117 lines
14 KiB
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117 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682" ID-Pensoft-Pub="2747-8106-97-53" ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47" ID-ZooBank="933EC356F21C45AF9CFA563E64D27953" ModsDocID="2747-8106-97-53" checkinTime="1702462880200" checkinUser="pensoft" docAuthor="Neubauer, Thomas A. & Wesselingh, Frank P." docDate="2023" docId="880543E811DC58D2B97FE14B5150BFA6" docLanguage="en" docName="Zitteliana 97: 53-88" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115682" docTitle="Valvata sp. 1" docType="treatment" docVersion="1" id="4A3318772B185D35B1E7EC9578EF4A47" lastPageNumber="53" masterDocId="4A3318772B185D35B1E7EC9578EF4A47" masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms" masterLastPageNumber="88" masterPageNumber="53" pageNumber="53" updateTime="1702462880200" updateUser="pensoft">
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<mods:title id="DAD6FA5F954D25A146661FF6AD641CEC">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
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<mods:namePart id="6F12D31D9AF560EFCBFD36C14642ECF4">Neubauer, Thomas A.</mods:namePart>
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<mods:nameIdentifier id="22920EBD34F57DCC2E24A09401E55317" type="ORCID">https://orcid.org/0000-0002-1398-9941</mods:nameIdentifier>
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<mods:affiliation id="9DEB798029327A8B67B0B20902DAA261">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:nameIdentifier id="F3A1AA9B39947EEC496FA3A266AC6D9E" type="email">neubauer@snsb.de</mods:nameIdentifier>
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<mods:namePart id="840029C8932C083FF8804EC24B8C48FC">Wesselingh, Frank P.</mods:namePart>
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<mods:nameIdentifier id="2216DFFA54579CD335CA61E2889CEB98" type="ORCID">https://orcid.org/0000-0003-3655-0701</mods:nameIdentifier>
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<mods:affiliation id="1B6C7E6509614C79210F19B8C711CDF2">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands & Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
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<mods:title id="33297A4BBC31C902C7E98F983FD3DFCB">Zitteliana</mods:title>
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<mods:date id="C29DDBCE7FB0366C29388EA521091F5F">2023</mods:date>
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<mods:number id="C3DBC2D0E2365CC4E49E199CD62B335B">2023-12-12</mods:number>
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<mods:number id="CAAB587BC1CAE32FFCA421065A539DD8">97</mods:number>
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<mods:url id="C9404C079619E1DFFCA9555602A26369">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:url>
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<mods:classification id="5F7F002895CC05C26922F5593DCBE7AE">journal article</mods:classification>
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<mods:identifier id="25C424A90B52A0A648CDDE33BC50453B" type="DOI">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:identifier>
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<treatment id="880543E811DC58D2B97FE14B5150BFA6" LSID="urn:lsid:plazi:treatment:880543E811DC58D2B97FE14B5150BFA6" httpUri="http://treatment.plazi.org/id/880543E811DC58D2B97FE14B5150BFA6" lastPageNumber="53" pageId="0" pageNumber="53">
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<subSubSection id="618F5E19673E36DCECAC68E2FA2A4350" pageId="0" pageNumber="53" type="nomenclature">
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<paragraph id="FA9AF724FE20E33D3E2B9B87688F0C13" pageId="0" pageNumber="53">
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<taxonomicName id="015BBE444AD616B9C0CA0E867AFDB867" ID-CoL="8V67M" LSID="880543E8-11DC-58D2-B97F-E14B5150BFA6" authority="sp. 1" authorityName="sp. 1" class="Gastropoda" family="Valvatidae" genus="Valvata" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Valvata" order="Ectobranchia" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">Valvata sp. 1</taxonomicName>
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</paragraph>
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<subSubSection id="9A6E578FB6B30D20BBF0DBE4741F7368" pageId="0" pageNumber="53" type="description">
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<paragraph id="113DC3D9EEEB6F865655075D7C690405" pageId="0" pageNumber="53">
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<figureCitation id="2BB3ED4F59D3026D338D12837EC90876" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Valvatidae of the Kolankaya I fauna. A-D. Valvata piscinalis (Mueller, 1774), RGM 962694, sample 3. E-H, U, V. Valvata sp. 1, RGM 1365352, sample 1. I-L, W. Valvata sp. 1, RGM 962695, sample 2. M-P. Valvata gregaria Bukowski, 1896, lectotype (designated by Willmann 1981), IGUW 1895 XII / 25, Skiadhi Monastery, Rhodes, Greece, Istrios Formation (Pliocene). Q-T. Valvata gregaria Bukowski, 1896, paralectotype, IGUW 1895 XII / 26, same locality and stratum. Scale bars: 1 mm (A-T); 100 μm (U-W)." figureDoi="10.3897/zitteliana.97.115682.figure14" httpUri="https://binary.pensoft.net/fig/950751" pageId="0" pageNumber="53">Fig. 14E-L, U-W</figureCitation>
