321 lines
37 KiB
XML
321 lines
37 KiB
XML
<document id="955448131498252B2AF5BF2DFE7B67CE" ID-CLB-Dataset="7627" ID-DOI="10.1206/0003-0090.438.1.1" ID-GBIF-Dataset="dde09a91-0a09-4f99-a10f-7bf4d47ba7b7" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613901" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615997249597" checkinUser="felipe" docAuthor="Álvarez-Padilla, Fernando, Kallal, Robert J. & Hormiga, Gustavo" docDate="2020" docId="881F35527677A35EFC8C6B99FF1FFE33" docLanguage="en" docName="BulAmeMusNatHis.2020.438.1-107.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2020 (438)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2020/issue-438/0003-0090.438.1.1/Taxonomy-and-Phylogenetics-of-Nanometinae-and-Other-Australasian-Orb-Weaving/10.1206/0003-0090.438.1.1.full" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Harlanethis Álvarez-Padilla & Kallal & Hormiga 2020, gen. nov." docType="treatment" docVersion="7" lastPageNumber="107" masterDocId="74264D2A7635A314FF9F6948FFEAFFF3" masterDocTitle="Taxonomy And Phylogenetics Of Nanometinae And Other Australasian Orb-Weaving Spiders (Araneae: Tetragnathidae)" masterLastPageNumber="107" masterPageNumber="1" pageNumber="1" updateTime="1698933942376" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="84C25316FF2D60E18918CFFEB27683D8">Taxonomy And Phylogenetics Of Nanometinae And Other Australasian Orb-Weaving Spiders (Araneae: Tetragnathidae)</mods:title>
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<mods:namePart id="CE0845ECB373BA8DEC07137BAE010562">Álvarez-Padilla, Fernando</mods:namePart>
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<mods:affiliation id="EA0604F061582B44D35231302F74A825">Departamento de Biología Comparada, Facultad de Ciencias, Universidad Nacional Autónoma de México</mods:affiliation>
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<mods:namePart id="4381C30983CBAB2A9EA189FA0E953D6E">Kallal, Robert J.</mods:namePart>
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<mods:affiliation id="583195BE5B60B26AE18A417DB8314DA1">Department of Biological Sciences, The George Washington University</mods:affiliation>
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<mods:namePart id="13AA79F2A2B0086E2BFBD0E01BEF90F7">Hormiga, Gustavo</mods:namePart>
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<mods:affiliation id="2BA983C5751BA310D8D168EC3CB3DD9E">Department of Biological Sciences, The George Washington University</mods:affiliation>
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<mods:title id="67CE94D7C0A68A933BE78A05630695EA">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="8931F108FE124F8E82E6AE2F558E4F95">2020</mods:date>
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<mods:number id="9FEE0BDB09571E2AE992F1BEAA410008">2020-02-28</mods:number>
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<treatment id="881F35527677A35EFC8C6B99FF1FFE33" ID-DOI="http://doi.org/10.5281/zenodo.4631733" ID-GBIF-Taxon="180670991" ID-Zenodo-Dep="4631733" LSID="urn:lsid:plazi:treatment:881F35527677A35EFC8C6B99FF1FFE33" httpUri="http://treatment.plazi.org/id/881F35527677A35EFC8C6B99FF1FFE33" lastPageId="74" pageId="66">
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<heading id="5B4133287677A356FC8C6B99FBBCFD1B" box="[787,1110,720,745]" centered="true" fontSize="9" level="2" pageId="66" reason="2">
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Genus
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<emphasis id="32C258567677A356FCFF6B98FC06FD1A" bold="true" box="[864,1004,720,745]" pageId="66">
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<taxonomicName id="C7B6FFC77677A356FCFF6B98FC0FFD1B" ID-CoL="98X5K" ID-ENA="2976741" authority="Álvarez-Padilla & Kallal & Hormiga, 2020" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[864,997,720,744]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="66" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="32C258567677A356FCFF6B98FC0FFD1B" bold="true" box="[864,997,720,744]" italics="true" pageId="66">Harlanethis</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="29F1E52D7677A356FC6B6B99FBBCFD1B" box="[1012,1110,721,745]" pageId="66" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="48ACD7CF7677A356FCCC6A4AFBFFFCE9" box="[851,1045,770,794]" pageId="66" type="description">
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<paragraph id="000984447677A356FCCC6A4AFBFFFCE9" blockId="66.[851,1045,770,794]" box="[851,1045,770,794]" pageId="66">
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<heading id="5B4133287677A356FCCC6A4AFBFFFCE9" box="[851,1045,770,794]" centered="true" fontSize="9" level="2" pageId="66" reason="2">
