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<document id="7414921678D3BB400626832E45951E69" ID="10.11646/zootaxa.4144.1.1" ID-DOI="10.11646/zootaxa.4144.1.1" ID-GBIF-Dataset="c9808a7d-3696-45cc-a337-5448ce845ec6" ID-ISSN="1175-5326" ID-Zenodo-Dep="257666" ID-ZooBank="0BDD56E3-A657-46B2-B5E5-3CF7F8E8D3A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469513537727" checkinUser="plazi" docAuthor="Winterbottom, Richard" docDate="2016" docId="8962992CBA11FFC5FF4658DAA8E5FDF0" docLanguage="en" docName="zootaxa.4144.1.1.pdf" docOrigin="Zootaxa 4144 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acropora species" docType="key" docVersion="10" lastPageNumber="4" masterDocId="755BE154BA13FFC6FFD15D11AD79FFB7" masterDocTitle="Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific" masterLastPageNumber="53" masterPageNumber="1" pageNumber="3" updateTime="1698366168299" updateUser="plazi">
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<mods:title id="E99557599FF4479C653333E9587332B5">Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific</mods:title>
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<mods:namePart id="BDF97D53CA5C495C261515F0B9277E0F">Winterbottom, Richard</mods:namePart>
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<mods:date id="B43E303AB82C94DE1805B2BA2303DA99">2016</mods:date>
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<treatment id="8962992CBA11FFC5FF4658DAA8E5FDF0" ID-DOI="http://doi.org/10.5281/zenodo.6087301" ID-GBIF-Taxon="123207573" ID-Zenodo-Dep="6087301" LSID="urn:lsid:plazi:treatment:8962992CBA11FFC5FF4658DAA8E5FDF0" httpUri="http://treatment.plazi.org/id/8962992CBA11FFC5FF4658DAA8E5FDF0" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="49D17BB1BA11FFC4FF4658DAAE13FA52" box="[151,874,1483,1509]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="0174283ABA11FFC4FF4658DAAE13FA52" blockId="2.[151,874,1483,1509]" box="[151,874,1483,1509]" pageId="2" pageNumber="3">
<heading id="5A3C9F56BA11FFC4FF4658DAAE13FA52" bold="true" box="[151,874,1483,1509]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="33BFF428BA11FFC4FF4658DAAE13FA52" bold="true" box="[151,874,1483,1509]" pageId="2" pageNumber="3">
Key to the
<taxonomicName id="C6CB53B9BA11FFC4FECE58DAAC0CFA52" box="[287,373,1483,1509]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="3" phylum="Cnidaria" rank="species">species</taxonomicName>
of the
<taxonomicName id="C6CB53B9BA11FFC4FE1B58DAAFF2FA52" box="[458,651,1483,1509]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="33BFF428BA11FFC4FE1B58DAAFF2FA52" bold="true" box="[458,651,1483,1509]" italics="true" pageId="2" pageNumber="3">Trimma tevegae</emphasis>
</taxonomicName>
complex (adults).
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="49D17BB1BA11FFC5FF465B00A8E5FDF0" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="key">
<keyStep id="BA3F309FBA11FFC4FF465B00A8E5F899" pageId="2" pageNumber="3">
<paragraph id="0174283ABA11FFC4FF465B00A8E2F90E" blockId="2.[151,1436,1553,2014]" pageId="2" pageNumber="3">
<materialsCitation id="B1A32267BA11FFC4FF465B00A8E2F90E" ID-GBIF-Occurrence="1288048887" country="Palau" location="Sulawesi" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Sulawesi">
<keyLead id="BA3A8B0FBA11FFC4FF465B00A8E2F90E" pageId="2" pageNumber="3">
1a Anterior predorsal scale in the midline cycloid, second dorsal spine usually reaches the middle of the second dorsal fin, only 2 cephalic sensory papillae in row
<emphasis id="33BFF428BA11FFC4FDF25B3FAF53F9F4" box="[547,554,1582,1603]" italics="true" pageId="2" pageNumber="3">f</emphasis>
on the chin and in row
<emphasis id="33BFF428BA11FFC4FCC75B3FAE66F9F4" box="[790,799,1582,1603]" italics="true" pageId="2" pageNumber="3">r</emphasis>
on the top of the snout, usually 4 papillae in row
<emphasis id="33BFF428BA11FFC4FADD5B3FA861F9F4" box="[1292,1304,1582,1603]" italics="true" pageId="2" pageNumber="3">d</emphasis>
ʹ immediately behind the maxilla, no third (ventralmost) row of cheek scales, live colour with no blue stripe from anterior tip of snout to origin of first dorsal fin, lateral stripe continuing anteriorly over eye narrowed over pupil to form two spots connected by a thin line, preserved specimens lacking darker stripe in centre of snout, four distinct groupings of small round black melanophores situated at the 3, 4:30, 6 and 9 oclock positions around the eye...........