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</paragraph>
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<subSubSection id="528E0169CE737973F8B42CB6093105C6" pageId="0" pageNumber="53" type="materials_examined">
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<paragraph id="8B1A398BA419967365BC1DCCBA08F764" pageId="0" pageNumber="53">Material.</paragraph>
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<paragraph id="9DED23C811246350C3B7903828962C1A" pageId="0" pageNumber="53">
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Sample 1:
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<specimenCount id="B1E0A61C018DB70CBF9F956CEBD76B7E" type="generic">1 specimen</specimenCount>
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(RGM 1365352),
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<specimenCount id="02E320EEA7D971228A11360E7E9764A6" type="generic">1 specimen</specimenCount>
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(SNSB-BSPG
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<date id="C8C86D05C4BCE835FA0FD919D8B7D677" value="2023-12-21">2023 XII 21</date>
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); sample 2:
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<specimenCount id="80F4649277101A58F75C641E8146A127" type="generic">1 specimen</specimenCount>
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(RGM 962695),
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<specimenCount id="B7B6FDA83910CA60C4275C723A02FBE2" type="generic">1 specimen</specimenCount>
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(SNSB-BSPG
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<date id="2B08A1A003082E552297D145AF390E1A" value="2023-12-22">2023 XII 22</date>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="CA553564558DC0806D6C24EA23EC9ABD" pageId="0" pageNumber="53" type="description">
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<paragraph id="C3A0C22CA1841DBA73EA457291955DDA" pageId="0" pageNumber="53">Dimensions.</paragraph>
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<paragraph id="03F3E3FC1510394B96FF27B96199D0DC" pageId="0" pageNumber="53">
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1.60
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<normalizedToken id="CF3D0A007F561F1859B13F34E6FAC0BE" originalValue="×">x</normalizedToken>
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2.48 mm (RGM 962695; Fig.
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<figureCitation id="F09CEA3729175F34B10B062144DFF7DA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Valvatidae of the Kolankaya I fauna. A-D. Valvata piscinalis (Mueller, 1774), RGM 962694, sample 3. E-H, U, V. Valvata sp. 1, RGM 1365352, sample 1. I-L, W. Valvata sp. 1, RGM 962695, sample 2. M-P. Valvata gregaria Bukowski, 1896, lectotype (designated by Willmann 1981), IGUW 1895 XII / 25, Skiadhi Monastery, Rhodes, Greece, Istrios Formation (Pliocene). Q-T. Valvata gregaria Bukowski, 1896, paralectotype, IGUW 1895 XII / 26, same locality and stratum. Scale bars: 1 mm (A-T); 100 μm (U-W)." figureDoi="10.3897/zitteliana.97.115682.figure14" httpUri="https://binary.pensoft.net/fig/950751" pageId="0" pageNumber="53">14I-L, W</figureCitation>
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), 1.48
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<normalizedToken id="10B97892FD09BC2F7BF2AE8B190C3638" originalValue="×">x</normalizedToken>
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2.18 mm (RGM 1365352; Fig.
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<figureCitation id="2CED514EB582E3716AD93B67AA23019D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Valvatidae of the Kolankaya I fauna. A-D. Valvata piscinalis (Mueller, 1774), RGM 962694, sample 3. E-H, U, V. Valvata sp. 1, RGM 1365352, sample 1. I-L, W. Valvata sp. 1, RGM 962695, sample 2. M-P. Valvata gregaria Bukowski, 1896, lectotype (designated by Willmann 1981), IGUW 1895 XII / 25, Skiadhi Monastery, Rhodes, Greece, Istrios Formation (Pliocene). Q-T. Valvata gregaria Bukowski, 1896, paralectotype, IGUW 1895 XII / 26, same locality and stratum. Scale bars: 1 mm (A-T); 100 μm (U-W)." figureDoi="10.3897/zitteliana.97.115682.figure14" httpUri="https://binary.pensoft.net/fig/950751" pageId="0" pageNumber="53">14E-H, U, V</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="41FF72AAD357DD0B2991E5910528619B" pageId="0" pageNumber="53" type="remarks">
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<paragraph id="72F815750D0979D9ADEAEFB78AD735E2" pageId="0" pageNumber="53">Remarks.</paragraph>
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<paragraph id="2EA9C938AA4F9D0EC91DE6875C636B44" pageId="0" pageNumber="53">
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This species is characterized by a small, low-trochiform shell with relatively large, nearly circular aperture, slightly raised, bulbous apex, wide umbilicus, and moderately distinct growth lines. The protoconch bears the typical valvatid pattern, with numerous distinct spiral ridges crossed by weaker axial, u-shaped ridges, creating a wavy impression (Fig.