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<figureCitation id="988D98C17677A356FCCC6A4AFC05FCE9" box="[851,1007,770,794]" captionStart-0="FIGURE 38" captionStart-1="FIGURE 39" captionStart-2="FIGURE 41" captionStart-3="FIGURE 42" captionStart-4="FIGURE 43" captionStart-5="FIGURE 44" captionStartId-0="67.[108,194,1525,1546]" captionStartId-1="68.[107,193,1620,1641]" captionStartId-2="71.[106,192,1624,1645]" captionStartId-3="72.[108,194,1653,1674]" captionStartId-4="73.[105,192,1677,1698]" captionStartId-5="75.[108,194,1636,1657]" captionTargetBox-0="[126,1195,224,1479]" captionTargetBox-1="[187,1130,224,1593]" captionTargetBox-2="[149,1171,224,1597]" captionTargetBox-3="[157,1155,224,1627]" captionTargetBox-4="[170,1143,248,1645]" captionTargetBox-5="[156,1181,267,1596]" captionTargetId-0="figure-16@67.[120,774,751,1066]" captionTargetId-1="figure-16@68.[655,1133,833,1216]" captionTargetId-2="figure-57@71.[147,800,590,1002]" captionTargetId-3="figure-107@72.[483,819,697,1170]" captionTargetId-4="figure-67@73.[547,866,637,1187]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionTargetPageId-2="71" captionTargetPageId-3="72" captionTargetPageId-4="73" captionTargetPageId-5="75" captionText-0="FIGURE 38. Harlanethis weintrauborum (male: TEAU016; female: TEAU017). A. Female, dorsal habitus. B. Female, frontal habitus. C. Female, ventral habitus. D. Female, lateral habitus. E. Male, frontal habitus. F. Female, chelicera ventral view. G. Male, frontal view of endites with stridulating cuticle highlighted with arrow. H. Male, chelicerae ventral view with striae highlighted with arrow. Scale bars: A, C, D, 1 mm; B, –E, 0.5 mm; F–H, 0.2 mm." captionText-1="FIGURE 39. Harlanethis lipscombae (female: TEAU019, TEAU048; male: TEAU038; A, C, E, G, H) and H. weintrauborum (TEAU017, TEAU050; B, D, F). A, B. Epigynum, lateral, with arrow indicating anterior margin of epigynal protrusion. C, D. Epigynum, ventral. E, F. Epigynum, dorsal digested. G. Pedipalp, expanded, apical. H. Pedipalp, expanded, ventral. Scale bars: 0.2 mm." captionText-2="FIGURE 41. Harlanethis lipscombae (TEAU038). A. Epigynum, ventral. B. Embolus and conductor detail. C. Conductor and embolus, apical. D. Epigynum, lateral. E. Epigynum, dorsolateral. F. Pedipalp, CDBP detail. G. Femur IV trichobothria. H. Male, cheliceral striae. Scale bars: A–G, 0.1 mm; H, 0.05 mm." captionText-3="FIGURE 42. Harlanethis lipscombae (KS 52425; TEAU038). A. Pedipalp, ventral. B. Pedipalp, ectal. C. Pedipalp, dorsal. D. Pedipalp, ectodorsal. E. Conductor and embolus, ventral detail. F. Pedipalp CDBP, ventral detail. G. Pedipalp expanded, apical. Scale bars: 0.5 mm." captionText-4="FIGURE 43. Harlanethis weintrauborum (KS 0503; TEAU016). A. Pedipalp, ventral. B. Pedipalp, ectal. C. Pedipalp, dorsal. D. Pedipalp, ectodorsal. E. Pedipalp, CDBP detail. F. Conductor and embolus ventral detail. Scale bars: 0.5 mm." captionText-5="FIGURE 44. Harlanethis epigynum illustrations (TEAU037; TEAU050). A. H. lipscombae, ventral. B. H. weintrauborum, ventral. C. H. lipscombae, dorsal. D. H. weintrauborum, dorsal. E. H. lipscombae, lateral. F. H. lipscombae, ventral. G. H. weintrauborum, lateral. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4613982" figureDoi-1="http://doi.org/10.5281/zenodo.4613984" figureDoi-2="http://doi.org/10.5281/zenodo.4613986" figureDoi-3="http://doi.org/10.5281/zenodo.4613990" figureDoi-4="http://doi.org/10.5281/zenodo.4613994" figureDoi-5="http://doi.org/10.5281/zenodo.4613996" httpUri-0="https://zenodo.org/record/4613982/files/figure.png" httpUri-1="https://zenodo.org/record/4613984/files/figure.png" httpUri-2="https://zenodo.org/record/4613986/files/figure.png" httpUri-3="https://zenodo.org/record/4613990/files/figure.png" httpUri-4="https://zenodo.org/record/4613994/files/figure.png" httpUri-5="https://zenodo.org/record/4613996/files/figure.png" pageId="66">Figures 38–44</figureCitation>
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,
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<figureCitation id="988D98C17677A356FC646A4AFBFFFCE9" box="[1019,1045,770,794]" captionStart="FIGURE 60" captionStartId="96.[108,194,804,825]" captionTargetBox="[108,1205,224,776]" captionTargetId="figure-387@96.[108,1213,224,776]" captionTargetPageId="96" captionText="FIGURE 60. Distribution of new genera in Queensland. A. Harlanethis. B. Iamarra." figureDoi="http://doi.org/10.5281/zenodo.4614031" httpUri="https://zenodo.org/record/4614031/files/figure.png" pageId="66">60</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="48ACD7CF7677A356FD536A7CFB81FCB8" box="[716,1131,820,844]" pageId="66" type="materials_examined">
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<paragraph id="000984447677A356FD536A7CFB81FCB8" blockId="66.[684,1213,820,1737]" box="[716,1131,820,844]" pageId="66">
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<typeStatus id="DF0D3AE67677A356FD536A7CFCE9FCB8" box="[716,771,820,843]" pageId="66">TYPE</typeStatus>
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SPECIES:
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<taxonomicName id="C7B6FFC77677A356FCF36A7CFB8FFCBF" baseAuthorityName="GH" baseAuthorityYear="1144" box="[876,1125,820,844]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="66" phylum="Arthropoda" rank="species" species="lipscombae">
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<emphasis id="32C258567677A356FCF36A7CFB8FFCBF" box="[876,1125,820,844]" italics="true" pageId="66">Harlanethis lipscombae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="48ACD7CF7677A356FD536A1DFBEFFA12" pageId="66" type="diagnosis">
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<paragraph id="000984447677A356FD536A1DFBEFFA12" blockId="66.[684,1213,820,1737]" pageId="66">
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DIAGNOSIS:
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<taxonomicName id="C7B6FFC77677A356FCCD6A1DFC25FC9E" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[850,975,853,877]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="66" phylum="Arthropoda" rank="genus">
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<emphasis id="32C258567677A356FCCD6A1DFC25FC9E" box="[850,975,853,877]" italics="true" pageId="66">Harlanethis</emphasis>
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</taxonomicName>
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species are similar to other leucaugine genera in having femoral trichobothria and silver guanine abdominal patches (fig. 38A, C–D), but can be easily separated from other leucaugines by the presence some unique features among tetragnathids such a whitish, lightly sclerotized triangular epigynal protrusion (fig. 39C, D); two anterior longitudinal deep grooves on the epigynum venter (fig. 41A); and by having the epigynum almost the same size as the sternum (fig. 38C). Males are diagnosed by their massive conductor, which is wider than long and formed by three sclerites united by a membrane (figs. 39G, H, 41B, 42G, 43F). The embolus morphology is diagnostic, being very thick, tubular, slightly curved, and apically covered with small scales (figs. 41B, C, 42G). In males, the ventral cuticle of the paturon has four deep grooves intercalated with transverse ridges (figs. 38F, H, 41H).
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</paragraph>
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</subSubSection>
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<subSubSection id="48ACD7CF7677A352FD536CA2FC17FA11" lastPageId="70" pageId="66" type="description">
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<paragraph id="000984447677A356FD536CA2FB57F93A" blockId="66.[684,1213,820,1737]" pageId="66">DESCRIPTION: Female total length 3.99–5.25. Cephalothorax length 1.72–1.87, width 1.30–1.38. Carapace glabrous, pale yellow to light brown, slightly darker over the cephalic region and carapace margins. Fovea deep, triangular in shape, carapace dorsal pits absent (fig. 38A). Clypeus height 0.9 to 0.7 AME diameter, cuticle darker</paragraph>
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<caption id="54C9D4CC7676A357FFF36CBDFEBCF973" ID-DOI="http://doi.org/10.5281/zenodo.4613982" ID-Zenodo-Dep="4613982" httpUri="https://zenodo.org/record/4613982/files/figure.png" pageId="67" startId="67.[108,194,1525,1546]" targetBox="[126,1195,224,1479]" targetPageId="67">
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<paragraph id="000984447676A357FFF36CBDFEBCF973" blockId="67.[108,1215,1523,1665]" pageId="67">
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FIGURE 38.
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<taxonomicName id="C7B6FFC77676A357FF6F6CBBFDECF9F8" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[240,518,1523,1547]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="67" phylum="Arthropoda" rank="species" species="weintrauborum">
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<emphasis id="32C258567676A357FF6F6CBBFDECF9F8" box="[240,518,1523,1547]" italics="true" pageId="67">Harlanethis weintrauborum</emphasis>
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</taxonomicName>
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(male: TEAU016; female: TEAU017).
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<emphasis id="32C258567676A357FC0C6CBBFC46F9F8" bold="true" box="[915,940,1523,1547]" pageId="67">A.</emphasis>
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Female, dorsal habitus.
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<emphasis id="32C258567676A357FB396CBCFB56F9FF" bold="true" box="[1190,1212,1524,1548]" pageId="67">B.</emphasis>
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Female, frontal habitus.
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<emphasis id="32C258567676A357FEF56F59FE68F9DA" bold="true" box="[362,386,1553,1577]" pageId="67">C.</emphasis>
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Female, ventral habitus.
