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<document id="30F064B5E00BE438E69F59C4ED3B5961" ID-CLB-Dataset="272022" ID-DOI="10.11646/phytotaxa.619.1.2" ID-GBIF-Dataset="1fa451ec-43d2-4b93-a99b-912b03ab5f82" ID-ISSN="1179-3163" ID-Zenodo-Dep="8425689" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1696937392554" checkinUser="plazi" docAuthor="Leme, Elton M. C., Gonella, Paulo M., Couto, Dayvid R., Fernandez, Eduardo P., De Carvalho, Jordano D. T., De Almeida, Pedro S. &amp; Mariath, Jorge E. A." docDate="2023" docId="9175B60B2829FFCCFF15FBA9FC1AEA4F" docLanguage="en" docName="phytotaxa.619.1.2.pdf" docOrigin="Phytotaxa 619 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.619.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Krenakanthus ribeiranus Leme, Gonella &amp; D. R. Couto 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="57" masterDocId="6D4CCE732839FFDEFF9DFFBAFFA6EC17" masterDocTitle="A “ hairy situation ” in Minas Gerais, Brazil: a striking new species of Krenakanthus (Bromeliaceae: Bromelioideae) covered with uniseriate trichomes" masterLastPageNumber="62" masterPageNumber="39" pageNumber="55" updateTime="1697029221272" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2E2DA8A863EC887AB744FD94D64D349D">A “ hairy situation ” in Minas Gerais, Brazil: a striking new species of Krenakanthus (Bromeliaceae: Bromelioideae) covered with uniseriate trichomes</mods:title>
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<mods:namePart id="B306E99AD95CA63C4DE2204444D64EF2">Leme, Elton M. C.</mods:namePart>
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<taxonomicName id="DEDC7C9E2829FFCEFF15FBA9FCA6E839" authority="Leme, Gonella &amp; D.R. Couto" authorityName="Leme, Gonella &amp; D. R. Couto" authorityYear="2023" box="[136,768,1043,1071]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="16" pageNumber="55" phylum="Tracheophyta" rank="species" species="ribeiranus" status="sp. nov.">
<emphasis id="2BA8DB0F2829FFCEFF15FBA9FE0AE839" bold="true" box="[136,428,1043,1070]" italics="true" pageId="16" pageNumber="55">Krenakanthus ribeiranus</emphasis>
Leme, Gonella &amp; D.R. Couto
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<emphasis id="2BA8DB0F2829FFCEFC96FBAEFCFAE839" bold="true" box="[779,860,1044,1070]" italics="true" pageId="16" pageNumber="55">sp. nov</emphasis>
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<figureCitation id="81E71B982829FFCEFCEEFBAEFC45E839" box="[883,995,1044,1070]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="2.[136,229,1657,1679]" captionStartId-1="3.[136,229,1048,1070]" captionStartId-10="13.[136,229,1848,1870]" captionStartId-2="4.[136,229,1996,2018]" captionStartId-3="6.[136,229,1457,1479]" captionStartId-4="7.[136,229,1992,2014]" captionStartId-5="8.[136,229,1953,1975]" captionStartId-6="9.[136,229,1953,1975]" captionStartId-7="10.[136,229,1952,1974]" captionStartId-8="11.[136,229,2025,2047]" captionStartId-9="12.[136,229,1995,2017]" captionTargetBox-0="[180,1407,291,1617]" captionTargetBox-1="[158,1428,197,1017]" captionTargetBox-10="[215,1372,190,1824]" captionTargetBox-2="[233,1354,389,1971]" captionTargetBox-3="[151,1436,190,1433]" captionTargetBox-4="[196,1390,281,1967]" captionTargetBox-5="[178,1409,190,1928]" captionTargetBox-6="[178,1409,190,1928]" captionTargetBox-7="[178,1409,190,1920]" captionTargetBox-8="[222,1366,1145,2000]" captionTargetBox-9="[190,1397,425,1970]" captionTargetId-0="figure-56@2.[167,1419,278,1630]" captionTargetId-1="figure-18@3.[151,1436,190,1024]" captionTargetId-10="figure-18@13.[215,1372,190,1824]" captionTargetId-2="figure-121@4.[233,1354,389,1971]" captionTargetId-3="figure-23@6.[151,1436,190,1433]" captionTargetId-4="figure-55@7.[196,1390,281,1967]" captionTargetId-5="figure-23@8.[178,1409,190,1928]" captionTargetId-6="figure-18@9.[178,1409,190,1928]" captionTargetId-7="figure-23@10.[178,1409,190,1928]" captionTargetId-8="figure-497@11.[222,1366,1145,2000]" captionTargetId-9="figure-144@12.[190,1398,425,1970]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="13" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIGURE 1. Distribution of Krenakanthus ribeiranus and K. roseolilacinus. The Area of Occupancy (AOO) of K. ribeiranus in presented alongside the results of the overlay analyses with Mapbiomas land use cover data series." captionText-1="FIGURE 2. The massif of Serra de Santa Maria immediate delimitation and with buffers of 1 km, 3 km, and 5 km of its immediate surroundings. The overlay analyses of these geometries with Mapbiomas historical land use cover data series is presented for each land use and vegetation cover classes remaining proportion." captionText-10="FIGURE 11. Pollenwithpartial viewofthe reticulumandthe sulcus.A–B.Krenakanthusribeiranus (Leme10220).A.Viewof thereticulum. B. View of the sulcus covered with condensed exine elements. C. Krenakanthus roseolilacinus (Leme 8922). D. Orthocryptanthus arcanus (Leme 9093). E. Orthocryptanthus santaritensis (Leme 8945). F. Orthocryptanthus vasconcelosianus (Leme 8673). G–J. Orthophytum. G. O. (subg. Orthophytum) fosterianum (Leme 5980). H. O. (subg. Orthophytum) macroflorum (Leme 6001). I. O. (subg. Capixabanthus) pseudovagans (Leme 7028). J. O. (subg. Clavanthus) mello-barretoi (Leme 7243). K. Rokautskyia scaposa (Leme 5219). L. Sincoraea ophiurioides (Leme 2282). M. Cryptanthus beuckeri (Leme 7341). N. Forzzaea flavipetala (Leme 9093). O. Hoplocryptanthus regius (Leme 6372). Bars = 5 μm. SEM photos: A–B. P.S. de Almeida. C–O. H. Halbritter." captionText-2="FIGURE 3. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. General aspect of the shaded rocky habitat alongside creeks at the type locality. B–D. Subpopulations at the type locality composed of individuals growing on organic-rich, shallow soils accumulated on sandstone rock surfaces among mosses. Photos: J.C.S. Ribeiro." captionText-3="FIGURE 4. Leaf margin and indumentum in Krenakanthus. A–B. Krenakanthus roseolilacinus. C–E. K. ribeiranus. A–B. Leaf margin with conspicuous spines. C–D. Leaf margin with inconspicuous spines and leaf blade with trichomes. E. Detail of a non-glandular uniseriate trichome. Arrowhead = spine. t = tector trichome. Bars: A–D = 1 mm. E = 0.5 mm." captionText-4="FIGURE 5. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A Habit in three stages of flower development. B. Habit, highlighting an individual in fruit stage. C. Small-sized aspect of the leaf rosette in an adult individual. D. Frontal view of the fan blade-like corolla. E. Lateral view of the corolla. F. Abundant seedlings growing not far from mother-plants. Photos: J.C.S. Ribeiro." captionText-5="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." captionText-6="FIGURE 7. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. Side view of the leaf blade, highlighting the spreading, uniseriate, hair-like trichomes on the abaxial and adaxial surfaces. B. Basal portion of the leaf, evidencing the inconspicuous marginal spines. C. The inconspicuous peduncle of the inflorescence. D. Fruits in different stages of maturation. E. Fruits. F. Seeds. Bars = 5 mm (A–B, D–E). Bars = 2 mm (C, F)." captionText-7="FIGURE 8. Leaf anatomy in Krenakanthus. A, C–D. Krenakanthus ribeiranus. B, E–F: K. roseolilacinus. A–B: leaf median crosssection. C–D. Non-glandular trichome anatomy. E–F: Glandular trichome anatomy. C, E. Trichomes in epidermis context. D. Detail of the stalk. F: detail of the glandular trichome. Arrowhead = stoma. ap = aquiferous parenchyma. bc = basal cells. ch = chlorenchyma. e = epidermis. gc = glandular apical cell. h = hypodermiss. st = stalk. Bars: A–B = 100 μm. C–F = 50 μm." captionText-8="FIGURE 9. Seed morphology in Krenakanthus, seeds in lateral view. A–B. Krenakanthus ribeiranus. C. K. roseolilacinus.. c = chalaza. m = micropyle. r = raphe. Bars = 0.5 mm." captionText-9="FIGURE 10. Seed anatomy. A, C. Krenakanthus ribeiranus. B, D. K. roseolilacinus. A–B. Seeds in median cross-section. C–D. Detail of the seed coat in anti-raphe. Arrowhead: tegmic cuticle. ar = anti-raphe. en = endosperm. pr = pre-raphe. The numbers indicate the different seed coat layers. 1 = endotegmen. 2 = exotegmen. 3 = endotesta. 4 = exotesta. Bars: A–B = 200 μm. C–D = 50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.8425693" figureDoi-1="http://doi.org/10.5281/zenodo.8425695" figureDoi-10="http://doi.org/10.5281/zenodo.8425717" figureDoi-2="http://doi.org/10.5281/zenodo.8425697" figureDoi-3="http://doi.org/10.5281/zenodo.8425699" figureDoi-4="http://doi.org/10.5281/zenodo.8425701" figureDoi-5="http://doi.org/10.5281/zenodo.8431275" figureDoi-6="http://doi.org/10.5281/zenodo.8425703" figureDoi-7="http://doi.org/10.5281/zenodo.8425705" figureDoi-8="http://doi.org/10.5281/zenodo.8425707" figureDoi-9="http://doi.org/10.5281/zenodo.8425713" httpUri-0="https://zenodo.org/record/8425693/files/figure.png" httpUri-1="https://zenodo.org/record/8425695/files/figure.png" httpUri-10="https://zenodo.org/record/8425717/files/figure.png" httpUri-2="https://zenodo.org/record/8425697/files/figure.png" httpUri-3="https://zenodo.org/record/8425699/files/figure.png" httpUri-4="https://zenodo.org/record/8425701/files/figure.png" httpUri-5="https://zenodo.org/record/8431275/files/figure.png" httpUri-6="https://zenodo.org/record/8425703/files/figure.png" httpUri-7="https://zenodo.org/record/8425705/files/figure.png" httpUri-8="https://zenodo.org/record/8425707/files/figure.png" httpUri-9="https://zenodo.org/record/8425713/files/figure.png" pageId="16" pageNumber="55">figs. 1–11</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="51C654962829FFCEFF15FBE5FC9CE93D" pageId="16" pageNumber="55" type="diagnosis">
<paragraph id="1963071D2829FFCEFF15FBE5FC9CE93D" blockId="16.[136,1452,1119,1430]" pageId="16" pageNumber="55">
<emphasis id="2BA8DB0F2829FFCEFF15FBE5FF4DE862" bold="true" box="[136,235,1119,1141]" pageId="16" pageNumber="55">Diagnosis</emphasis>
:—This new species is morphologically related to 
<taxonomicName id="DEDC7C9E2829FFCEFD5AFBDAFCCCE862" authorityName="Leme, Zizka &amp; Paule (Leme et al." authorityYear="2022" baseAuthorityName="Leme" baseAuthorityYear="2015" box="[711,874,1120,1141]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="16" pageNumber="55" phylum="Tracheophyta" rank="species" species="roseolilacinus">
<emphasis id="2BA8DB0F2829FFCEFD5AFBDAFCCCE862" box="[711,874,1120,1141]" italics="true" pageId="16" pageNumber="55">K. roseolilacinus</emphasis>
</taxonomicName>
, but distinctly differs by stemless habit (
<emphasis id="2BA8DB0F2829FFCEFB76FBDAFB59E862" box="[1259,1279,1120,1141]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. long caulescent), leaf blades subdensely to densely covered by spreading, hair-like trichomes (
<emphasis id="2BA8DB0F2829FFCEFC5CFB3EFC73E88E" box="[961,981,1156,1177]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. glabrescent or glabrous), margins straight (
<emphasis id="2BA8DB0F2829FFCEFA0CFB3EFA03E88E" box="[1425,1445,1156,1177]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. undulate mainly toward the base), inconspicuously spinulose and appearing entire (
<emphasis id="2BA8DB0F2829FFCEFC73FB12FBAFE8AA" box="[1006,1033,1192,1213]" italics="true" pageId="16" pageNumber="55">vs.</emphasis>
distinctly spinulose), inflorescence shortly pedunculate to subsessile (
<emphasis id="2BA8DB0F2829FFCEFE59FB76FE7EE8F6" box="[452,472,1228,1249]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. sessile), simple (
<emphasis id="2BA8DB0F2829FFCEFD16FB76FD39E8F6" box="[651,671,1228,1249]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. sparsely branched only at the base), flowers distinctly smaller (
<emphasis id="2BA8DB0F2829FFCEFA82FB76FA90E8F6" box="[1311,1334,1228,1249]" italics="true" pageId="16" pageNumber="55">ca</emphasis>
. 