<taxonomicName id="C6CB53B9BA11FFC4FC075BB5A935F90E" ID-CoL="58V57" box="[982,1100,1700,1721]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="corerefum" status="sp. nov.">
<emphasis id="33BFF428BA11FFC4FC075BB5A935F90E" box="[982,1100,1700,1721]" italics="true" pageId="2" pageNumber="3">T. corerefum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="288C4953BA11FFC4FB855BB2A9FCF90E" box="[1108,1157,1699,1721]" pageId="2" pageNumber="3" rank="species">
<emphasis id="33BFF428BA11FFC4FB855BB2A9FCF90E" bold="true" box="[1108,1157,1699,1721]" pageId="2" pageNumber="3">n.sp.</emphasis>
</taxonomicNameLabel>
(
<collectingRegion id="C30FE6D8BA11FFC4FB415BB5A991F90E" box="[1168,1256,1700,1721]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="3">Sulawesi</collectingRegion>
to northern
<collectingCountry id="79DC68AABA11FFC4FA8F5BB5A8ECF90E" box="[1374,1429,1700,1721]" name="Palau" pageId="2" pageNumber="3">Palau</collectingCountry>
)
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="0174283ABA11FFC4FF465BD0A8E5F899" blockId="2.[151,1436,1553,2014]" pageId="2" pageNumber="3">
<keyLead id="BA3A8B0FBA11FFC4FF465BD0A8E5F899" pageId="2" pageNumber="3">
1b Anterior rows of midline predorsal scales cycloid or ctenoid, usually 4 or more cephalic sensory papillae in row
<emphasis id="33BFF428BA11FFC4FAF25BD0A853F961" box="[1315,1322,1729,1750]" italics="true" pageId="2" pageNumber="3">f</emphasis>
on the chin and in row
<emphasis id="33BFF428BA11FFC4FE9D5BCFAC2CF944" box="[332,341,1758,1779]" italics="true" pageId="2" pageNumber="3">r</emphasis>
on the top of the snout and 6 or more papillae in row
<emphasis id="33BFF428BA11FFC4FC825BCFAE26F944" box="[851,863,1758,1779]" italics="true" pageId="2" pageNumber="3">d</emphasis>
ʹ immediately behind the maxilla, third (ventralmost) row of cheek scales usually present, lateral stripe (if present) continuing anteriorly over eye, not narrowed over pupil to form two spots connected by a thin line, pigmentation around eye not in four distinct groups of small, round, black melanophores.... 2
</keyLead>
</paragraph>
</keyStep>
<paragraph id="0174283ABA11FFC4FF465A27A8E2F814" blockId="2.[151,1436,1553,2014]" pageId="2" pageNumber="3">
<keyStep id="BA3F309FBA11FFC4FF465A27A8E2F814" pageId="2" pageNumber="3">
<keyLead id="BA3A8B0FBA11FFC4FF465A27A8E2F814" pageId="2" pageNumber="3">
2a Second spine of first dorsal fin not reaching beyond base of third ray of second dorsal fin, no blue (live) or dark (preserved) stripe present on anterior midline of head, top and sides of snout densely pigmented with small, dark, rounded chromatophores mixed with slightly larger, amorphous, lighter brown chromatophores and a sprinkling of much larger, darker, rounded chromatophores, a dark spot behind symphysis of lower jaw..........................