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<emphasis id="32C258567676A357FD146F59FD4EF9DA" bold="true" box="[651,676,1553,1577]" pageId="67">D.</emphasis>
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Female, lateral habitus.
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<emphasis id="32C258567676A357FC3A6F59FC50F9DA" bold="true" box="[933,954,1553,1577]" pageId="67">E.</emphasis>
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Male, frontal habitus.
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<emphasis id="32C258567676A357FB356F59FB55F9DA" bold="true" box="[1194,1215,1553,1577]" pageId="67">F.</emphasis>
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Female, chelicera ventral view.
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<emphasis id="32C258567676A357FE2A6F66FE24F9B5" bold="true" box="[437,462,1582,1606]" pageId="67">G.</emphasis>
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Male, frontal view of endites with stridulating cuticle highlighted with arrow.
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<emphasis id="32C258567676A357FF2D6F04FF27F997" bold="true" box="[178,205,1612,1636]" pageId="67">H.</emphasis>
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Male, chelicerae ventral view with striae highlighted with arrow. Scale bars: A, C, D, 1 mm; B, –E, 0.5 mm; F–H, 0.2 mm.
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</paragraph>
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</caption>
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<caption id="54C9D4CC7671A350FFF46F1CFD2AF931" ID-DOI="http://doi.org/10.5281/zenodo.4613984" ID-Zenodo-Dep="4613984" httpUri="https://zenodo.org/record/4613984/files/figure.png" pageId="68" startId="68.[107,193,1620,1641]" targetBox="[187,1130,224,1593]" targetPageId="68">
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<paragraph id="000984447671A350FFF46F1CFD2AF931" blockId="68.[107,1212,1618,1731]" pageId="68">
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FIGURE 39.
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<taxonomicName id="C7B6FFC77671A350FF6F6F1AFE33F999" baseAuthorityName="GH" baseAuthorityYear="1144" box="[240,473,1618,1642]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="68" phylum="Arthropoda" rank="species" species="lipscombae">
|
||
<emphasis id="32C258567671A350FF6F6F1AFE33F999" box="[240,473,1618,1642]" italics="true" pageId="68">Harlanethis lipscombae</emphasis>
|
||
</taxonomicName>
|
||
(female: TEAU019, TEAU048; male: TEAU038;
|
||
<emphasis id="32C258567671A350FC4D6F1AFB89F998" bold="true" box="[978,1123,1618,1643]" pageId="68">A, C, E, G, H</emphasis>
|
||
) and
|
||
<taxonomicName id="C7B6FFC77671A350FB3D6F1BFEEFF97B" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="68" phylum="Arthropoda" rank="species" species="weintrauborum">
|
||
<emphasis id="32C258567671A350FB3D6F1BFEEFF97B" italics="true" pageId="68">H. weintrauborum</emphasis>
|
||
</taxonomicName>
|
||
(TEAU017, TEAU050;
|
||
<emphasis id="32C258567671A350FE686F38FDAAF97B" bold="true" box="[503,576,1648,1672]" pageId="68">B, D, F</emphasis>
|
||
).
|
||
<emphasis id="32C258567671A350FDCA6F27FD60F97B" bold="true" box="[597,650,1647,1672]" pageId="68">A, B.</emphasis>
|
||
Epigynum, lateral, with arrow indicating anterior margin of epigynal protrusion.
|
||
<emphasis id="32C258567671A350FE1F6FC5FE5DF956" bold="true" box="[384,439,1677,1701]" pageId="68">C, D.</emphasis>
|
||
Epigynum, ventral.
|
||
<emphasis id="32C258567671A350FD1B6FC5FD5BF956" bold="true" box="[644,689,1677,1701]" pageId="68">E, F.</emphasis>
|
||
Epigynum, dorsal digested.
|
||
<emphasis id="32C258567671A350FC4F6FC5FC00F956" bold="true" box="[976,1002,1677,1701]" pageId="68">G.</emphasis>
|
||
Pedipalp, expanded, apical.
|
||
<emphasis id="32C258567671A350FF2D6FE3FF26F930" bold="true" box="[178,204,1707,1731]" pageId="68">H.</emphasis>
|
||
Pedipalp, expanded, ventral. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="000984447670A351FFF46F64FBDBF9AC" blockId="69.[107,1210,1578,1631]" pageId="69">
|
||
FIGURE 40.
