B04A819EC12FF84509F3CCDB90BFE22" blockId="7.[151,599,476,535]" box="[151,599,476,502]" pageId="7" pageNumber="512">
<heading id="704C1F75EC12FF84509F3CCDB90BFE22" bold="true" box="[151,599,476,502]" fontSize="11" level="1" pageId="7" pageNumber="512" reason="1">
<taxonomicName id="ECBBD39AEC12FF84509F3CCDB90BFE22" authority="Kleine, 1926" authorityName="Kleine" authorityYear="1926" box="[151,599,476,502]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="19CF740BEC12FF84509F3CCDB90BFE22" bold="true" box="[151,599,476,502]" pageId="7" pageNumber="512">
<emphasis id="19CF740BEC12FF84509F3CCDBAE9FE22" bold="true" box="[151,437,476,502]" italics="true" pageId="7" pageNumber="512">Cladophorus humeralis</emphasis>
<bibRefCitation id="4F2AD5E8EC12FF8451B43CCDB90BFE22" author="Kleine" box="[444,599,476,502]" pageId="7" pageNumber="512" refString="Kleine, R. (1926) Coleoptera Lycidae. Nova Guinea. Resultats des expeditions scientifiques a la Nouvelle Guinee, 15, 91 - 195." type="journal article" year="1926">Kleine, 1926</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="2B04A819EC12FF84509F3CEFBAF0FDC3" blockId="7.[151,599,476,535]" box="[151,428,510,535]" pageId="7" pageNumber="512">
(
<figureCitation id="B380B49CEC12FF8450973CEFBA7AFDC3" box="[159,294,510,535]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="7" pageNumber="512">Figs. 10, 24</figureCitation>
,
<figureCitation id="B380B49CEC12FF84513A3CEEBA0DFDC3" box="[306,337,511,535]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">40</figureCitation>
,
<figureCitation id="B380B49CEC12FF8451553CEFBAF8FDC3" box="[349,420,510,535]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="7" pageNumber="512">6162</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC12FF84509F3F57B924FD8B" box="[151,632,582,607]" pageId="7" pageNumber="512" type="reference_group">
<paragraph id="2B04A819EC12FF84509F3F57B924FD8B" blockId="7.[151,1436,582,1399]" box="[151,632,582,607]" pageId="7" pageNumber="512">
<treatmentCitationGroup id="0BAB8F37EC12FF84509F3F57B924FD8B" box="[151,632,582,607]" pageId="7" pageNumber="512">
<taxonomicName id="ECBBD39AEC12FF84509F3F57B92FFD8B" authority="Kleine, 1926: 130" authorityName="Kleine" authorityPageNumber="130" authorityYear="1926" box="[151,627,582,607]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="19CF740BEC12FF84509F3F57BAFCFD8A" box="[151,416,582,606]" italics="true" pageId="7" pageNumber="512">Cladophorus humeralis</emphasis>
<treatmentCitation id="AA1A8E08EC12FF8451AF3F57B92FFD8B" author="Kleine" box="[423,627,582,607]" page="130" pageId="7" pageNumber="512" year="1926">
<bibRefCitation id="4F2AD5E8EC12FF8451AF3F57B960FD8B" author="Kleine" box="[423,572,582,607]" pageId="7" pageNumber="512" refString="Kleine, R. (1926) Coleoptera Lycidae. Nova Guinea. Resultats des expeditions scientifiques a la Nouvelle Guinee, 15, 91 - 195." type="journal article" year="1926">Kleine, 1926</bibRefCitation>
: 130
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC12FF84509F3F7BBED8FD73" pageId="7" pageNumber="512" type="materials_examined">
<paragraph id="2B04A819EC12FF84509F3F7BBED8FD73" blockId="7.[151,1436,582,1399]" pageId="7" pageNumber="512">
<materialsCitation id="9BD3A244EC12FF84509F3F7BB950FD73" ID-GBIF-Occurrence="1638540609" collectionCode="MIZW" country="Guinea-Bissau" county="Co-Typus'" location="Bomberai Peninsula" municipality="Male" pageId="7" pageNumber="512" specimenCount="2" typeStatus="syntype">
<emphasis id="19CF740BEC12FF84509F3F7BBA1AFD57" bold="true" box="[151,326,618,643]" pageId="7" pageNumber="512">
<typeStatus id="F40016BBEC12FF84509F3F7BBB8EFD57" box="[151,210,618,643]" pageId="7" pageNumber="512">Type</typeStatus>
material.
</emphasis>
<collectingCounty id="C265D095EC12FF84515A3F7ABA92FD57" box="[338,462,619,643]" pageId="7" pageNumber="512">Co-Typus</collectingCounty>
[
<typeStatus id="F40016BBEC12FF8451D53F7AB966FD57" box="[477,570,619,643]" pageId="7" pageNumber="512" type="syntype">syntype</typeStatus>
, No. 274].
<collectingMunicipality id="CB603263EC12FF8452B23F7BB9AAFD57" box="[698,758,618,643]" pageId="7" pageNumber="512">Male</collectingMunicipality>
, New
<collectingCountry id="53ACE889EC12FF8453483F7BB8C8FD57" box="[832,916,618,643]" name="Guinea-Bissau" pageId="7" pageNumber="512">Guinea</collectingCountry>
,
<location id="2E64FEC2EC12FF8453973F7BBFDAFD57" LSID="urn:lsid:plazi:treatment:A312190FEC12FF84509F3CCDBF9AFAA3:2E64FEC2EC12FF8453973F7BBFDAFD57" box="[927,1158,618,643]" country="Guinea-Bissau" county="Co-Typus'" municipality="Male" name="Bomberai Peninsula" pageId="7" pageNumber="512">Bomberai Peninsula</location>
,
<location id="2E64FEC2EC12FF84549A3F7ABFB9FD56" LSID="urn:lsid:plazi:treatment:A312190FEC12FF84509F3CCDBF9AFAA3:2E64FEC2EC12FF84549A3F7ABFB9FD56" box="[1170,1253,619,642]" country="Guinea-Bissau" county="Co-Typus'" municipality="Male" name="Kapaur" pageId="7" pageNumber="512">Kapaur</location>
(
<collectionCode id="4DAA30DCEC12FF8454FD3F7ABE15FD57" box="[1269,1353,619,643]" pageId="7" pageNumber="512">MIZW</collectionCode>
). Other material examined. Male, female
</materialsCitation>
.
<materialsCitation id="9BD3A244EC12FF84521E3F9FBED8FD73" ID-GBIF-Occurrence="1638540607" box="[534,1412,654,679]" collectionCode="LMBC" country="Indonesia" elevation="150" location="West Papua" pageId="7" pageNumber="512" specimenCount="1" stateProvince="West Papua" typeStatus="syntype">
<collectingCountry id="53ACE889EC12FF84521E3F9FB9DAFD73" box="[534,646,654,679]" name="Indonesia" pageId="7" pageNumber="512">Indonesia</collectingCountry>
,
<collectingRegion id="E97F66FBEC12FF8452993F9EB848FD72" box="[657,788,655,678]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="512">West Papua</collectingRegion>
prov., DMP-Wasior,
<elevation id="A0964F2AEC12FF84540B3F9EBFDEFD72" box="[1027,1154,655,679]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="512" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<quantity id="EC4305FCEC12FF84540B3F9EBFDEFD72" box="[1027,1154,655,679]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="512" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200 m</quantity>
</elevation>
(A0043435,
<collectionCode id="4DAA30DCEC12FF84552C3F9EBE24FD73" box="[1316,1400,655,679]" pageId="7" pageNumber="512">LMBC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC12FF8450CF3FA3BA86FCAB" pageId="7" pageNumber="512" type="diagnosis">
<paragraph id="2B04A819EC12FF8450CF3FA3BA86FCAB" blockId="7.[151,1436,582,1399]" pageId="7" pageNumber="512">
<emphasis id="19CF740BEC12FF8450CF3FA3BA1DFD1F" bold="true" box="[199,321,690,715]" pageId="7" pageNumber="512">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC12FF8451403FA2BA85FD1F" box="[328,473,691,715]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="19CF740BEC12FF8451403FA2BA85FD1F" box="[328,473,691,715]" italics="true" pageId="7" pageNumber="512">C. humeralis</emphasis>
</taxonomicName>
has a unique elytral colour pattern with brown humeri and a longitudinal patch in the posterior part of each elytron (
<figureCitation id="B380B49CEC12FF8452093FC7B905FD3B" box="[513,601,726,751]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Fig. 40</figureCitation>
). Further, the phallus of this species is robust and gradually acuminate apically (
<figureCitation id="B380B49CEC12FF84510C3FEBBACEFCC7" box="[260,402,762,787]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="7" pageNumber="512">Figs. 6162</figureCitation>
).
<taxonomicName id="ECBBD39AEC12FF8451A23FEAB921FCC7" authorityName="Bocak" authorityYear="2017" box="[426,637,763,787]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="manokwarensis">
<emphasis id="19CF740BEC12FF8451A23FEAB921FCC7" box="[426,637,763,787]" italics="true" pageId="7" pageNumber="512">C. manokwarensis</emphasis>
</taxonomicName>
was identified as a sister species of
<taxonomicName id="ECBBD39AEC12FF8454323FEABF8CFCC7" box="[1082,1232,763,787]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="19CF740BEC12FF8454323FEABF8CFCC7" box="[1082,1232,763,787]" italics="true" pageId="7" pageNumber="512">C. humeralis</emphasis>
</taxonomicName>
in the molecular phylogeny (
<figureCitation id="B380B49CEC12FF8451143E0FBA3DFCE3" box="[284,353,798,823]" captionStart="FIGURES 4 5. 4" captionStartId="3.[151,264,1220,1243]" captionTargetBox="[159,1417,255,1198]" captionTargetId="figure@3.[151,1436,255,1210]" captionTargetPageId="3" captionText="FIGURES 4 5. 4, Phylogenetic hypothesis of Papuan Cladophorus inferred from the cox 1 mtDNA dataset using maximum likelihood criterion. 5, Cladophorus sp., general appearance." httpUri="https://zenodo.org/record/1051955/files/figure.png" pageId="7" pageNumber="512">Fig. 4</figureCitation>
). Although these species are very close each to other, they differ besides the colour pattern in the shape of the antennomere 3 and the relative size of eyes. The shape of phallus differs only slightly in this pair of species (
<figureCitation id="B380B49CEC12FF8450FF3E77BA8EFCAB" box="[247,466,870,895]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="7" pageNumber="512">Figs. 6162, 6566</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC12FF8450CF3E9BBF50FA86" pageId="7" pageNumber="512" type="description">
<paragraph id="2B04A819EC12FF8450CF3E9BB851FADF" blockId="7.[151,1436,582,1399]" pageId="7" pageNumber="512">
<emphasis id="19CF740BEC12FF8450CF3E9BBA28FC77" bold="true" box="[199,372,906,931]" pageId="7" pageNumber="512">Redescription.</emphasis>
Male. Body large,
<quantity id="EC4305FCEC12FF8452673E9BB847FC76" box="[623,795,906,931]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" metricValueMax="1.5699999999999998" metricValueMin="1.55" pageId="7" pageNumber="512" unit="mm" value="15.6" valueMax="15.7" valueMin="15.5">15.515.7 mm</quantity>
long, dark brown to black including pronotum and substantial part of elytra (
<figureCitation id="B380B49CEC12FF8451C93EBFB94BFC13" box="[449,535,942,967]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Fig. 40</figureCitation>
), humeri light brown, bright humeral patches separated by dark coloured area along inner margin of elytra; additionally long, light patch in posterior part of each elytron, posterior patch with gradual transition between dark and light coloured areas of elytron (
<figureCitation id="B380B49CEC12FF8453A53EE7BF59FBDB" box="[941,1029,1014,1039]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Fig. 40</figureCitation>
). Head small, with hemispherically prominent eyes, frontal interocular distance 1.92 times maximum diameter; antennae long, reaching beyond two thirds of elytral length, flabellate, antennomere 3 with lamella 2.55 times longer than stem, lamella attached to apical part of antennomere (
<figureCitation id="B380B49CEC12FF8451DF3973B970FBAF" box="[471,556,1122,1147]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="7" pageNumber="512">Fig. 10</figureCitation>
). Pronotum transverse, 1.43 times wider than long at midline, lateral margins apparently elevated, frontal margin widely rounded, frontal angles inconspicuous, rounded, lateral margins slightly convex, posterior angles prominent, all carinae well-developed (
<figureCitation id="B380B49CEC12FF84538839BBB884FB17" box="[896,984,1194,1219]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="7" pageNumber="512">Fig. 24</figureCitation>
). Elytra flat, slender, subparallel-sided, transverse costae well-developed, regular, elytral areoles slightly transverse (
<figureCitation id="B380B49CEC12FF8453F839DFBF18FB33" box="[1008,1092,1230,1255]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Fig. 40</figureCitation>
). Phallus robust, parallel-sided in middle, with gradually acuminate apex (
<figureCitation id="B380B49CEC12FF84527F39E3B85DFADF" box="[631,769,1266,1291]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="7" pageNumber="512">Figs. 6162</figureCitation>
).
</paragraph>
<paragraph id="2B04A819EC12FF8450CF3807BF50FA86" blockId="7.[151,1436,582,1399]" pageId="7" pageNumber="512">
<emphasis id="19CF740BEC12FF8450CF3807BA21FAFB" bold="true" box="[199,381,1302,1327]" pageId="7" pageNumber="512">Measurements.</emphasis>
BL
<quantity id="EC4305FCEC12FF8451BD3807B901FAFA" box="[437,605,1302,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" metricValueMax="1.5699999999999998" metricValueMin="1.55" pageId="7" pageNumber="512" unit="mm" value="15.6" valueMax="15.7" valueMin="15.5">15.515.7 mm</quantity>
, PL
<quantity id="EC4305FCEC12FF84529F3806B85CFAFA" box="[663,768,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="512" unit="mm" value="2.1">2.10 mm</quantity>
, PW 3.00
<quantity id="EC4305FCEC12FF84538B3807B8B0FAFA" box="[899,1004,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="7" pageNumber="512" unit="mm" value="3.05">3.05 mm</quantity>
, WH
<quantity id="EC4305FCEC12FF84543B3807BF87FAFA" box="[1075,1243,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.75" metricValueMin="1.7" pageId="7" pageNumber="512" unit="mm" value="1.725" valueMax="1.75" valueMin="1.7">1.701.75 mm</quantity>
, Edist
<quantity id="EC4305FCEC12FF8455263807BEC4FAFA" box="[1326,1432,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="512" unit="mm" value="1.15">1.15 mm</quantity>
, Ediam
<quantity id="EC4305FCEC12FF8450EF382BBA11FA86" box="[231,333,1338,1363]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="512" unit="mm" value="0.6">0.60 mm</quantity>
, LE
<quantity id="EC4305FCEC12FF84518A382ABAB4FA86" box="[386,488,1339,1363]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="7" pageNumber="512" unit="mm" value="13.2">13.2 mm</quantity>
, antennomere 3: stem 1.0 mm, lamella
<quantity id="EC4305FCEC12FF8453A9382BBF54FA86" box="[929,1032,1338,1363]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="7" pageNumber="512" unit="mm" value="2.55">2.55 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC12FF8450CF384FBF9AFAA3" box="[199,1222,1374,1399]" pageId="7" pageNumber="512" type="distribution">
<paragraph id="2B04A819EC12FF8450CF384FBF9AFAA3" blockId="7.[151,1436,582,1399]" box="[199,1222,1374,1399]" pageId="7" pageNumber="512">
<emphasis id="19CF740BEC12FF8450CF384FBA3CFAA3" bold="true" box="[199,352,1374,1399]" pageId="7" pageNumber="512">Distribution.</emphasis>
<collectingRegion id="E97F66FBEC12FF84516F384EBA9EFAA2" box="[359,450,1375,1398]" country="Papua New Guinea" name="Western" pageId="7" pageNumber="512">Western</collectingRegion>
New
<collectingCountry id="53ACE889EC12FF84520D384FB905FAA3" box="[517,601,1374,1399]" name="Guinea-Bissau" pageId="7" pageNumber="512">Guinea</collectingCountry>
, Bomberai Peninsula, Teluk Wondama regency (
<figureCitation id="B380B49CEC12FF845472384FBFE1FAA3" box="[1146,1213,1374,1399]" captionStart="FIGURES 1 3. 1 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 3. 1 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="7" pageNumber="512">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>