<taxonomicName id="C6CB53B9BA11FFC4FBEB5A9FA9E2F814" ID-CoL="58V7Q" box="[1082,1179,1934,1955]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="33BFF428BA11FFC4FBEB5A9FA9E2F814" box="[1082,1179,1934,1955]" italics="true" pageId="2" pageNumber="3">T. tevegae</emphasis>
</taxonomicName>
(Solomons to Philippines)
</keyLead>
</keyStep>
</paragraph>
<paragraph id="0174283ABA11FFC4FF465ABDA8E2F869" blockId="2.[151,1436,1553,2014]" pageId="2" pageNumber="3">
<keyLead id="BA3A8B0FBA11FFC4FF465ABDA8E2F869" pageId="2" pageNumber="3">
2b Second spine of first dorsal fin usually reaching well beyond base of third ray of second dorsal fin (except in
<taxonomicName id="C6CB53B9BA11FFC4FAD55ABDA805F876" ID-CoL="58V67" box="[1284,1404,1964,1985]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="hollemani">
<emphasis id="33BFF428BA11FFC4FAD55ABDA805F876" box="[1284,1404,1964,1985]" italics="true" pageId="2" pageNumber="3">T. hollemani</emphasis>
</taxonomicName>
), a blue (live) or dark (preserved) stripe present on anterior midline of head, top and sides of snout not as above, a dark spot
</keyLead>
</paragraph>
<paragraph id="0174283ABA10FFC5FF305D86A8E2FE95" blockId="3.[151,1436,151,584]" pageId="3" pageNumber="4">
behind symphysis of lower jaw usually absent............................................................... 3
<keyStep id="BA3F309FBA10FFC5FF465DA4A8E2FE95" pageId="3" pageNumber="4">
<keyLead id="BA3A8B0FBA10FFC5FF465DA4A8E2FE95" pageId="3" pageNumber="4">
3a Live and freshly collected specimens with a blue spot in front of eye, another on opercle behind ventral margin of pupil and a blue stripe bordered dorsally with red below eye from tip of maxilla to at least bend of preopercle, these areas dark in preserved specimens, second dorsal spine elongate, usually reaching beyond end of second dorsal fin.......................................................................
<taxonomicName id="C6CB53B9BA10FFC5FD325C1CAEE6FE95" ID-CoL="58V4Z" box="[739,927,269,290]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="caudomaculatum">
<emphasis id="33BFF428BA10FFC5FD325C1CAEE6FE95" box="[739,927,269,290]" italics="true" pageId="3" pageNumber="4">T. caudomaculatum</emphasis>
</taxonomicName>
(Solomons to Japan, apparently Saipan to Maldives)
</keyLead>
</keyStep>
</paragraph>
<paragraph id="0174283ABA10FFC5FF465C3BA8E5FEEB" blockId="3.[151,1436,151,584]" pageId="3" pageNumber="4">
<materialsCitation id="B1A32267BA10FFC5FF465C3BA8E5FEEB" ID-GBIF-Occurrence="1288048864" location="No" pageId="3" pageNumber="4" specimenCount="1" stateProvince="No">
<keyLead id="BA3A8B0FBA10FFC5FF465C3BA8E5FEEB" pageId="3" pageNumber="4">
3b
<collectingRegion id="C30FE6D8BA10FFC5FF305C3BAD87FE88" box="[225,254,298,319]" country="Maldives" name="Uthuru" pageId="3" pageNumber="4">No</collectingRegion>
blue spot on opercle or blue stripe below eye (may be a white stripe) in live/fresh specimens, these areas not dark in preserved material, second spine of first dorsal fin usually not reaching to end of second dorsal fin...................... 4
</keyLead>
</materialsCitation>
</paragraph>
<keyStep id="BA3F309FBA10FFC5FF465C74A8E5FDF0" pageId="3" pageNumber="4">
<paragraph id="0174283ABA10FFC5FF465C74A8E2FE65" blockId="3.[151,1436,151,584]" pageId="3" pageNumber="4">