|
||
<taxonomicName id="C7B6FFC77670A351FF716F62FE3CF9B1" baseAuthorityName="GH" baseAuthorityYear="1144" box="[238,470,1578,1602]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="69" phylum="Arthropoda" rank="species" species="lipscombae">
|
||
<emphasis id="32C258567670A351FF716F62FE3CF9B1" box="[238,470,1578,1602]" italics="true" pageId="69">Harlanethis lipscombae</emphasis>
|
||
</taxonomicName>
|
||
(TEAU038;
|
||
<emphasis id="32C258567670A351FDC76F62FD29F9B1" bold="true" box="[600,707,1578,1602]" pageId="69">A, C, E, G</emphasis>
|
||
), and
|
||
<taxonomicName id="C7B6FFC77670A351FC986F63FC2BF9B1" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[775,961,1578,1602]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="69" phylum="Arthropoda" rank="species" species="weintrauborum">
|
||
<emphasis id="32C258567670A351FC986F63FC2BF9B1" box="[775,961,1578,1602]" italics="true" pageId="69">H. weintrauborum</emphasis>
|
||
</taxonomicName>
|
||
(TEUA016;
|
||
<emphasis id="32C258567670A351FBDC6F62FB46F9B1" bold="true" box="[1091,1196,1578,1602]" pageId="69">B, D, F, H</emphasis>
|
||
).
|
||
<emphasis id="32C258567670A351FFF46F0FFF4AF9AC" bold="true" box="[107,160,1607,1631]" pageId="69">A, B.</emphasis>
|
||
Pedipalp, ventral.
|
||
<emphasis id="32C258567670A351FEC16F0FFE7FF9AC" bold="true" box="[350,405,1607,1631]" pageId="69">C, D.</emphasis>
|
||
Pedipalp, dorsal.
|
||
<emphasis id="32C258567670A351FDD56F0FFD92F9AC" bold="true" box="[586,632,1607,1631]" pageId="69">E, F.</emphasis>
|
||
Pedipalp, mesal.
|
||
<emphasis id="32C258567670A351FCB56F0FFC8EF9AC" bold="true" box="[810,868,1607,1631]" pageId="69">G, H.</emphasis>
|
||
Scale bars:
|
||
<quantity id="C74E29A17670A351FC436F0FFBC7F9AC" box="[988,1069,1607,1631]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="69" unit="mm" value="0.2">0.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="000984447673A352FFF469AAFEEDF9B6" blockId="70.[107,636,226,1738]" pageId="70">under the median eyes and paler under the lateral eyes (figs. 38B). AME slightly larger than PME. Lateral eyes on the same tubercle, one AME diameter apart from median eyes and approximately half that size. Chelicerae with few scattered setae, cuticle smooth, without ventral stridulatory ridges (fig. 38F), with three promarginal and two retromarginal teeth (figs. 38B, F). Endites longer than wide, dark brown, internal margins pale yellow. Labium rectangular, wider than long, colored as the endites. Sternum dark brown, trapezoidal in shape, wider between the first two legs (fig. 38C). Abdomen dorsally covered with silvery guanine patches over a background pale to dark gray, with a dark median longitudinal line, lateral surfaces with diffused dark vertical stripes; ventrally dark brown, with guanine patches concentrated in four spots forming a central rectangle (figs. 38A, C–D). Booklung covers without stridulatory striae. Spinnerets pale yellow to brown, lighter pattern on internal surfaces. Leg formula 1-2-4-3. Femur I length 2.96–3.67. First pair considerably larger, femora and tibia brown-yellow with dark-brown annuli. Second pair femora and tibiae pale yellow without annulation. Metatarsi and tarsi dark yellow. All leg segments dark brown apically. Femora III and IV with ca. six prolateral trichobothria, distributed around the base, flagellum smooth (fig. 41G). Femora with few setae, other segments hirsute increasing after tibiae. All leg segments with few macrosetae, except tarsi, which are glabrous. Epigynum: trapezoidal shape wider than long. Copulatory openings medially located on both sides of posterior margin of triangular epigynal protrusion (fig. 41A, D). Spermathecae membranous, internal surfaces covered with accessory gland ducts (figs. 39E, F, 41E). Copulatory ducts short, enclosing the spermathecae entrance, both copulatory ducts converging into a common sclerotized base where the fertilization ducts also originate (figs. 39E, F, 44C–D). Fertilization ducts short, straight.</paragraph>
|
||
<paragraph id="000984447673A352FF146F06FC97FC01" blockId="70.[107,636,226,1738]" lastBlockId="70.[684,1213,226,1738]" pageId="70">Male same as female except as noted. Total length 3.86–4.15. Cephalothorax length 1.71– 1.76, width 1.35–1.46. Carapace dorsal coloration white to dark yellow. Clypeus height 0.83–0.91 AME diameter, pale yellow. Sternum pale yellow to brown. Chelicerae pale to dark brown, slightly longer and narrower, dorsal cuticle rugose, setal bases enlarged and more abundant at base; ectal cuticle less rugose toward the ventral and apical surfaces (fig. 38E); ventral cuticle with four deep grooves intercalated with transverse ridges (figs. 38H, 41H); with three promarginal and three to four retromarginal teeth (fig. 38E, H). Endites brown; dorsal cuticle without obvious complementary structures to the cheliceral striae (fig. 38G). Abdominal pattern as in female, but lighter in coloration. Femur I length 3.65–4.17. Legs pale yellow, annulation less pronounced than in female. Pedipalpal tibia triangular, as long as wide, apical margin wider. CDBP shaped as a long curved ridge with its basal portion forming a small triangular spur (figs. 40C–F, 41F, 42C). Paracymbium cylindrical, almost glabrous, slightly curved, apically swollen, its cymbium attachment membranous (fig. 38G, H). Embolus very thick, tubular, slightly curved and apically covered with small scales (fig. 41B, C).</paragraph>