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<figureCitation id="10A67B396847DCFDE152C7C492BA5838" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Valvatidae of the Kolankaya I fauna. A-D. Valvata piscinalis (Mueller, 1774), RGM 962694, sample 3. E-H, U, V. Valvata sp. 1, RGM 1365352, sample 1. I-L, W. Valvata sp. 1, RGM 962695, sample 2. M-P. Valvata gregaria Bukowski, 1896, lectotype (designated by Willmann 1981), IGUW 1895 XII / 25, Skiadhi Monastery, Rhodes, Greece, Istrios Formation (Pliocene). Q-T. Valvata gregaria Bukowski, 1896, paralectotype, IGUW 1895 XII / 26, same locality and stratum. Scale bars: 1 mm (A-T); 100 μm (U-W)." figureDoi="10.3897/zitteliana.97.115682.figure14" httpUri="https://binary.pensoft.net/fig/950751" pageId="0" pageNumber="53">14U, V</figureCitation>
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).
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</paragraph>
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<paragraph id="CD29080F99B15EA9BC861123BAC15539" pageId="0" pageNumber="53">
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The material closely resembles the Pliocene
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<taxonomicName id="B303CC2F9D5BBC5DD07D11CF8458325E" authorityName="Bukowski" authorityYear="1896" class="Gastropoda" family="Valvatidae" genus="Valvata" kingdom="Animalia" lsidName="Valvata gregaria" order="Ectobranchia" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="gregaria">
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<emphasis id="B461A2E07E8A80132E16AF4A1A3D2FA7" italics="true" pageId="0" pageNumber="53">Valvata gregaria</emphasis>
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</taxonomicName>
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Bukowski, 1896 (p. 25-28, pl. 8, figs 7, 8) from Rhodes in terms of the shell size and shape, the relatively large aperture, and low number of whorls (see also
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<bibRefCitation id="A084A324766DE6E22211963D54942812" author="Willmann, R" journalOrPublisher="A" pageId="0" pageNumber="53" pagination="10 - 235" refId="B179" refString="Willmann, R, 1981. Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos/Aegaeis. Palaeontographica Abt. A 174: 10 - 235" title="Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos / Aegaeis. Palaeontographica Abt." volume="174" year="1981">Willmann 1981</bibRefCitation>
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, p. 77, textfig. 25). However, the lectotype and paralectotype of that species (designated by
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<bibRefCitation id="BA7A399AD13E193D05D87D0A6E06BE1E" author="Willmann, R" journalOrPublisher="A" pageId="0" pageNumber="53" pagination="10 - 235" refId="B179" refString="Willmann, R, 1981. Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos/Aegaeis. Palaeontographica Abt. A 174: 10 - 235" title="Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos / Aegaeis. Palaeontographica Abt." volume="174" year="1981">Willmann 1981</bibRefCitation>
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) show a depressed and unevenly coiled spire (Fig.
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<figureCitation id="3350A0D022F0D7B3D9495B201B7A38A0" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Valvatidae of the Kolankaya I fauna. A-D. Valvata piscinalis (Mueller, 1774), RGM 962694, sample 3. E-H, U, V. Valvata sp. 1, RGM 1365352, sample 1. I-L, W. Valvata sp. 1, RGM 962695, sample 2. M-P. Valvata gregaria Bukowski, 1896, lectotype (designated by Willmann 1981), IGUW 1895 XII / 25, Skiadhi Monastery, Rhodes, Greece, Istrios Formation (Pliocene). Q-T. Valvata gregaria Bukowski, 1896, paralectotype, IGUW 1895 XII / 26, same locality and stratum. Scale bars: 1 mm (A-T); 100 μm (U-W)." figureDoi="10.3897/zitteliana.97.115682.figure14" httpUri="https://binary.pensoft.net/fig/950751" pageId="0" pageNumber="53">14M-T</figureCitation>
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), while the Denizli specimens has a bulbous, almost pointy apex and a slightly more ovoid aperture.
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</paragraph>
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<paragraph id="41D2F28F71312BC4EB89FE5806B7AC13" pageId="0" pageNumber="53">Given the low amount of specimens available for comparison and assessing morphological variability, the perhaps not fully adult state of the specimens, as well as the low number of features, we do not describe a new species, although we could not detect any extant or late Cenozoic species that matches our material.</paragraph>
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</subSubSection>
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</treatment>
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</document> |