<materialsCitation id="B1A32267BA10FFC5FF465C74A8E2FE65" ID-GBIF-Occurrence="1288048881" country="Philippines" location="Sulawesi" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Sulawesi">
<keyLead id="BA3A8B0FBA10FFC5FF465C74A8E2FE65" pageId="3" pageNumber="4">
4a Second spine of second dorsal fin not reaching beyond base of third ray of second dorsal fin, usually 5 papillae in row
<emphasis id="33BFF428BA10FFC5FA8D5C74A81DFECD" box="[1372,1380,357,378]" italics="true" pageId="3" pageNumber="4">r</emphasis>
, dark stripe in midline of snout in preserved material often made up of amorphous large brown chromatophores, no broad diffuse internal dark stripe over the abdominal cavity which narrows and continues posteriorly on and just below the vertebral column...............................................
<taxonomicName id="C6CB53B9BA10FFC5FCDC5CACAEF1FE65" ID-CoL="58V67" box="[781,904,445,466]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="hollemani" status="sp. nov.">
<emphasis id="33BFF428BA10FFC5FCDC5CACAEF1FE65" box="[781,904,445,466]" italics="true" pageId="3" pageNumber="4">T. hollemani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="288C4953BA10FFC5FC5E5CADAEBFFE65" box="[911,966,444,466]" pageId="3" pageNumber="4" rank="species">
<emphasis id="33BFF428BA10FFC5FC5E5CADAEBFFE65" bold="true" box="[911,966,444,466]" pageId="3" pageNumber="4">n. sp.</emphasis>
</taxonomicNameLabel>
(Solomons to
<collectingRegion id="C30FE6D8BA10FFC5FB835CACA9D1FE65" box="[1106,1192,445,466]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="4">Sulawesi</collectingRegion>
and north to
<collectingCountry id="79DC68AABA10FFC5FAF95CACA8EFFE65" box="[1320,1430,445,466]" name="Philippines" pageId="3" pageNumber="4">Philippines</collectingCountry>
)
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="0174283ABA10FFC5FF465CCBA8E5FDF0" blockId="3.[151,1436,151,584]" pageId="3" pageNumber="4">
<keyLead id="BA3A8B0FBA10FFC5FF465CCBA8E5FDF0" pageId="3" pageNumber="4">
4b Second spine of second dorsal fin usually reaching to base of last ray of second dorsal fin or beyond, usually 4 papillae in row
<emphasis id="33BFF428BA10FFC5FF305CE6AD90FDBB" box="[225,233,503,524]" italics="true" pageId="3" pageNumber="4">r</emphasis>
, dark stripe in midline of snout in preserved material often made up of tiny round black melanophores, a broad diffuse internal dark stripe over the abdominal cavity which narrows and continues posteriorly on and just below the vertebral column..............................................................................
<taxonomicName id="C6CB53B9BA10FFC5FBBD5F23A99AFDF0" box="[1132,1251,562,583]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="burridgae" status="sp. nov.">
<emphasis id="33BFF428BA10FFC5FBBD5F23A99AFDF0" box="[1132,1251,562,583]" italics="true" pageId="3" pageNumber="4">T. burridgae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="288C4953BA10FFC5FB3B5F23A858FDFF" box="[1258,1313,562,584]" pageId="3" pageNumber="4" rank="species">
<emphasis id="33BFF428BA10FFC5FB3B5F23A858FDFF" bold="true" box="[1258,1313,562,584]" pageId="3" pageNumber="4">n. sp.</emphasis>
</taxonomicNameLabel>
(Palau only)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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