|
||
<paragraph id="000984447673A352FD536AB3FCB2FBA6" blockId="70.[684,1213,226,1738]" pageId="70">
|
||
COMPOSITION: This new genus includes two species:
|
||
<taxonomicName id="C7B6FFC77673A352FCBF6D54FBABFBC7" baseAuthorityName="GH" baseAuthorityYear="1144" box="[800,1089,1052,1076]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="70" phylum="Arthropoda" rank="species" species="lipscombae">
|
||
<emphasis id="32C258567673A352FCBF6D54FBABFBC7" box="[800,1089,1052,1076]" italics="true" pageId="70">Harlanethis lipscombae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C7B6FFC77673A352FB3F6D55FCB8FBA6" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="70" phylum="Arthropoda" rank="species" species="weintrauborum">
|
||
<emphasis id="32C258567673A352FB3F6D55FCB8FBA6" italics="true" pageId="70">H. weintrauborum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="000984447673A352FD536D17FC17FA11" blockId="70.[684,1213,226,1738]" pageId="70">
|
||
SYSTEMATICS: This genus is placed in the subfamily
|
||
<taxonomicName id="C7B6FFC77673A352FD616D37FC60FB64" authorityName="CAPORIACCO" authorityYear="1955" box="[766,906,1151,1175]" pageId="70" rank="subFamily" subFamily="Leucauginae">Leucauginae</taxonomicName>
|
||
(figs. 61–63), nested in a clade including
|
||
<taxonomicName id="C7B6FFC77673A352FCE06DE9FB5CFB4A" authority="Kulczynski, 1911" authorityName="Kulczynski" authorityYear="1911" box="[895,1206,1185,1209]" class="Arachnida" family="Tetragnathidae" genus="Mesida" kingdom="Animalia" order="Araneae" pageId="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="32C258567673A352FCE06DE9FC3AFB4A" box="[895,976,1185,1209]" italics="true" pageId="70">Mesida</emphasis>
|
||
Kulczyński, 1911
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7B6FFC77673A352FD336D8AFC7FFB29" authority="Simon, 1894" authorityName="Simon" authorityYear="1894" box="[684,917,1218,1242]" class="Arachnida" family="Tetragnathidae" genus="Tylorida" kingdom="Animalia" order="Araneae" pageId="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="32C258567673A352FD336D8AFCECFB29" box="[684,774,1218,1242]" italics="true" pageId="70">Tylorida</emphasis>
|
||
Simon, 1894
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C7B6FFC77673A352FC506D8AFBDDFB2A" authorityName="Thorell" authorityYear="1890" box="[975,1079,1218,1241]" class="Arachnida" family="Tetragnathidae" genus="Orsinome" kingdom="Animalia" order="Araneae" pageId="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="32C258567673A352FC506D8AFBDDFB2A" box="[975,1079,1218,1241]" italics="true" pageId="70">Orsinome</emphasis>
|
||
</taxonomicName>
|
||
. The monophyly of
|
||
<taxonomicName id="C7B6FFC77673A352FC916DABFC61FB08" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[782,907,1251,1275]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="32C258567673A352FC916DABFC61FB08" box="[782,907,1251,1275]" italics="true" pageId="70">Harlanethis</emphasis>
|
||
</taxonomicName>
|
||
is supported by the following morphological synapomorphies: massive highly sclerotized conductor, wider than long and formed by three sclerites united by a membrane (figs. 38G, H, 42E, G) and the large lightly sclerotized triangular epigynal protrusion with two anterior longitudinal deep grooves on the epigynum venter (fig. 39A–D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="48ACD7CF7673A35EFD536CA3FE8DFEE8" lastPageId="74" pageId="70" type="etymology">
|
||
<paragraph id="000984447673A352FD536CA3FB56F939" blockId="70.[684,1213,226,1738]" pageId="70">
|
||
ETYMOLOGY: TThis new genus is named by the third author (G.H.) to honor Wilbur B. (Bill) Harlan, whose endowment to the Department Biological Sciences of the George
|
||
<collectingRegion id="C2724AA67673A352FC556F06FBA2F995" box="[970,1096,1614,1638]" country="United States of America" name="Washington" pageId="70">Washington</collectingRegion>
|
||
University has empowered numerous young biologists scholarly work. This name is compounded with
|
||
<emphasis id="32C258567673A352FBE56FD8FB5CF95B" box="[1146,1206,1680,1704]" italics="true" pageId="70">nethis</emphasis>
|
||
, the Greek work for spinster, i.e., a woman who
|
||
</paragraph>
|
||
<caption id="54C9D4CC7672A353FFF56F10FBF4F95B" ID-DOI="http://doi.org/10.5281/zenodo.4613986" ID-Zenodo-Dep="4613986" httpUri="https://zenodo.org/record/4613986/files/figure.png" pageId="71" startId="71.[106,192,1624,1645]" targetBox="[149,1171,224,1597]" targetPageId="71">
|
||
<paragraph id="000984447672A353FFF56F10FBF4F95B" blockId="71.[106,1215,1621,1704]" pageId="71">
|
||
FIGURE 41.