<quantity id="DE24AAF82829FFCEFAD9FB76FA2FE8F6" box="[1348,1417,1228,1249]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="55" unit="mm" value="30.0">30 mm</quantity>
<emphasis id="2BA8DB0F2829FFCEFA0CFB76FA03E8F6" box="[1425,1445,1228,1249]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. 
<quantity id="DE24AAF82829FFCEFF20FB55FE83E912" box="[189,293,1263,1285]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="5.5" metricValueMin="4.2" pageId="16" pageNumber="55" unit="mm" value="48.5" valueMax="55.0" valueMin="42.0">42–55 mm</quantity>
long), sepals smaller (11–11.5 × 
<quantity id="DE24AAF82829FFCEFDF4FB55FD14E912" box="[617,690,1263,1285]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="55" unit="mm" value="2.5">2.5 mm</quantity>
<emphasis id="2BA8DB0F2829FFCEFD24FB4AFD6BE912" box="[697,717,1264,1285]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. 20–27 × 
<quantity id="DE24AAF82829FFCEFCADFB55FC22E912" box="[816,900,1263,1285]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="16" pageNumber="55" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
), petals smaller (25–26 × 
<quantity id="DE24AAF82829FFCEFB1DFB4AFB11E912" box="[1152,1207,1264,1285]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="55" unit="mm" value="8.0">8 mm</quantity>
<emphasis id="2BA8DB0F2829FFCEFB23FB4AFB74E912" box="[1214,1234,1264,1285]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. 35–44 × 
<quantity id="DE24AAF82829FFCEFAA8FB55FA07E912" box="[1333,1441,1263,1285]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="16" pageNumber="55" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
), and exappendiculate (
<emphasis id="2BA8DB0F2829FFCEFE0DFAAEFE02E93E" box="[400,420,1300,1321]" italics="true" pageId="16" pageNumber="55">vs</emphasis>
. bearing distinct cupuliform appendages).
</paragraph>
</subSubSection>
<subSubSection id="51C654962829FFCEFF15FA82FC8EE982" pageId="16" pageNumber="55" type="materials_examined">
<paragraph id="1963071D2829FFCEFF15FA82FC8EE982" blockId="16.[136,1452,1119,1430]" pageId="16" pageNumber="55">
<emphasis id="2BA8DB0F2829FFCEFF15FA82FF1CE95A" bold="true" box="[136,186,1336,1357]" pageId="16" pageNumber="55">Type</emphasis>
:— 
<materialsCitation id="A9B40D402829FFCEFF4AFA82FE68E982" collectingDate="2023-03-01" collectionCode="RB" collectorName="P. M. Gonella &amp; J. C. S. Ribeiro &amp; D. P. Cordeiro &amp; L. D. Martins &amp; J. Martins" country="Brazil" elevation="650" location="Serra de Santa Maria" municipality="Alvarenga" pageId="16" pageNumber="55" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="61CB478D2829FFCEFF4AFA82FE95E95A" box="[215,307,1336,1357]" name="Brazil" pageId="16" pageNumber="55">BRAZIL</collectingCountry>
. 
<collectingRegion id="DB18C9FF2829FFCEFEA3FA82FE63E959" box="[318,453,1336,1358]" country="Brazil" name="Minas Gerais" pageId="16" pageNumber="55">Minas Gerais</collectingRegion>
: 
<collectingMunicipality id="F9079D672829FFCEFE4DFA82FD90E959" box="[464,566,1336,1358]" pageId="16" pageNumber="55">Alvarenga</collectingMunicipality>
, 
<location id="1C0351C62829FFCEFDDEFA82FCBEE959" LSID="urn:lsid:plazi:treatment:9175B60B2829FFCCFF15FBA9FC1AEA4F:1C0351C62829FFCEFDDEFA82FCBEE959" box="[579,792,1336,1358]" country="Brazil" municipality="Alvarenga" name="Serra de Santa Maria" pageId="16" pageNumber="55" stateProvince="Minas Gerais">Serra de Santa Maria</location>
, maciço quartzítico a leste da sede do município, [precise locality withheld for conservation purposes], 
<quantity id="DE24AAF82829FFCEFD82FAE1FDFDE966" box="[543,603,1371,1393]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="16" pageNumber="55" unit="m" value="650.0">
<elevation id="92F1E02E2829FFCEFD82FAE1FDFDE966" box="[543,603,1371,1393]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="16" pageNumber="55" unit="m" value="650.0">650 m</elevation>
</quantity>
elevation, 
<date id="6D6221DD2829FFCEFD5EFAE6FCECE966" box="[707,842,1372,1394]" pageId="16" pageNumber="55" value="2023-03-01">
<collectingDate id="7D26D8352829FFCEFD5EFAE6FCECE966" box="[707,842,1372,1394]" pageId="16" pageNumber="55" value="2023-03-01">1 March 2023</collectingDate>
</date>
, 
<emphasis id="2BA8DB0F2829FFCEFCCEFAE6FE9FE982" italics="true" pageId="16" pageNumber="55">
<collectorName id="B42962CB2829FFCEFCCEFAE6FC76E966" box="[851,976,1372,1393]" pageId="16" pageNumber="55">P.M. Gonella</collectorName>
3760, 
<collectorName id="B42962CB2829FFCEFB8CFAE6FB31E966" box="[1041,1175,1372,1393]" pageId="16" pageNumber="55">J.C.S. Ribeiro</collectorName>
, 
<collectorName id="B42962CB2829FFCEFB3DFAE6FA81E966" box="[1184,1319,1372,1393]" pageId="16" pageNumber="55">D.P. Cordeiro</collectorName>
, 
<collectorName id="B42962CB2829FFCEFAADFAE6FA0DE966" box="[1328,1451,1372,1393]" pageId="16" pageNumber="55">L.D. Martins</collectorName>
&amp; 
<collectorName id="B42962CB2829FFCEFF4BFA3AFE9FE982" box="[214,313,1408,1429]" pageId="16" pageNumber="55">J. Martins</collectorName>
</emphasis>
(
<typeStatus id="C667B9BF2829FFCEFED8FA3AFE3DE981" box="[325,411,1408,1430]" pageId="16" pageNumber="55">holotype</typeStatus>
<collectionCode id="7FCD9FD82829FFCEFE3CFA3AFE61E982" box="[417,455,1408,1429]" pageId="16" pageNumber="55">RB</collectionCode>
!,
</materialsCitation>
<materialsCitation id="A9B40D402829FFCEFE48FA3AFC82E982" box="[469,804,1408,1430]" collectingDate="2023-03-01" collectionCode="BHCB, CESJ, MBML" collectorName="P. M. Gonella &amp; J. C. S. Ribeiro &amp; D. P. Cordeiro &amp; L. D. Martins &amp; J. Martins" country="Brazil" elevation="650" location="Serra de Santa Maria" municipality="Alvarenga" pageId="16" pageNumber="55" specimenCount="3" stateProvince="Minas Gerais" typeStatus="isotype">
<typeStatus id="C667B9BF2829FFCEFE48FA3AFD82E981" box="[469,548,1408,1430]" pageId="16" pageNumber="55">isotypes</typeStatus>
<collectionCode id="7FCD9FD82829FFCEFDB4FA3AFDD7E982" box="[553,625,1408,1429]" pageId="16" pageNumber="55">BHCB</collectionCode>
!, 
<collectionCode id="7FCD9FD82829FFCEFDE2FA3AFD1DE982" box="[639,699,1408,1429]" collectionName="CESJ" pageId="16" pageNumber="55">CESJ</collectionCode>
!, 
<collectionCode id="7FCD9FD82829FFCEFD55FA3AFCBCE982" box="[712,794,1408,1429]" collectionName="MBML" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15042" lsid="urn:lsid:biocol.org:col:15042" name="Museu de Biologia Mello Leitão" pageId="16" pageNumber="55" type="Herbarium">MBML</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="51C654962829FFCFFF15FA7EFC55EE7F" lastPageId="17" lastPageNumber="56" pageId="16" pageNumber="55" type="description">
<paragraph id="1963071D2829FFCFFF15FA7EFC55EE7F" blockId="16.[136,1452,1475,2079]" lastBlockId="17.[136,1452,158,2092]" lastPageId="17" lastPageNumber="56" pageId="16" pageNumber="55">
<emphasis id="2BA8DB0F2829FFCEFF15FA7EFEB4E9C9" bold="true" box="[136,274,1476,1502]" pageId="16" pageNumber="55">Description</emphasis>
:— 
<emphasis id="2BA8DB0F2829FFCEFEABFA79FED9E9C9" bold="true" box="[310,383,1475,1502]" italics="true" pageId="16" pageNumber="55">Plants</emphasis>
saxicolous, stemless, flowering 
<quantity id="DE24AAF82829FFCEFD7AFA7EFCE8E9C8" box="[743,846,1476,1503]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="16" pageNumber="55" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 cm</quantity>
heigh, propagating by short basal shoots. 