|
||
<taxonomicName id="C7B6FFC77672A353FF726F1EFE3FF99D" baseAuthorityName="GH" baseAuthorityYear="1144" box="[237,469,1622,1646]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="71" phylum="Arthropoda" rank="species" species="lipscombae">
|
||
<emphasis id="32C258567672A353FF726F1EFE3FF99D" box="[237,469,1622,1646]" italics="true" pageId="71">Harlanethis lipscombae</emphasis>
|
||
</taxonomicName>
|
||
(TEAU038).
|
||
<emphasis id="32C258567672A353FDC06F1DFD92F99E" bold="true" box="[607,632,1621,1645]" pageId="71">A.</emphasis>
|
||
Epigynum, ventral.
|
||
<emphasis id="32C258567672A353FCD76F1EFCB4F99D" bold="true" box="[840,862,1622,1646]" pageId="71">B.</emphasis>
|
||
Embolus and conductor detail.
|
||
<emphasis id="32C258567672A353FB396F1DFB54F99E" bold="true" box="[1190,1214,1621,1645]" pageId="71">C.</emphasis>
|
||
Conductor and embolus, apical.
|
||
<emphasis id="32C258567672A353FE246F3BFE3FF978" bold="true" box="[443,469,1651,1675]" pageId="71">D.</emphasis>
|
||
Epigynum, lateral.
|
||
<emphasis id="32C258567672A353FD006F3BFD5EF978" bold="true" box="[671,692,1651,1675]" pageId="71">E.</emphasis>
|
||
Epigynum, dorsolateral.
|
||
<emphasis id="32C258567672A353FC266F3BFC24F978" bold="true" box="[953,974,1651,1675]" pageId="71">F.</emphasis>
|
||
Pedipalp, CDBP detail.
|
||
<emphasis id="32C258567672A353FFF56FD8FF69F95B" bold="true" box="[106,131,1680,1704]" pageId="71">G.</emphasis>
|
||
Femur IV trichobothria.
|
||
<emphasis id="32C258567672A353FE166FD8FE49F95B" bold="true" box="[393,419,1680,1704]" pageId="71">H.</emphasis>
|
||
Male, cheliceral striae. Scale bars: A–G, 0.1 mm; H, 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="54C9D4CC767DA35CFFF36F3DFD46F936" ID-DOI="http://doi.org/10.5281/zenodo.4613990" ID-Zenodo-Dep="4613990" httpUri="https://zenodo.org/record/4613990/files/figure.png" pageId="72" startId="72.[108,194,1653,1674]" targetBox="[157,1155,224,1627]" targetPageId="72">
|
||
<paragraph id="00098444767DA35CFFF36F3DFD46F936" blockId="72.[108,1212,1651,1734]" pageId="72">
|
||
FIGURE 42.
|
||
<taxonomicName id="C7B6FFC7767DA35CFF6F6F3BFE32F978" baseAuthorityName="GH" baseAuthorityYear="1144" box="[240,472,1651,1675]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="72" phylum="Arthropoda" rank="species" species="lipscombae">
|
||
<emphasis id="32C25856767DA35CFF6F6F3BFE32F978" box="[240,472,1651,1675]" italics="true" pageId="72">Harlanethis lipscombae</emphasis>
|
||
</taxonomicName>
|
||
(KS 52425; TEAU038).
|
||
<emphasis id="32C25856767DA35CFD4D6F3BFD00F978" bold="true" box="[722,746,1651,1675]" pageId="72">A.</emphasis>
|
||
Pedipalp, ventral.
|
||
<emphasis id="32C25856767DA35CFC366F3BFC2AF978" bold="true" box="[937,960,1651,1675]" pageId="72">B.</emphasis>
|
||
Pedipalp, ectal.
|
||
<emphasis id="32C25856767DA35CFBF76F3BFB95F978" bold="true" box="[1128,1151,1651,1675]" pageId="72">C.</emphasis>
|
||
Pedipalp, dorsal.
|
||
<emphasis id="32C25856767DA35CFF6C6FD8FEE6F95B" bold="true" box="[243,268,1680,1704]" pageId="72">D.</emphasis>
|
||
Pedipalp, ectodorsal.
|
||
<emphasis id="32C25856767DA35CFE6E6FD8FDEDF95B" bold="true" box="[497,519,1680,1704]" pageId="72">E.</emphasis>
|
||
Conductor and embolus, ventral detail.