<emphasis id="2BA8DB0F2829FFCEFA83FA7EFACBE9C9" bold="true" box="[1310,1389,1476,1502]" italics="true" pageId="16" pageNumber="55">Leaves</emphasis>
<quantity id="DE24AAF82829FFCEFAEEFA7EFF38EA15" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="3.048" metricValueMin="1.778" pageId="16" pageNumber="55" unit="in" value="9.5" valueMax="12.0" valueMin="7.0">7–12 in</quantity>
number, very thin in texture, spreading, prostrate, forming a lax rosette without water impounding capacity; 
<emphasis id="2BA8DB0F2829FFCEFAC9FA5DFA0DEA15" bold="true" box="[1364,1451,1511,1538]" italics="true" pageId="16" pageNumber="55">sheaths</emphasis>
inconspicuous, 
<emphasis id="2BA8DB0F2829FFCEFEA7F9B7FEFAEA31" box="[314,348,1549,1574]" italics="true" pageId="16" pageNumber="55">ca.</emphasis>
9 × 
<quantity id="DE24AAF82829FFCEFE07F9B7FE46EA30" box="[410,480,1549,1575]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="55" unit="mm" value="7.0">7 mm</quantity>
, remotely spinulose, nerved, with sparse, long hair-like trichomes; 
<emphasis id="2BA8DB0F2829FFCEFB6FF9B1FA9DEA31" bold="true" box="[1266,1339,1547,1574]" italics="true" pageId="16" pageNumber="55">blades</emphasis>
narrowly lanceolate, narrowed toward the base, apex acuminate-caudate, 7–25 × 
<quantity id="DE24AAF82829FFCEFC4FF98AFBF1EA5C" box="[978,1111,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="16" pageNumber="55" unit="cm" value="1.3" valueMax="1.8" valueMin="0.8">0.8–1.8 cm</quantity>
, green, concolorous, opaque, finely nerved, subdensely to densely covered by spreading, uniseriate hair-like trichomes mainly abaxially, margins straight, with dense long hair-like trichomes, inconspicuously spinulose and appearing entire; 
<emphasis id="2BA8DB0F2829FFCEFB04F9C2FB47EA85" box="[1177,1249,1656,1682]" italics="true" pageId="16" pageNumber="55">
<emphasis id="2BA8DB0F2829FFCEFB04F9C2FB70EA85" bold="true" box="[1177,1238,1656,1682]" italics="true" pageId="16" pageNumber="55">spine</emphasis>
s
</emphasis>
whitish, narrowly triangular, 
<quantity id="DE24AAF82829FFCEFE99F926FE2BEAA0" box="[260,397,1692,1719]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="16" pageNumber="55" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
long, 
<quantity id="DE24AAF82829FFCEFE45F926FD91EAA0" box="[472,567,1692,1719]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="16" pageNumber="55" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
apart, spreading (upper ones) to slightly antrorse (basal ones). 
<emphasis id="2BA8DB0F2829FFCEFA93F921FA0DEAA1" bold="true" box="[1294,1451,1691,1718]" italics="true" pageId="16" pageNumber="55">Inflorescence</emphasis>
shortly pedunculate to subessile, 
<quantity id="DE24AAF82829FFCEFD9CF97AFDCCEACC" box="[513,618,1728,1755]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="16" pageNumber="55" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity>
long (not including the petals), 
<emphasis id="2BA8DB0F2829FFCEFC44F97BFC55EACD" box="[985,1011,1729,1754]" italics="true" pageId="16" pageNumber="55">ca</emphasis>
. 
<quantity id="DE24AAF82829FFCEFB9EF97AFBF6EACC" box="[1027,1104,1728,1755]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="55" unit="cm" value="1.5">1.5 cm</quantity>
in diameter (not including the petals), simple, corymbose; 
<emphasis id="2BA8DB0F2829FFCEFE59F959FD8AEAE9" bold="true" box="[452,556,1763,1790]" italics="true" pageId="16" pageNumber="55">peduncle</emphasis>
<quantity id="DE24AAF82829FFCEFDA9F95EFD3EEAE8" box="[564,664,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="16" pageNumber="55" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
long, 
<emphasis id="2BA8DB0F2829FFCEFD42F95FFD5FEAE9" box="[735,761,1765,1790]" italics="true" pageId="16" pageNumber="55">ca</emphasis>
. 
<quantity id="DE24AAF82829FFCEFC95F95EFCEFEAE8" box="[776,841,1764,1791]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="55" unit="mm" value="2.0">2 mm</quantity>
in diameter, greenish-white, with spreading uniseriate hair-like trichomes; 
<emphasis id="2BA8DB0F2829FFCEFEFBF8BDFDBEEB35" bold="true" box="[358,536,1799,1826]" italics="true" pageId="16" pageNumber="55">peduncle bracts</emphasis>
foliaceous; 
<emphasis id="2BA8DB0F2829FFCEFD01F8BDFC81EB35" bold="true" box="[668,807,1799,1826]" italics="true" pageId="16" pageNumber="55">floral bracts</emphasis>
the basal ones foliaceous, distinctly exceeding the flowers, the upper ones lanceolate, acuminate-caudate, very thin in texture, green, spreading-recurved, shorter than the sepals, with spreading uniseriate hair-like trichomes, finely nerved, ecarinate, 5–10 × 
<quantity id="DE24AAF82829FFCEFC75F8EAFBFCEB7C" box="[1000,1114,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="16" pageNumber="55" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 mm</quantity>
, margins entire. 
<emphasis id="2BA8DB0F2829FFCEFA8DF8F5FACBEB7D" bold="true" box="[1296,1389,1871,1898]" italics="true" pageId="16" pageNumber="55">Flowers</emphasis>
<quantity id="DE24AAF82829FFCEFAEEF8EAFF38EB99" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.54" metricValueMin="1.27" pageId="16" pageNumber="55" unit="in" value="7.5" valueMax="10.0" valueMin="5.0">5–10 in</quantity>
number, all perfect, sessile, 
<emphasis id="2BA8DB0F2829FFCEFE7AF8CFFDA7EB99" box="[487,513,1909,1934]" italics="true" pageId="16" pageNumber="55">ca</emphasis>
. 
<quantity id="DE24AAF82829FFCEFD8CF8CEFDC4EB98" box="[529,610,1908,1935]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="55" unit="mm" value="30.0">30 mm</quantity>
long (with the petals extended), fragrance not detected; 
<emphasis id="2BA8DB0F2829FFCEFB76F8C9FA97EB99" bold="true" box="[1259,1329,1907,1934]" italics="true" pageId="16" pageNumber="55">sepals</emphasis>
11–11.5 × 
<quantity id="DE24AAF82829FFCEFF15F822FF46EBA4" box="[136,224,1944,1971]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="55" unit="mm" value="2.5">2.5 mm</quantity>
, lanceolate, acuminate, entire, connate at the base for 
<quantity id="DE24AAF82829FFCEFCDDF822FC13EBA4" box="[832,949,1944,1971]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="16" pageNumber="55" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
, green but drying dark castaneous soon after anthesis, with sparse spreading hair-like trichomes, thin in texture, the adaxial ones carinate with the keels continuing on the ovary, the abaxial one obtusely if at all carinate; 
<emphasis id="2BA8DB0F2829FFCEFC9EF865FCE0EBED" bold="true" box="[771,838,2015,2042]" italics="true" pageId="16" pageNumber="55">petals</emphasis>
broadly spathulate from a very narrow base, 25–26 × 
<quantity id="DE24AAF82829FFCEFF15F7BEFF6AE408" box="[136,204,2052,2079]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="55" unit="mm" value="8.0">8 mm</quantity>
, 3.1–3.2 times longer than wide, free, green at the base and lilac toward the apex, the blades broadly elliptic to broadly obovate, acute, spreading at anthesis and forming a fan blade-like corolla, flaccidescent afterwards, at the base with 2 longitudinal callosities equaling the antepetalous filaments, with sparse and inconspicuous glandular trichomes, exappendiculate; 
<emphasis id="2BA8DB0F2828FFCFFED6FF5DFE0EED16" bold="true" box="[331,424,231,257]" italics="true" pageId="17" pageNumber="56">stamens</emphasis>
deeply included and not visible; 
<emphasis id="2BA8DB0F2828FFCFFC89FF5CFCD9ED16" bold="true" box="[788,895,230,257]" italics="true" pageId="17" pageNumber="56">filaments</emphasis>
distinctly unequal in length, the antepetalous ones 
<emphasis id="2BA8DB0F2828FFCFFF15FEB1FF04ED33" box="[136,162,267,292]" italics="true" pageId="17" pageNumber="56">ca</emphasis>
. 
<quantity id="DE24AAF82828FFCFFF2CFEB0FEA1ED32" box="[177,263,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="56" unit="mm" value="3.5">3.5 mm</quantity>
long, adnate to the petals for 
<emphasis id="2BA8DB0F2828FFCFFDCBFEB1FDD6ED33" box="[598,624,267,292]" italics="true" pageId="17" pageNumber="56">ca</emphasis>
. 
<quantity id="DE24AAF82828FFCFFDE3FEB0FD71ED32" box="[638,727,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="56" unit="mm" value="0.5">0.5 mm</quantity>
, the antesepalous ones 
<emphasis id="2BA8DB0F2828FFCFFC40FEB1FC51ED33" box="[989,1015,267,292]" italics="true" pageId="17" pageNumber="56">ca</emphasis>
. 
<quantity id="DE24AAF82828FFCFFB9BFEB0FBE1ED32" box="[1030,1095,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="56" unit="mm" value="5.0">5 mm</quantity>
long, free, slightly complanate, greenish-white toward the distal end; 
<emphasis id="2BA8DB0F2828FFCFFDD8FE94FD3AED5E" bold="true" box="[581,668,302,329]" italics="true" pageId="17" pageNumber="56">anthers</emphasis>
<quantity id="DE24AAF82828FFCFFD3AFE94FCB8ED5E" box="[679,798,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="17" pageNumber="56" unit="mm" value="1.6" valueMax="2.0" valueMin="1.2">1.2–2 mm</quantity>
long, oblong, dorsifixed at 
<emphasis id="2BA8DB0F2828FFCFFBF7FE95FB22ED5F" box="[1130,1156,303,328]" italics="true" pageId="17" pageNumber="56">ca</emphasis>
. 1/4 from the base, base bilobed, apex distinctly apiculate; 
<emphasis id="2BA8DB0F2828FFCFFD93FEE8FDF0ED7A" bold="true" box="[526,598,338,365]" italics="true" pageId="17" pageNumber="56">pollen</emphasis>
oblate, 25–35 μm in diameter, sulcate, sulcus narrow, with exine elements, margins weakly defined to indistinct, exine reticulate, reticulum broadly meshed proximally and meshes decreasing in size towards the sulcus margins; 
<emphasis id="2BA8DB0F2828FFCFFDBCFE20FDF2EDA2" bold="true" box="[545,596,410,437]" italics="true" pageId="17" pageNumber="56">style</emphasis>
about equaling the antepetalous stamens and shorter than the antesepalous ones, white; 
<emphasis id="2BA8DB0F2828FFCFFE87FE04FEC0EDCF" bold="true" box="[282,358,446,472]" italics="true" pageId="17" pageNumber="56">stigma</emphasis>
conduplicate-spiral, whitish, blades elongate, papillose, erect; 
<emphasis id="2BA8DB0F2828FFCFFBADFE05FBD6EDCE" bold="true" box="[1072,1136,447,473]" italics="true" pageId="17" pageNumber="56">ovary</emphasis>
<quantity id="DE24AAF82828FFCFFBE4FE04FB71EDCE" box="[1145,1239,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="17" pageNumber="56" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, 
<emphasis id="2BA8DB0F2828FFCFFABCFE05FA9DEDCF" box="[1313,1339,447,472]" italics="true" pageId="17" pageNumber="56">ca</emphasis>
. 
<quantity id="DE24AAF82828FFCFFAD6FE04FA2BEDCE" box="[1355,1421,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="56" unit="mm" value="3.0">3 mm</quantity>
in diameter distally, trigonous, white, with sparse hair-like trichomes; 
<emphasis id="2BA8DB0F2828FFCFFCE7FE58FB81EDEA" bold="true" box="[890,1063,482,509]" italics="true" pageId="17" pageNumber="56">epigynous tube</emphasis>
lacking; 
<emphasis id="2BA8DB0F2828FFCFFB13FE58FB71EDEA" bold="true" box="[1166,1239,482,509]" italics="true" pageId="17" pageNumber="56">ovules</emphasis>
numerous, obtuse; 
<emphasis id="2BA8DB0F2828FFCFFF15FDBCFEB1EE36" bold="true" box="[136,279,518,545]" italics="true" pageId="17" pageNumber="56">placentation</emphasis>
axial, median to apical. 
<emphasis id="2BA8DB0F2828FFCFFDB1FDBCFDD2EE37" bold="true" box="[556,628,518,544]" italics="true" pageId="17" pageNumber="56">Fruits</emphasis>
subglobose, greenish becoming pale yellowish castaneous to castaneous, 4 × 
<quantity id="DE24AAF82828FFCFFF28FD90FE8FEE52" box="[181,297,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="17" pageNumber="56" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 mm</quantity>
, with persistent sepals, 2.7–2.9 times longer than the fruit length; 
<emphasis id="2BA8DB0F2828FFCFFB8CFD90FBE8EE52" bold="true" box="[1041,1102,554,581]" italics="true" pageId="17" pageNumber="56">seeds</emphasis>
at least 
<quantity id="DE24AAF82828FFCFFB31FD90FB43EE53" box="[1196,1253,554,580]" metricMagnitude="0" metricUnit="m" metricValue="2.286" pageId="17" pageNumber="56" unit="in" value="90.0">90 in</quantity>
number, obovoid to subtrapeziform, 0.8–1.2 × 
<quantity id="DE24AAF82828FFCFFE53FDF4FD80EE7E" box="[462,550,590,617]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="56" unit="mm" value="0.6">0.6 mm</quantity>
, yellow, distinctly longitudinally sulcate.
</paragraph>
</subSubSection>
<subSubSection id="51C654962828FFCFFF20FDC8FF49EAB6" pageId="17" pageNumber="56" type="distribution">
<paragraph id="1963071D2828FFCFFF20FDC8FA20EEEF" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
<emphasis id="2BA8DB0F2828FFCFFF20FDC8FE4CEE9B" bold="true" box="[189,490,626,652]" pageId="17" pageNumber="56">Distribution and habitat</emphasis>
:— 
<taxonomicName id="DEDC7C9E2828FFCFFD90FDC8FC8FEE9B" authorityName="Leme, Gonella &amp; D. R. Couto" authorityYear="2023" box="[525,809,626,652]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="17" pageNumber="56" phylum="Tracheophyta" rank="species" species="ribeiranus">
<emphasis id="2BA8DB0F2828FFCFFD90FDC8FC8FEE9B" box="[525,809,626,652]" italics="true" pageId="17" pageNumber="56">Krenakanthus ribeiranus</emphasis>
</taxonomicName>
is a sciadophytic, saxicolous species known from two small subpopulations located in the 
<typeStatus id="C667B9BF2828FFCFFD84FD2DFDEFEEA6" box="[537,585,663,689]" pageId="17" pageNumber="56">type</typeStatus>
region, the Serra de Santa Maria, Alvarenga county, 
<collectingRegion id="DB18C9FF2828FFCFFB07FD2CFACCEEA6" box="[1178,1386,662,689]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="56">Minas Gerais state</collectingRegion>
(
<figureCitation id="81E71B982828FFCFFAE4FD2CFF13EEC3" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1657,1679]" captionStartId-1="3.[136,229,1048,1070]" captionTargetBox-0="[180,1407,291,1617]" captionTargetBox-1="[158,1428,197,1017]" captionTargetId-0="figure-56@2.[167,1419,278,1630]" captionTargetId-1="figure-18@3.[151,1436,190,1024]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Distribution of Krenakanthus ribeiranus and K. roseolilacinus. The Area of Occupancy (AOO) of K. ribeiranus in presented alongside the results of the overlay analyses with Mapbiomas land use cover data series." captionText-1="FIGURE 2. The massif of Serra de Santa Maria immediate delimitation and with buffers of 1 km, 3 km, and 5 km of its immediate surroundings. The overlay analyses of these geometries with Mapbiomas historical land use cover data series is presented for each land use and vegetation cover classes remaining proportion." figureDoi-0="http://doi.org/10.5281/zenodo.8425693" figureDoi-1="http://doi.org/10.5281/zenodo.8425695" httpUri-0="https://zenodo.org/record/8425693/files/figure.png" httpUri-1="https://zenodo.org/record/8425695/files/figure.png" pageId="17" pageNumber="56">figs. 1–2</figureCitation>
). The species habitat is a riparian forest in a matrix of Semideciduous Seasonal Forest and Campos Rupestres in the Atlantic Forest domain, where it is found in a small fragment of an advanced regeneration of secondary forest.
</paragraph>
<paragraph id="1963071D2828FFCFFF20FCB8FA9FEFBB" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
The only known two subpopulations of the species are located very close to the forest edges, which directly adjoins pasture areas. The first subpopulation, located near a waterfall, contains approximately 200 mature individuals and a large number of juveniles. The species appears to have a high recruitment rate, with many germinating seeds and young plants taking root (
<figureCitation id="81E71B982828FFCFFE31FCD4FDA2EF9F" box="[428,516,878,905]" captionStart="FIGURE 5" captionStartId="7.[136,229,1992,2014]" captionTargetBox="[196,1390,281,1967]" captionTargetId="figure-55@7.[196,1390,281,1967]" captionTargetPageId="7" captionText="FIGURE 5. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A Habit in three stages of flower development. B. Habit, highlighting an individual in fruit stage. C. Small-sized aspect of the leaf rosette in an adult individual. D. Frontal view of the fan blade-like corolla. E. Lateral view of the corolla. F. Abundant seedlings growing not far from mother-plants. Photos: J.C.S. Ribeiro." figureDoi="http://doi.org/10.5281/zenodo.8425701" httpUri="https://zenodo.org/record/8425701/files/figure.png" pageId="17" pageNumber="56">fig. 5 F</figureCitation>
). Seed germination occurs in close proximity with mother plants, and occasionally within the fruit, indicating limited dispersal capacity, which may explain the restricted range of the species.
</paragraph>
<paragraph id="1963071D2828FFCFFF20FC0CFE2FE82B" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
The second subpopulation contains approximately 100 mature individuals, located on a rock wall by the river about 
<quantity id="DE24AAF82828FFCFFF50FC60FE96EFE2" box="[205,304,986,1013]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="17" pageNumber="56" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
from a pasture. This subpopulation is particularly vulnerable to fire events that could seriously impact the species, not only directly but also by altering the deeply shaded environment that is critical for its survival, given its high shade demand.
</paragraph>
<paragraph id="1963071D2828FFCFFF20FBFCFE8DE8BF" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
Both subpopulations are located at about 
<quantity id="DE24AAF82828FFCFFD1AFBFCFCA3E876" box="[647,773,1094,1121]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="17" pageNumber="56" unit="m" value="600.0" valueMax="650.0" valueMin="550.0">550–650 m</quantity>
elevation and are composed of scattered individuals growing on organic-rich, shallow soils accumulated on sandstone rock surfaces among mosses, in shaded rocky spots alongside creeks (
<figureCitation id="81E71B982828FFCFFF42FB34FEBBE8BF" box="[223,285,1166,1192]" captionStart="FIGURE 3" captionStartId="4.[136,229,1996,2018]" captionTargetBox="[233,1354,389,1971]" captionTargetId="figure-121@4.[233,1354,389,1971]" captionTargetPageId="4" captionText="FIGURE 3. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. General aspect of the shaded rocky habitat alongside creeks at the type locality. B–D. Subpopulations at the type locality composed of individuals growing on organic-rich, shallow soils accumulated on sandstone rock surfaces among mosses. Photos: J.C.S. Ribeiro." figureDoi="http://doi.org/10.5281/zenodo.8425697" httpUri="https://zenodo.org/record/8425697/files/figure.png" pageId="17" pageNumber="56">fig. 3</figureCitation>
).
</paragraph>
<paragraph id="1963071D2828FFCFFF20FB08FA02E94B" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">The Serra de Santa Maria range is situated on the boundary of Alvarenga, Inhapim, and Tarumirim counties. Despite sharing a similar geomorphological history with other ranges in the area, such as Pico da Aliança in Alvarenga county, it is geographically isolated and predominantly composed of quartzitic Campos Rupestres and semideciduous forest remnants. The range is inadequately protected by any designated protected area and has been poorly sampled botanically. Human activities have resulted in severe habitat conversion, exacerbating the range’s biota vulnerability.</paragraph>
<paragraph id="1963071D2828FFCFFF20FADCFF49EAB6" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
The applied overlay analyses (Jord„o 
<emphasis id="2BA8DB0F2828FFCFFDFDFADDFD3EE997" box="[608,664,1382,1408]" italics="true" pageId="17" pageNumber="56">et al.</emphasis>
2022) showed that the massif is suffering from intense human-induced disturbances and that only a small fraction of the range is suitable for the species (
<figureCitation id="81E71B982828FFCFFBBDFA30FBF9E9B3" box="[1056,1119,1418,1444]" captionStart="FIGURE 2" captionStartId="3.[136,229,1048,1070]" captionTargetBox="[158,1428,197,1017]" captionTargetId="figure-18@3.[151,1436,190,1024]" captionTargetPageId="3" captionText="FIGURE 2. The massif of Serra de Santa Maria immediate delimitation and with buffers of 1 km, 3 km, and 5 km of its immediate surroundings. The overlay analyses of these geometries with Mapbiomas historical land use cover data series is presented for each land use and vegetation cover classes remaining proportion." figureDoi="http://doi.org/10.5281/zenodo.8425695" httpUri="https://zenodo.org/record/8425695/files/figure.png" pageId="17" pageNumber="56">fig. 2</figureCitation>
). Although 48.7% of the total massif area is still forested, and 13.81% is rocky outcrops, we have only located 
<taxonomicName id="DEDC7C9E2828FFCFFB94FA15FB38E9DF" authorityName="Leme, Gonella &amp; D. R. Couto" authorityYear="2023" box="[1033,1182,1454,1480]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="17" pageNumber="56" phylum="Tracheophyta" rank="species" species="ribeiranus">
<emphasis id="2BA8DB0F2828FFCFFB94FA15FB38E9DF" box="[1033,1182,1454,1480]" italics="true" pageId="17" pageNumber="56">K. ribeiranus</emphasis>
</taxonomicName>
near small creeks under gallery forests, with a much smaller actually occupied area than the total AOO and the area of the range. This makes it highly vulnerable to the direct and indirect effects of agriculture and pasture expansion, which comprise nearly 26% of the total massif area. Additionally, 10.71% of the massif is utilized for temporary crops, which uses fire to clear the land and eventually promote intense fire events. Even in this scenario, however, the massif is a strong candidate for the creation of a private or public conservation unit, as the region’s deficit in terms of 
<emphasis id="2BA8DB0F2828FFCFFBC1F9D9FB04EA6B" box="[1116,1186,1635,1660]" italics="true" pageId="17" pageNumber="56">in situ</emphasis>
conservation strategies suggests.
</paragraph>
</subSubSection>
<subSubSection id="51C654962828FFCFFF20F910FC94EB42" pageId="17" pageNumber="56" type="etymology">
<paragraph id="1963071D2828FFCFFF20F910FC94EB42" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
<emphasis id="2BA8DB0F2828FFCFFF20F910FE98EAD3" bold="true" box="[189,318,1706,1732]" pageId="17" pageNumber="56">Etymology</emphasis>
:—The epithet of this new species honors its discoverer, the young naturalist Júlio Cesar dos Santos Ribeiro, that lives at the foothills of Serra do Padre Ângelo, Conselheiro Pena, 
<collectingRegion id="DB18C9FF2828FFCFFC68F974FB6BEAFF" box="[1013,1229,1742,1768]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="56">Minas Gerais State</collectingRegion>
. With his sharp eye, escalating skills, and great curiosity for the natural world, Mr. Ribeiro has dedicated himself intensively to the field survey of the species that make up the extremely rich flora of his homeland, having participated in the discovery of many of the novelties from the region, such as 
<taxonomicName id="DEDC7C9E2828FFCFFD0BF881FC8DEB43" authorityName="Leme, Gonella &amp; D. R. Couto" authorityYear="2023" box="[662,811,1850,1876]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="17" pageNumber="56" phylum="Tracheophyta" rank="species" species="ribeiranus">
<emphasis id="2BA8DB0F2828FFCFFD0BF881FC8DEB43" box="[662,811,1850,1876]" italics="true" pageId="17" pageNumber="56">K. ribeiranus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="51C654962828FFCCFF20F8E4FB6BEE7F" lastPageId="18" lastPageNumber="57" pageId="17" pageNumber="56" type="conservation">
<paragraph id="1963071D2828FFCFFF20F8E4FBD2E43B" blockId="17.[136,1452,158,2092]" pageId="17" pageNumber="56">
<emphasis id="2BA8DB0F2828FFCFFF20F8E4FE0BEB6F" bold="true" box="[189,429,1886,1912]" pageId="17" pageNumber="56">Conservation status</emphasis>
:—Critically Endangered—CR B1ab (ii,iii,v) + B2ab (ii,iii,v). Several threats affect both subpopulations, such as agriculture (coffee plantations and other temporary crops), deforestation due to cattle ranching and grazing, fires, which already affected one subpopulation, invasion by alien grasses (
<taxonomicName id="DEDC7C9E2828FFCFFB1AF81CFABCEBD6" box="[1159,1306,1958,1985]" class="Liliopsida" family="Poaceae" genus="Urochloa" kingdom="Plantae" order="Poales" pageId="17" pageNumber="56" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2BA8DB0F2828FFCFFB1AF81CFAB2EBD7" box="[1159,1300,1958,1984]" italics="true" pageId="17" pageNumber="56">Urochloa sp</emphasis>
.
</taxonomicName>
, used in the surrounding grazing areas), which can synergistically intensify fire events. Although targeted surveys in other areas of similar habitat may potentially reveal unrecorded subpopulations, so far, the species can be considered a narrow endemic bromeliad from the last significantly continuously forested massif of this region.
</paragraph>
<paragraph id="1963071D282BFFCCFF20FF24FD33EDEA" blockId="18.[136,1452,158,1625]" pageId="18" pageNumber="57">
Since the species is currently known only from two nearby collection points or subpopulations, it does not have an associated EOO polygon and has an estimated AOO of 
<quantity id="DE24AAF8282BFFCCFD71FF78FC81ECC8" box="[748,807,194,223]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="57" unit="km" value="4.0">4 km</quantity>
². Land use and land cover data retrieved from Mapbiomas (2022a, b) applied for the overlay analyses (Jord„o 
<emphasis id="2BA8DB0F282BFFCCFD47FF5DFCB3ED17" box="[730,789,230,256]" italics="true" pageId="18" pageNumber="57">et al.</emphasis>
2022) indicates that over 31% of its AOO was converted to pastures, and that 17.9% is currently designated as a mosaic of agriculture and pastureland formations (
<figureCitation id="81E71B98282BFFCCFADBFEB0FADBED33" box="[1350,1405,266,292]" captionStart="FIGURE 1" captionStartId="2.[136,229,1657,1679]" captionTargetBox="[180,1407,291,1617]" captionTargetId="figure-56@2.[167,1419,278,1630]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Krenakanthus ribeiranus and K. roseolilacinus. The Area of Occupancy (AOO) of K. ribeiranus in presented alongside the results of the overlay analyses with Mapbiomas land use cover data series." figureDoi="http://doi.org/10.5281/zenodo.8425693" httpUri="https://zenodo.org/record/8425693/files/figure.png" pageId="18" pageNumber="57">fig 1</figureCitation>
, 
<figureCitation id="81E71B98282BFFCCFA10FEB0FA38ED33" box="[1421,1438,266,292]" captionStart="FIGURE 2" captionStartId="3.[136,229,1048,1070]" captionTargetBox="[158,1428,197,1017]" captionTargetId="figure-18@3.[151,1436,190,1024]" captionTargetPageId="3" captionText="FIGURE 2. The massif of Serra de Santa Maria immediate delimitation and with buffers of 1 km, 3 km, and 5 km of its immediate surroundings. The overlay analyses of these geometries with Mapbiomas historical land use cover data series is presented for each land use and vegetation cover classes remaining proportion." figureDoi="http://doi.org/10.5281/zenodo.8425695" httpUri="https://zenodo.org/record/8425695/files/figure.png" pageId="18" pageNumber="57">2</figureCitation>
). Considering the intensity of the described stress vectors, one single location is considered, as these threats may extirpate the whole population if one single intense stress event occurs within its narrow and fragile habitat. Although we have an indication of the putative number of mature individuals for each known subpopulation, no data on generation length and the population trend is known, so the species could not be assessed based on its (
<emphasis id="2BA8DB0F282BFFCCFBCFFE21FB0FEDA3" box="[1106,1193,411,436]" italics="true" pageId="18" pageNumber="57">a priori</emphasis>
small) population size and reduction rate. All known subpopulations are outside protected areas, which might increase the vulnerability of the species in face of severe anthropogenic events.
</paragraph>
<paragraph id="1963071D282BFFCCFF20FDBCFB6BEE7F" blockId="18.[136,1452,158,1625]" pageId="18" pageNumber="57">Therefore, based on the minimum values of EOO and AOO, combined with one single location and an estimated continuing decline in its AOO, extent and quality of habitat, and possibly in the number of mature individuals, the species is here declared as Critically Endangered under the aforementioned IUCN (2012) criteria.</paragraph>
</subSubSection>
<subSubSection id="51C65496282BFFCCFF20FDC8FCB5EEEF" pageId="18" pageNumber="57" type="materials_examined">
<paragraph id="1963071D282BFFCCFF20FDC8FCB5EEEF" blockId="18.[136,1452,158,1625]" pageId="18" pageNumber="57">
<emphasis id="2BA8DB0F282BFFCCFF20FDC8FD15EE9B" bold="true" box="[189,691,626,652]" pageId="18" pageNumber="57">Additional specimens examined(paraypes)</emphasis>
:— 
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<collectingCountry id="61CB478D282BFFCCFD49FDC8FC99EE9B" box="[724,831,626,652]" name="Brazil" pageId="18" pageNumber="57">BRAZIL</collectingCountry>
. 
<collectingRegion id="DB18C9FF282BFFCCFCDBFDC8FC7AEE9B" box="[838,988,626,652]" country="Brazil" name="Minas Gerais" pageId="18" pageNumber="57">Minas Gerais</collectingRegion>
: 
<collectingCounty id="F0027F91282BFFCCFC7FFDC8FBFCEE9B" box="[994,1114,626,652]" pageId="18" pageNumber="57">Alvarenga</collectingCounty>
, 
<collectingMunicipality id="F9079D67282BFFCCFBFDFDC8FAE2EE9B" box="[1120,1348,626,652]" pageId="18" pageNumber="57">Serra de Santa Maria</collectingMunicipality>
, 
<location id="1C0351C6282BFFCCFAD6FDC8FF4EEEA7" LSID="urn:lsid:plazi:treatment:9175B60B2829FFCCFF15FBA9FC1AEA4F:1C0351C6282BFFCCFAD6FDC8FF4EEEA7" country="Brazil" county="Alvarenga" municipality="Serra de Santa Maria" name="Sobreiro de Cima" pageId="18" pageNumber="57" stateProvince="Minas Gerais">Sobreiro de Cima</location>
, maciço quartizítico a leste da sede do Município, [precise locality withheld for conservation purposes], 
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<elevation id="92F1E02E282BFFCCFAFBFD2CFA0AEEA6" box="[1382,1452,662,689]" metricMagnitude="2" metricUnit="m" metricValue="5.95" pageId="18" pageNumber="57" unit="m" value="595.0">595 m</elevation>
</quantity>
elevation, 
<date id="6D6221DD282BFFCCFF61FD00FE70EEC3" box="[252,470,698,724]" pageId="18" pageNumber="57" value="2022-11-27">
<collectingDate id="7D26D835282BFFCCFF61FD00FE70EEC3" box="[252,470,698,724]" pageId="18" pageNumber="57" value="2022-11-27">27 November 2022</collectingDate>
</date>
, 
<emphasis id="2BA8DB0F282BFFCCFE7DFD01FD0AEEC3" box="[480,684,698,725]" italics="true" pageId="18" pageNumber="57">
<collectorName id="B42962CB282BFFCCFE7DFD01FDDDEEC3" box="[480,635,698,725]" pageId="18" pageNumber="57">J.C.S. Ribeiro</collectorName>
001
</emphasis>
, cult. 
<emphasis id="2BA8DB0F282BFFCCFD6DFD01FC3EEEC3" box="[752,920,698,724]" italics="true" pageId="18" pageNumber="57">
<collectorName id="B42962CB282BFFCCFD6DFD01FCEAEEC3" box="[752,844,699,724]" pageId="18" pageNumber="57">E. Leme</collectorName>
10220
</emphasis>
(
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!);
</materialsCitation>
<materialsCitation id="A9B40D40282BFFCCFC70FD00FCA9EEEF" collectingDate="2022-11-27" collectingDateMax="2023-03-01" collectingDateMin="2022-11-27" collectionCode="RB" collectorName="P. M. Gonella &amp; D. P. Cordeiro" country="Brazil" county="Alvarenga" elevation="595" location="Sobreiro de Cima" municipality="Serra de Santa Maria" pageId="18" pageNumber="57" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
ibidem, 
<quantity id="DE24AAF8282BFFCCFBD5FD00FB29EEC2" box="[1096,1167,698,725]" metricMagnitude="2" metricUnit="m" metricValue="6.15" pageId="18" pageNumber="57" unit="m" value="615.0">
<elevation id="92F1E02E282BFFCCFBD5FD00FB29EEC2" box="[1096,1167,698,725]" metricMagnitude="2" metricUnit="m" metricValue="6.15" pageId="18" pageNumber="57" unit="m" value="615.0">615 m</elevation>
</quantity>
elevation, 
<date id="6D6221DD282BFFCCFA94FD00FA01EEC3" box="[1289,1447,698,724]" pageId="18" pageNumber="57" value="2023-03-01">
<collectingDate id="7D26D835282BFFCCFA94FD00FA01EEC3" box="[1289,1447,698,724]" pageId="18" pageNumber="57" value="2023-03-01">1 March 2023</collectingDate>
</date>
, 
<emphasis id="2BA8DB0F282BFFCCFF15FD65FD63EEEF" box="[136,709,734,761]" italics="true" pageId="18" pageNumber="57">
<collectorName id="B42962CB282BFFCCFF15FD65FEBAEEEF" box="[136,284,734,760]" pageId="18" pageNumber="57">P.M. Gonella</collectorName>
3800, 
<collectorName id="B42962CB282BFFCCFEF4FD65FDA3EEEF" box="[361,517,734,761]" pageId="18" pageNumber="57">J.C.S. Ribeiro</collectorName>
&amp; 
<collectorName id="B42962CB282BFFCCFDB4FD65FD63EEEF" box="[553,709,734,760]" pageId="18" pageNumber="57">D.P. Cordeiro</collectorName>
</emphasis>
(
<collectionCode id="7FCD9FD8282BFFCCFD48FD64FCA5EEEF" box="[725,771,734,760]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="18" pageNumber="57" type="Herbarium">RB</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="51C65496282BFFCCFF20FCB8FC1AEA4F" pageId="18" pageNumber="57" type="diagnosis">
<paragraph id="1963071D282BFFCCFF20FCB8FC61E903" blockId="18.[136,1452,158,1625]" pageId="18" pageNumber="57">
<emphasis id="2BA8DB0F282BFFCCFF20FCB8FE63EF0B" bold="true" box="[189,453,770,796]" pageId="18" pageNumber="57">Distinctive characters</emphasis>
:—The set of morphological features of this new species makes its circumscription in 
<taxonomicName id="DEDC7C9E282BFFCCFF15FC9CFE82EF57" box="[136,292,806,832]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="2BA8DB0F282BFFCCFF15FC9CFE82EF57" box="[136,292,806,832]" italics="true" pageId="18" pageNumber="57">Krenakanthus</emphasis>
</taxonomicName>
apparently challenging. This is true due to the several morphological differences when compared to the single known species of the genus, 
<taxonomicName id="DEDC7C9E282BFFCCFD80FCF1FD7BEF73" authorityName="Leme, Zizka &amp; Paule (Leme et al." authorityYear="2022" baseAuthorityName="Leme" baseAuthorityYear="2015" box="[541,733,842,868]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="species" species="roseolilacinus">
<emphasis id="2BA8DB0F282BFFCCFD80FCF1FD7BEF73" box="[541,733,842,868]" italics="true" pageId="18" pageNumber="57">K. roseolilacinus</emphasis>
</taxonomicName>
. These differences are: (a) stemless habit (
<figureCitation id="81E71B98282BFFCCFB58FCF0FAE8EF73" box="[1221,1358,842,868]" captionStart="FIGURE 3" captionStartId="4.[136,229,1996,2018]" captionTargetBox="[233,1354,389,1971]" captionTargetId="figure-121@4.[233,1354,389,1971]" captionTargetPageId="4" captionText="FIGURE 3. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. General aspect of the shaded rocky habitat alongside creeks at the type locality. B–D. Subpopulations at the type locality composed of individuals growing on organic-rich, shallow soils accumulated on sandstone rock surfaces among mosses. Photos: J.C.S. Ribeiro." figureDoi="http://doi.org/10.5281/zenodo.8425697" httpUri="https://zenodo.org/record/8425697/files/figure.png" pageId="18" pageNumber="57">figs. 3 C–D</figureCitation>
, 
<figureCitation id="81E71B98282BFFCCFAC7FCF0FF3AEF9E" captionStart="FIGURE 5" captionStartId="7.[136,229,1992,2014]" captionTargetBox="[196,1390,281,1967]" captionTargetId="figure-55@7.[196,1390,281,1967]" captionTargetPageId="7" captionText="FIGURE 5. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A Habit in three stages of flower development. B. Habit, highlighting an individual in fruit stage. C. Small-sized aspect of the leaf rosette in an adult individual. D. Frontal view of the fan blade-like corolla. E. Lateral view of the corolla. F. Abundant seedlings growing not far from mother-plants. Photos: J.C.S. Ribeiro." figureDoi="http://doi.org/10.5281/zenodo.8425701" httpUri="https://zenodo.org/record/8425701/files/figure.png" pageId="18" pageNumber="57">5 B–C, E</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFF36FCD5FF6FEF9F" box="[171,201,879,904]" italics="true" pageId="18" pageNumber="57">vs.</emphasis>
long caulescent) and much fewer leaves (7–12 
<emphasis id="2BA8DB0F282BFFCCFD6AFCD5FCA8EF9F" box="[759,782,879,904]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 
<quantity id="DE24AAF8282BFFCCFC82FCD4FC23EF9F" box="[799,901,878,904]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.858" metricValueMin="5.08" pageId="18" pageNumber="57" unit="in" value="23.5" valueMax="27.0" valueMin="20.0">20–27 in</quantity>
number); (b) leaf blades subdensely to densely covered by spreading, hair-like trichomes (
<figureCitation id="81E71B98282BFFCCFDE7FC28FCA0EFBB" box="[634,774,914,940]" captionStart="FIGURE 4" captionStartId="6.[136,229,1457,1479]" captionTargetBox="[151,1436,190,1433]" captionTargetId="figure-23@6.[151,1436,190,1433]" captionTargetPageId="6" captionText="FIGURE 4. Leaf margin and indumentum in Krenakanthus. A–B. Krenakanthus roseolilacinus. C–E. K. ribeiranus. A–B. Leaf margin with conspicuous spines. C–D. Leaf margin with inconspicuous spines and leaf blade with trichomes. E. Detail of a non-glandular uniseriate trichome. Arrowhead = spine. t = tector trichome. Bars: A–D = 1 mm. E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8425699" httpUri="https://zenodo.org/record/8425699/files/figure.png" pageId="18" pageNumber="57">figs. 4 C–D</figureCitation>
, 
<figureCitation id="81E71B98282BFFCCFC88FC29FCE5EFBB" box="[789,835,914,941]" captionStart="FIGURE 7" captionStartId="9.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-18@9.[178,1409,190,1928]" captionTargetPageId="9" captionText="FIGURE 7. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. Side view of the leaf blade, highlighting the spreading, uniseriate, hair-like trichomes on the abaxial and adaxial surfaces. B. Basal portion of the leaf, evidencing the inconspicuous marginal spines. C. The inconspicuous peduncle of the inflorescence. D. Fruits in different stages of maturation. E. Fruits. F. Seeds. Bars = 5 mm (A–B, D–E). Bars = 2 mm (C, F)." figureDoi="http://doi.org/10.5281/zenodo.8425703" httpUri="https://zenodo.org/record/8425703/files/figure.png" pageId="18" pageNumber="57">7 A</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFCCFFC29FCCFEFBB" box="[850,873,915,940]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. glabrescent, with minute glandular trichomes, or glabrous, 
<figureCitation id="81E71B98282BFFCCFF66FC0CFEDAEFC7" box="[251,380,950,976]" captionStart="FIGURE 4" captionStartId="6.[136,229,1457,1479]" captionTargetBox="[151,1436,190,1433]" captionTargetId="figure-23@6.[151,1436,190,1433]" captionTargetPageId="6" captionText="FIGURE 4. Leaf margin and indumentum in Krenakanthus. A–B. Krenakanthus roseolilacinus. C–E. K. ribeiranus. A–B. Leaf margin with conspicuous spines. C–D. Leaf margin with inconspicuous spines and leaf blade with trichomes. E. Detail of a non-glandular uniseriate trichome. Arrowhead = spine. t = tector trichome. Bars: A–D = 1 mm. E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8425699" httpUri="https://zenodo.org/record/8425699/files/figure.png" pageId="18" pageNumber="57">fig. 4 A–B</figureCitation>
), smaller (7–25 × 
<quantity id="DE24AAF8282BFFCCFDC4FC0CFD7CEFC6" box="[601,730,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="18" pageNumber="57" unit="cm" value="1.3" valueMax="1.8" valueMin="0.8">0.8–1.8 cm</quantity>
<emphasis id="2BA8DB0F282BFFCCFD78FC0DFCA2EFC7" box="[741,772,951,976]" italics="true" pageId="18" pageNumber="57">vs.</emphasis>
17–27 × 
<quantity id="DE24AAF8282BFFCCFCE6FC0CFBA6EFC7" box="[891,1024,950,976]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="18" pageNumber="57" unit="cm" value="2.1" valueMax="2.4" valueMin="1.8">1.8–2.4 cm</quantity>
), thinner in texture, margins straight (
<figureCitation id="81E71B98282BFFCCFF0DFC60FEAAEFE3" box="[144,268,986,1013]" captionStart="FIGURE 5" captionStartId="7.[136,229,1992,2014]" captionTargetBox="[196,1390,281,1967]" captionTargetId="figure-55@7.[196,1390,281,1967]" captionTargetPageId="7" captionText="FIGURE 5. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A Habit in three stages of flower development. B. Habit, highlighting an individual in fruit stage. C. Small-sized aspect of the leaf rosette in an adult individual. D. Frontal view of the fan blade-like corolla. E. Lateral view of the corolla. F. Abundant seedlings growing not far from mother-plants. Photos: J.C.S. Ribeiro." figureDoi="http://doi.org/10.5281/zenodo.8425701" httpUri="https://zenodo.org/record/8425701/files/figure.png" pageId="18" pageNumber="57">fig. 5 A–B</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFE84FC61FE96EFE3" box="[281,304,987,1012]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. undulate mainly toward the base), inconspicuously spinulose and appearing entire (
<figureCitation id="81E71B98282BFFCCFB73FC60FAEEEFE3" box="[1262,1352,986,1012]" captionStart="FIGURE 4" captionStartId="6.[136,229,1457,1479]" captionTargetBox="[151,1436,190,1433]" captionTargetId="figure-23@6.[151,1436,190,1433]" captionTargetPageId="6" captionText="FIGURE 4. Leaf margin and indumentum in Krenakanthus. A–B. Krenakanthus roseolilacinus. C–E. K. ribeiranus. A–B. Leaf margin with conspicuous spines. C–D. Leaf margin with inconspicuous spines and leaf blade with trichomes. E. Detail of a non-glandular uniseriate trichome. Arrowhead = spine. t = tector trichome. Bars: A–D = 1 mm. E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8425699" httpUri="https://zenodo.org/record/8425699/files/figure.png" pageId="18" pageNumber="57">fig. 4 C</figureCitation>
, 
<figureCitation id="81E71B98282BFFCCFAC9FC61FA26EFE2" box="[1364,1408,987,1013]" captionStart="FIGURE 7" captionStartId="9.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-18@9.[178,1409,190,1928]" captionTargetPageId="9" captionText="FIGURE 7. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. Side view of the leaf blade, highlighting the spreading, uniseriate, hair-like trichomes on the abaxial and adaxial surfaces. B. Basal portion of the leaf, evidencing the inconspicuous marginal spines. C. The inconspicuous peduncle of the inflorescence. D. Fruits in different stages of maturation. E. Fruits. F. Seeds. Bars = 5 mm (A–B, D–E). Bars = 2 mm (C, F)." figureDoi="http://doi.org/10.5281/zenodo.8425703" httpUri="https://zenodo.org/record/8425703/files/figure.png" pageId="18" pageNumber="57">7 B</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFA10FC61FA0DEFE3" box="[1421,1451,987,1012]" italics="true" pageId="18" pageNumber="57">vs.</emphasis>
distinctly spinulose, 
<figureCitation id="81E71B98282BFFCCFEF3FC44FE40E80F" box="[366,486,1022,1048]" captionStart="FIGURE 4" captionStartId="6.[136,229,1457,1479]" captionTargetBox="[151,1436,190,1433]" captionTargetId="figure-23@6.[151,1436,190,1433]" captionTargetPageId="6" captionText="FIGURE 4. Leaf margin and indumentum in Krenakanthus. A–B. Krenakanthus roseolilacinus. C–E. K. ribeiranus. A–B. Leaf margin with conspicuous spines. C–D. Leaf margin with inconspicuous spines and leaf blade with trichomes. E. Detail of a non-glandular uniseriate trichome. Arrowhead = spine. t = tector trichome. Bars: A–D = 1 mm. E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8425699" httpUri="https://zenodo.org/record/8425699/files/figure.png" pageId="18" pageNumber="57">fig. 4 A–B</figureCitation>
), spines inconspicuous [
<quantity id="DE24AAF8282BFFCCFD69FC44FCDBE80E" box="[756,893,1022,1049]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="18" pageNumber="57" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
<emphasis id="2BA8DB0F282BFFCCFC1EFC45FC3CE80F" box="[899,922,1023,1048]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 0.3 (apical ones)– 
<quantity id="DE24AAF8282BFFCCFBECFC44FB16E80E" box="[1137,1200,1022,1049]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="57" unit="mm" value="3.0">3 mm</quantity>
(basal ones) long]; (c) inflorescence shortly pedunculate to subsessile (
<figureCitation id="81E71B98282BFFCCFD3FFB98FCBBE82B" box="[674,797,1058,1085]" captionStart="FIGURE 7" captionStartId="9.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-18@9.[178,1409,190,1928]" captionTargetPageId="9" captionText="FIGURE 7. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A. Side view of the leaf blade, highlighting the spreading, uniseriate, hair-like trichomes on the abaxial and adaxial surfaces. B. Basal portion of the leaf, evidencing the inconspicuous marginal spines. C. The inconspicuous peduncle of the inflorescence. D. Fruits in different stages of maturation. E. Fruits. F. Seeds. Bars = 5 mm (A–B, D–E). Bars = 2 mm (C, F)." figureDoi="http://doi.org/10.5281/zenodo.8425703" httpUri="https://zenodo.org/record/8425703/files/figure.png" pageId="18" pageNumber="57">fig. 7 C–D</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFCB5FB99FC99E82B" box="[808,831,1059,1084]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. sessile), simple (
<emphasis id="2BA8DB0F282BFFCCFB95FB99FBB9E82B" box="[1032,1055,1059,1084]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. sparsely branched only at the base and simples toward the apex); (d) flowers distinctly smaller (
<emphasis id="2BA8DB0F282BFFCCFCABFBFDFCF6E877" box="[822,848,1095,1120]" italics="true" pageId="18" pageNumber="57">ca</emphasis>
. 
<quantity id="DE24AAF8282BFFCCFCC3FBFCFC0BE876" box="[862,941,1094,1121]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="57" unit="mm" value="30.0">30 mm</quantity>
<emphasis id="2BA8DB0F282BFFCCFC29FBFDFC6DE877" box="[948,971,1095,1120]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 
<quantity id="DE24AAF8282BFFCCFC47FBFCFBF5E876" box="[986,1107,1094,1121]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="5.5" metricValueMin="4.2" pageId="18" pageNumber="57" unit="mm" value="48.5" valueMax="55.0" valueMin="42.0">42–55 mm</quantity>
long); sepals smaller (11–11.5 × 
<quantity id="DE24AAF8282BFFCCFF02FBD0FF50E892" box="[159,246,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="57" unit="mm" value="2.5">2.5 mm</quantity>
<emphasis id="2BA8DB0F282BFFCCFF60FBD1FEB2E893" box="[253,276,1131,1156]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 20–27 × 
<quantity id="DE24AAF8282BFFCCFE15FBD0FE4FE893" box="[392,489,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="18" pageNumber="57" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
), shorter connate at the base for 
<quantity id="DE24AAF8282BFFCCFCC5FBD0FC6CE892" box="[856,970,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="18" pageNumber="57" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
(
<figureCitation id="81E71B98282BFFCCFC47FBD0FB93E893" box="[986,1077,1130,1156]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 G</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFBDFFBD1FBFFE893" box="[1090,1113,1131,1156]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 
<quantity id="DE24AAF8282BFFCCFBF5FBD0FB61E892" box="[1128,1223,1130,1157]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="18" pageNumber="57" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
, 
<figureCitation id="81E71B98282BFFCCFB4EFBD0FA89E893" box="[1235,1327,1130,1156]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 H</figureCitation>
); (e) petals smaller (25–26 × 
<quantity id="DE24AAF8282BFFCCFECDFB34FE34E8BE" box="[336,402,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="57" unit="mm" value="8.0">8 mm</quantity>
<emphasis id="2BA8DB0F282BFFCCFE04FB35FE16E8BF" box="[409,432,1167,1192]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 35–44 × 
<quantity id="DE24AAF8282BFFCCFDBEFB34FD07E8BF" box="[547,673,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="18" pageNumber="57" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
), apex acute (
<figureCitation id="81E71B98282BFFCCFCA6FB34FC07E8BE" box="[827,929,1166,1193]" captionStart="FIGURE 5" captionStartId="7.[136,229,1992,2014]" captionTargetBox="[196,1390,281,1967]" captionTargetId="figure-55@7.[196,1390,281,1967]" captionTargetPageId="7" captionText="FIGURE 5. A–F. Krenakanthus ribeiranus (J.C.S. Ribeiro 001): A Habit in three stages of flower development. B. Habit, highlighting an individual in fruit stage. C. Small-sized aspect of the leaf rosette in an adult individual. D. Frontal view of the fan blade-like corolla. E. Lateral view of the corolla. F. Abundant seedlings growing not far from mother-plants. Photos: J.C.S. Ribeiro." figureDoi="http://doi.org/10.5281/zenodo.8425701" httpUri="https://zenodo.org/record/8425701/files/figure.png" pageId="18" pageNumber="57">figs. 5 D</figureCitation>
, 
<figureCitation id="81E71B98282BFFCCFC30FB34FC5CE8BF" box="[941,1018,1166,1192]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">6 A, C</figureCitation>
; 
<emphasis id="2BA8DB0F282BFFCCFB9BFB35FBBBE8BF" box="[1030,1053,1167,1192]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. acute to rounded, 
<figureCitation id="81E71B98282BFFCCFB6BFB34FAC8E8BF" box="[1270,1390,1166,1193]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 B, E</figureCitation>
), and exappendiculate (
<emphasis id="2BA8DB0F282BFFCCFED3FB09FEC3E8DB" box="[334,357,1203,1228]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. bearing distinct cupuliform appendages); (f) fruits smaller, 4 × 
<quantity id="DE24AAF8282BFFCCFBA9FB08FB00E8DA" box="[1076,1190,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="18" pageNumber="57" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 mm</quantity>
(
<emphasis id="2BA8DB0F282BFFCCFB28FB09FB6AE8DB" box="[1205,1228,1203,1228]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 8–10 × 
<quantity id="DE24AAF8282BFFCCFAADFB08FA39E8DB" box="[1328,1439,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="18" pageNumber="57" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">7–12 mm</quantity>
); (g) seeds smaller, 0.8–1.2 × 
<quantity id="DE24AAF8282BFFCCFE54FB6CFD86E8E6" box="[457,544,1238,1265]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="57" unit="mm" value="0.6">0.6 mm</quantity>
(
<emphasis id="2BA8DB0F282BFFCCFDACFB6DFDEEE8E7" box="[561,584,1239,1264]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. 2–3 × 
<quantity id="DE24AAF8282BFFCCFD3FFB6CFCBFE8E7" box="[674,793,1238,1264]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="18" pageNumber="57" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 mm</quantity>
), costal bands 8–12 cells wide, intercostal bands 1–3 cells wide (
<emphasis id="2BA8DB0F282BFFCCFF52FB41FF40E903" box="[207,230,1275,1300]" italics="true" pageId="18" pageNumber="57">vs</emphasis>
. costal bands 15–20 cells wide, intercostal bands 4–6 cells wide).
</paragraph>
<paragraph id="1963071D282BFFCCFF20FAA4FC1AEA4F" blockId="18.[136,1452,158,1625]" pageId="18" pageNumber="57">
Despite the relevant differences that distinguish these two species of 
<taxonomicName id="DEDC7C9E282BFFCCFC33FAA4FBECE92F" box="[942,1098,1310,1336]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="genus">
<emphasis id="2BA8DB0F282BFFCCFC33FAA4FBECE92F" box="[942,1098,1310,1336]" italics="true" pageId="18" pageNumber="57">Krenakanthus</emphasis>
</taxonomicName>
from each other, the similarities that unite 
<taxonomicName id="DEDC7C9E282BFFCCFF6AFAF9FE2BE94B" authorityName="Leme, Gonella &amp; D. R. Couto" authorityYear="2023" box="[247,397,1346,1372]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="species" species="ribeiranus">
<emphasis id="2BA8DB0F282BFFCCFF6AFAF9FE2BE94B" box="[247,397,1346,1372]" italics="true" pageId="18" pageNumber="57">K. ribeiranus</emphasis>
</taxonomicName>
and 
<taxonomicName id="DEDC7C9E282BFFCCFE5FFAF9FD26E94B" authorityName="Leme, Zizka &amp; Paule (Leme et al." authorityYear="2022" baseAuthorityName="Leme" baseAuthorityYear="2015" box="[450,640,1346,1372]" class="Liliopsida" family="Bromeliaceae" genus="Krenakanthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="18" pageNumber="57" phylum="Tracheophyta" rank="species" species="roseolilacinus">
<emphasis id="2BA8DB0F282BFFCCFE5FFAF9FD26E94B" box="[450,640,1346,1372]" italics="true" pageId="18" pageNumber="57">K. roseolilacinus</emphasis>
</taxonomicName>
are: (a) leaves thin to very thin in texture; (b) peduncle inconspicuous to absent); (c) sepals distinctly connate at the base; (d) petals similarly free, broadly spathulate from a very narrow blade (
<figureCitation id="81E71B98282BFFCCFF0DFA30FF4DE9B3" box="[144,235,1418,1444]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 C</figureCitation>
), blades spreading at anthesis and forming a fan blade-like corolla, and flaccidescent afterwards (
<figureCitation id="81E71B98282BFFCCFADFFA30FA38E9B3" box="[1346,1438,1418,1444]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 A</figureCitation>
); (e) stamens deeply included and not visible at anthesis (
<figureCitation id="81E71B98282BFFCCFC9EFA14FC26E9DF" box="[771,896,1454,1480]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 A–B</figureCitation>
); (f) filaments distinctly unequal in length (
<figureCitation id="81E71B98282BFFCCFAF3FA14FF64E9FB" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 C–D</figureCitation>
); (g) stigma conduplicate-spiral (
<figureCitation id="81E71B98282BFFCCFDA9FA68FD10E9FB" box="[564,694,1490,1516]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 M–N</figureCitation>
), being the only two representatives of the “Cryptanthoid complex” with such a stigma 
<typeStatus id="C667B9BF282BFFCCFEFCFA4DFE33EA06" box="[353,405,1527,1553]" pageId="18" pageNumber="57">type</typeStatus>
; (h) epigynous tube lacking; and (i) fruits with persistent sepals 2.5–2.9 times longer than the fruit length (
<figureCitation id="81E71B98282BFFCCFE8EF9A0FEDCEA23" box="[275,378,1562,1588]" captionStart="FIGURE 6" captionStartId="8.[136,229,1953,1975]" captionTargetBox="[178,1409,190,1928]" captionTargetId="figure-23@8.[178,1409,190,1928]" captionTargetPageId="8" captionText="FIGURE 6. Krenakanthus ribeiranus (J.C.S. Ribeiro 001; A, C, F–G, I, K, M) and K. roseolilacinus (Leme 8922; B, D–E, H, J, L, N): A–B. Frontal view of the corolla. C–D. Petals and stamens. E. Petal. F. Pistil and the stamens, highlighting the unequal filaments. G–H. Sepals. I–J. Fruits. K–L. Anthers. M–N. Stigma. Bars = 5 mm (C–E, J). Bars = 2 mm (F–I). Bars = 1 mm (K–N). Photos: A. J.C.S. Ribeiro. B–N. E. Leme." figureDoi="http://doi.org/10.5281/zenodo.8431275" httpUri="https://zenodo.org/record/8431275/files/figure.png" pageId="18" pageNumber="57">fig. 6 I–J</figureCitation>
). In addition, both species present similar habitats, occurring in the understory of Semideciduous Seasonal Forest in the region of Alvarenga and Conselheiro Pena (
<figureCitation id="81E71B98282BFFCCFCEDF984FC0DEA4F" box="[880,939,1598,1624]" captionStart="FIGURE 1" captionStartId="2.[136,229,1657,1679]" captionTargetBox="[180,1407,291,1617]" captionTargetId="figure-56@2.[167,1419,278,1630]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Krenakanthus ribeiranus and K. roseolilacinus. The Area of Occupancy (AOO) of K. ribeiranus in presented alongside the results of the overlay analyses with Mapbiomas land use cover data series." figureDoi="http://doi.org/10.5281/zenodo.8425693" httpUri="https://zenodo.org/record/8425693/files/figure.png" pageId="18" pageNumber="57">fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>