|
||
<emphasis id="32C25856767DA35CFC306FD8FC2EF95B" bold="true" box="[943,964,1680,1704]" pageId="72">F.</emphasis>
|
||
Pedipalp CDBP, ventral detail.
|
||
<emphasis id="32C25856767DA35CFF2E6FE5FF20F936" bold="true" box="[177,202,1709,1733]" pageId="72">G.</emphasis>
|
||
Pedipalp expanded, apical. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="00098444767CA35DFEF969AEFE9EFF09" box="[358,372,230,250]" pageId="73">E</paragraph>
|
||
<caption id="54C9D4CC767CA35DFFF66FC5FEDAF92E" ID-DOI="http://doi.org/10.5281/zenodo.4613994" ID-Zenodo-Dep="4613994" httpUri="https://zenodo.org/record/4613994/files/figure.png" pageId="73" startId="73.[105,192,1677,1698]" targetBox="[170,1143,248,1645]" targetPageId="73">
|
||
<paragraph id="00098444767CA35DFFF66FC5FEDAF92E" blockId="73.[105,1216,1675,1758]" pageId="73">
|
||
FIGURE 43.
|
||
<taxonomicName id="C7B6FFC7767CA35DFF6E6FC3FDE6F950" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[241,524,1675,1699]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="73" phylum="Arthropoda" rank="species" species="weintrauborum">
|
||
<emphasis id="32C25856767CA35DFF6E6FC3FDE6F950" box="[241,524,1675,1699]" italics="true" pageId="73">Harlanethis weintrauborum</emphasis>
|
||
</taxonomicName>
|
||
(KS 0503; TEAU016).
|
||
<emphasis id="32C25856767CA35DFC9D6FC3FCF1F950" bold="true" box="[770,795,1675,1699]" pageId="73">A.</emphasis>
|
||
Pedipalp, ventral.
|
||
<emphasis id="32C25856767CA35DFC7D6FC3FC12F950" bold="true" box="[994,1016,1675,1699]" pageId="73">B.</emphasis>
|
||
Pedipalp, ectal.
|
||
<emphasis id="32C25856767CA35DFB386FC3FB55F950" bold="true" box="[1191,1215,1675,1699]" pageId="73">C.</emphasis>
|
||
Pedipalp, dorsal.
|
||
<emphasis id="32C25856767CA35DFE876FE1FED8F932" bold="true" box="[280,306,1705,1729]" pageId="73">D.</emphasis>
|
||
Pedipalp, ectodorsal.
|
||
<emphasis id="32C25856767CA35DFD8E6FE1FDCFF932" bold="true" box="[529,549,1705,1729]" pageId="73">E.</emphasis>
|
||
Pedipalp, CDBP detail.
|
||
<emphasis id="32C25856767CA35DFC836FE1FCDBF932" bold="true" box="[796,817,1705,1729]" pageId="73">F.</emphasis>
|
||
Conductor and embolus ventral detail. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="00098444767FA35EFFF369AAFE8DFEE8" blockId="74.[108,636,226,448]" pageId="74">
|
||
spins.
|
||
<taxonomicName id="C7B6FFC7767FA35EFF3069AAFECDFF09" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[175,295,226,250]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="32C25856767FA35EFF3069AAFECDFF09" box="[175,295,226,250]" italics="true" pageId="74">Harlanethis</emphasis>
|
||
</taxonomicName>
|
||
is an undeclinable proper name and feminine in gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="48ACD7CF767FA35EFF13686CFF1FFE33" pageId="74" type="distribution">
|
||
<paragraph id="00098444767FA35EFF13686CFE0CFEAE" blockId="74.[108,636,226,448]" pageId="74">
|
||
DISTRIBUTION: This genus can be found in northeastern
|
||
<collectingRegion id="C2724AA6767FA35EFF60680DFE6BFEAE" box="[255,385,325,349]" country="Australia" name="Queensland" pageId="74">Queensland</collectingRegion>
|
||
(fig. 60).
|
||
</paragraph>
|
||
<paragraph id="00098444767FA35EFF13682EFF1FFE33" blockId="74.[108,636,226,448]" pageId="74">
|
||
NATURAL HISTORY:
|
||
<taxonomicName id="C7B6FFC7767FA35EFEF6682EFE0CFE8D" authorityName="Álvarez-Padilla & Kallal & Hormiga" authorityYear="2020" box="[361,486,358,382]" class="Arachnida" family="Tetragnathidae" genus="Harlanethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="32C25856767FA35EFEF6682EFE0CFE8D" box="[361,486,358,382]" italics="true" pageId="74">Harlanethis</emphasis>
|
||
</taxonomicName>
|
||
appears to be endemic to the tropical northern zone of
|
||
<collectingRegion id="C2724AA6767FA35EFFF368E0FF1BFE33" box="[108,241,424,448]" country="Australia" name="Queensland" pageId="74">Queensland</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |