265 lines
44 KiB
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265 lines
44 KiB
XML
<document ID-CLB-Dataset="229647" ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-GBIF-Dataset="6ba6d2ca-5dad-4270-914f-1042327d503c" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz & Cavichioli, Rodney Ramiro" docDate="2023" docId="8163D5A361105F87B85132B47B068843" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics & amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Effossana intrinseca Domahovski & Cavichioli 2023, sp. nov." docType="treatment" docUuid="932FADA1-5DB6-46DA-8CC4-A68231D872C1" docUuidSource="ZooBank" docVersion="5" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong & Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1732669275789" updateUser="ExternalLinkService">
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<mods:title id="80AD20E586F09DCCA48BEEC684A0E5F8">Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong & Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:namePart id="74930AD751B0DE8D837CFB706DF04835">Domahovski, Alexandre Cruz</mods:namePart>
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<mods:affiliation id="6CB84770EBD957E7390CD69413B245AB">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:nameIdentifier id="C59A6BF58CC3A433DCA9233B49B7E248" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="5A2FEBD3A386B668EF7CD853B11B7EF8">Cavichioli, Rodney Ramiro</mods:namePart>
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<mods:affiliation id="660F3E91F13A698280C55E43CC7F2795">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title id="C8D2F92919CBFFD436196207BA20A0AB">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="14E8C207D11A90C98A42EEE298C943FF">2023</mods:date>
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<mods:number id="F30D39F8B3D1CCC8610A3126A472F033">2023-01-20</mods:number>
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<mods:number id="1B4782FB0240F6AEAEE1DFF8C3BB9FE2">81</mods:number>
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<treatment id="8163D5A361105F87B85132B47B068843" ID-GBIF-Taxon="209013710" LSID="urn:lsid:zoobank.org:act:932FADA1-5DB6-46DA-8CC4-A68231D872C1" httpUri="http://treatment.plazi.org/id/8163D5A361105F87B85132B47B068843" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Cicadellidae" scope_order="Hemiptera" scope_tribe="Gyponini">
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<subSubSection id="A3058022B9D50F4061A95EFBE3AB3227" pageId="0" pageNumber="79" type="nomenclature">
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<paragraph id="67ECD3A1EE6707AEA5BB7831FB0656BD" pageId="0" pageNumber="79">
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<taxonomicName id="38F21CE1D74736C5A4F24B81A2A08CD6" LSID="https://zoobank.org/932FADA1-5DB6-46DA-8CC4-A68231D872C1" authority="Domahovski & Cavichioli, 2023" authorityName="Domahovski & Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Effossana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Effossana intrinseca" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="intrinseca" status="sp. nov.">Effossana intrinseca</taxonomicName>
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<taxonomicNameLabel id="059CE43A753770C686E015FFAA8775E2" pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="3808F3A83F3AD8F528BFA403497D2A85" pageId="0" pageNumber="79" type="description">
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<paragraph id="8E371198F08849C5C20A5F7B1EAE2895" pageId="0" pageNumber="79">
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<figureCitation id="F69F033A88F0CA1E2C5DF56C072B5D83" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">Figs 39</figureCitation>
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<figureCitation id="0BBFC61715FB7240BEBECC8BF33A84A6" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">, 40</figureCitation>
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<figureCitation id="1ABE946AA790461EF315D4F88923DEA7" captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Species of Effossana gen. nov., dorsal (left) and lateral (right) views. A, B E. gloma comb. nov., male, DZUP. C, D E. intrinseca sp. nov., holotype male, DZUP. E, F E. scapa comb. nov., holotype male, BMNH. G, H E. tenuemarginata sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure43" httpUri="https://binary.pensoft.net/fig/798196" pageId="0" pageNumber="79">, 43C, D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="84368BCCC566C54C0133E47FDA26D8D9" pageId="0" pageNumber="79" type="diagnosis">
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<paragraph id="5FD118C45DF6807166F1D7B18F4DA18E" pageId="0" pageNumber="79">Diagnosis.</paragraph>
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<paragraph id="E514D1C8087232941C36DE14A0D59CB7" pageId="0" pageNumber="79">
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Head, in lateral view (Fig.
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<figureCitation id="8AFB7D49EAF2319E9B79CF231CBBD78A" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39C</figureCitation>
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), crown-face transition moderately thick Pygofer, in dorsal view (Fig.
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<figureCitation id="9C4AF5A847BC15881AE04EA61A5CA73E" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39g</figureCitation>
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), with acute subapical process, directed inward. Style, in lateral view (Fig.
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<figureCitation id="FCF65F387CA438D1F4681C0C05515615" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39K</figureCitation>
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), with blade very narrow near base and strongly widened towards apex; apex truncated with thin and acute process directed dorsally. Aedeagus (Fig.
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<figureCitation id="630ECD9BCFE9A02EE5ACE5D3BE0D215C" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39L</figureCitation>
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) with shaft very long, basal portion strongly curved forming wide ring.
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</paragraph>
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</subSubSection>
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<subSubSection id="53F80716BE19EA9C822C5F23AAD9D20D" pageId="0" pageNumber="79" type="description">
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<paragraph id="CCF88DBD9CC6263AED4A232BB830E92C" pageId="0" pageNumber="79">Measurements.</paragraph>
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<paragraph id="9C45778B4E33C9A2B34026ED73A3B02E" pageId="0" pageNumber="79">Total length: holotype male 8.6 mm; paratypes, males (n = 4) 8.6-9.2 mm, females (n = 2) 9.3-9.4 mm.</paragraph>
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</subSubSection>
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<subSubSection id="163EE7BD72309C9762CF1773E0C17CE4" pageId="0" pageNumber="79" type="coloration">
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<paragraph id="005E2D14F8D10E445B7207B163A61FBE" pageId="0" pageNumber="79">Coloration.</paragraph>
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<paragraph id="421DA1D03BBD526B1E483E7F2135C278" pageId="0" pageNumber="79">
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Head and thorax yellowish brown. Crown (Fig.
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<figureCitation id="29D76B84CB39A233520ACA7681247465" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39A</figureCitation>
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) with coronal suture black. Face (Fig.
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<figureCitation id="DA5D220CEC2D288494B6BE9C124540F8" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39B</figureCitation>
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) gena yellow, brown markings near subgenal suture, ventral and inner margins of eye. Pronotum (Fig.
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<figureCitation id="C3D536615CCB750427D3A01BEDD7F446" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39A</figureCitation>
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) with brown punctures, pair of elongated and oblique black maculae behind ocelli, near anterior margin; posterior margin with black transverse band; proepimeron (Fig.
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<figureCitation id="E101AAFF5599CC457E679A8EDAB3A096" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39C</figureCitation>
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), with black band below lateral carina. Mesonotum (Fig.
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<figureCitation id="D0275110E4F64EAAB60C857DDD9D8F40" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39A</figureCitation>
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) with yellow markings, pair of black ring-shaped maculae near lateral angles and pair of rounded black spots, near scutoscutellar suture. Scutellum (Fig.
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<figureCitation id="C51446AEB36EBB000EBBA328981F866A" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39A</figureCitation>
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) black with pair of large yellow spots. Forewing (Fig.
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<figureCitation id="C00BE3D4F76D84F3873B45B9D3A7346D" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39D</figureCitation>
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) with dark-brown mottling; membrane translucent light yellow; veins brown outlined by dark brown. Metatibia (Fig.
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<figureCitation id="6B9B9CDCDF2312C3DE493572DAD620E7" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39C</figureCitation>
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) with cucullate bases of setae blacks.
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</paragraph>
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</subSubSection>
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<subSubSection id="8195A2A3FFA938EC5AE5C77DCF32BACD" pageId="0" pageNumber="79" type="description">
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<paragraph id="9DB531AA68BC759DE8631264FDAFF9FE" pageId="0" pageNumber="79">Description.</paragraph>
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<paragraph id="F2B592D2D4A4013F4523926A50C09229" pageId="0" pageNumber="79">
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Head, in dorsal view (Fig.
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<figureCitation id="6CCEC98A0C0984553D0ADD3662E3E403" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39A</figureCitation>
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), transocular width 8.1 tenths of humeral width of pronotum. Head, in frontal view (Fig.
|
||
<figureCitation id="6AEACC8DFDE126862B2CB99560D4B12B" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39B</figureCitation>
|
||
), frons 1.4
|
||
<normalizedToken id="C68A5540171A7C596E5E99B781992A90" originalValue="×">x</normalizedToken>
|
||
longer than wide; clypeus 1.4
|
||
<normalizedToken id="4060B4833CBC6C1815792DFB23B3B48D" originalValue="×">x</normalizedToken>
|
||
longer than wide; lateral margins parallel. Head, in lateral view (Fig.
|
||
<figureCitation id="5E06EF843BADB596483B0D45DEEA1634" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39C</figureCitation>
|
||
), crown-face transition moderately thick, smooth medially and with few striae near of eye. Forewing (Fig.
|
||
<figureCitation id="4C8FD52C82B3D6919DAC980C013575A6" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39D</figureCitation>
|
||
) with appendix reduced, narrower than width of first apical cell. Protibia with AD row with 2-4 differentiated setae. Metatibia PD, AD and AV rows with 23-24, 12-13 and 16-18 macrosetae respectively. Metatarsomere I moderately elongated, about 3
|
||
<normalizedToken id="F4C10206C13DAAE4AC7937F9258B1BC2" originalValue="×">x</normalizedToken>
|
||
longer than apical width; inner row of ventral surface with 5-6 setae; pecten with 6-7 platellae. Metatarsomere II pecten with 3 platellae. Other characteristics as in generic description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DAFDAD4BAEC8E8CCFA36088DA67CE03" pageId="0" pageNumber="79" type="male terminalia">
|
||
<paragraph id="CED66F555CCDF39C52150C80B1B26C15" pageId="0" pageNumber="79">Male terminalia.</paragraph>
|
||
<paragraph id="3BFC30D67F3581FC07BD1BA2F4060A0E" pageId="0" pageNumber="79">
|
||
Sternite VIII (Fig.
|
||
<figureCitation id="7160FDF8E70D647FBBC420F6F16D334E" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39E</figureCitation>
|
||
) 1.4
|
||
<normalizedToken id="66D6E184EDE2FE9C44B3EDD1FD8DDE2D" originalValue="×">x</normalizedToken>
|
||
wider than long, lateral margins rounded; posterior margin with median third deeply excavated, bearing short median projection. Valve (Fig.
|
||
<figureCitation id="F3C660495278FAC56148A4782B6D33C9" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39F</figureCitation>
|
||
) 2.5
|
||
<normalizedToken id="2A2FBD07006D1EF6B9075C7A3D167816" originalValue="×">x</normalizedToken>
|
||
wider than long; posterior margin narrowly excavated; in lateral view, strongly convex. Pygofer, in lateral view (Fig.
|
||
<figureCitation id="CA78D716E1940ACE86124B92E5A9F169" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39G</figureCitation>
|
||
), 2.2
|
||
<normalizedToken id="30F3CEBD87117F2D7902916595895E36" originalValue="×">x</normalizedToken>
|
||
longer than maximum height; anteroventral margin sinuous; posterodorsal and posteroventral margins slightly rounded; apex narrowed and rounded; in dorsal view (Fig.
|
||
<figureCitation id="BC8292E9EF3856E1F0304470DE2CCCE1" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39g</figureCitation>
|
||
), with acute subapical process, directed inward. Subgenital plate, in ventral view (Fig.
|
||
<figureCitation id="A0FB2368E41FC3EFF048B5748F190ED7" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39H</figureCitation>
|
||
), elongated, 3.4
|
||
<normalizedToken id="B5099E07703AD5B8690AAA069F852DB6" originalValue="×">x</normalizedToken>
|
||
longer than wide, maximum width near half length; inner margin straight; outer margin broadly rounded; apex rounded. Connective (Fig.
|
||
<figureCitation id="C85072558F6ED7D28CCD1F301FAD754D" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39I</figureCitation>
|
||
) with thin arms. Style, in lateral view (Fig.
|
||
<figureCitation id="82D927D357A01A234C7434AD36BB2CB5" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39K</figureCitation>
|
||
), with blade very narrow near base and strongly widened towards apex; dorsal margin excavated; ventral margin straight, with apical third serrated; apex truncated and very wide, with thin and acute process directed dorsally. Aedeagus (Fig.
|
||
<figureCitation id="4F383D51271ECD83108DAAFD352EDF1A" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39L, M</figureCitation>
|
||
) dorsal apodeme reduced, dorsal margin excavated, lateral margins not projected laterally; apodemal process elongated and curved dorsally, dorsal margin slightly excavated medially, apex slightly broadened and rounded; shaft cylindrical and very long, basal portion strongly curved forming wide ring, portion after the ring straight; apex slightly twisted laterally, with two pair of processes directed dorsally: apical pair very short, near lateral margins and subapical pair 4
|
||
<normalizedToken id="98DC3A6CEDB3FF4F7E6FE37EAAE317AC" originalValue="×">x</normalizedToken>
|
||
longer than the apical pair, arising from the posterior surface. Other characteristics as in generic description.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="494315AB50248A842225E640FB288B4A" pageId="0" pageNumber="79" type="female terminalia">
|
||
<paragraph id="0881EA5AC3C080A34F0637AD3EBD84E5" pageId="0" pageNumber="79">Female terminalia.</paragraph>
|
||
<paragraph id="3B74BED7DFF484F5B60C9540A77D55C0" pageId="0" pageNumber="79">
|
||
Sternite VII (Fig.
|
||
<figureCitation id="F3355487C962A170AC5F51265637380B" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">40A</figureCitation>
|
||
) 2
|
||
<normalizedToken id="32DB7838EB129594F88111D94263BD42" originalValue="×">x</normalizedToken>
|
||
wider than long; posterolateral angles produced as far as posterior margin; posterior margin deeply excavated each side of rounded median lobe which occupies more than median third. Internal sternite VIII membranous. Pygofer (Fig.
|
||
<figureCitation id="C129CABEBCFC8B08F77B5BFC3CB3880E" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">40A, B</figureCitation>
|
||
) 1.9
|
||
<normalizedToken id="14E653EA3F66751A5A119BA4BA5688AD" originalValue="×">x</normalizedToken>
|
||
longer than maximum height; apex rounded. First valvifer (Fig.
|
||
<figureCitation id="7EC4AFCB06983FF4722C0A7CAD48DB8E" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">40C</figureCitation>
|
||
) sub-rectangular 1.6
|
||
<normalizedToken id="87266AC8F28479A190FF33C615980EA2" originalValue="×">x</normalizedToken>
|
||
wider than long; dorsal margin straight; posterior margin rounded. First valvula (Fig.
|
||
<figureCitation id="D5B2490D517B12D814D12C3FD68A6414" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">40C</figureCitation>
|
||
) 4
|
||
<normalizedToken id="1C2A2839E686EAC60BB010EDC89F2AF8" originalValue="×">x</normalizedToken>
|
||
longer than wide. Second valvula (Fig.
|
||
<figureCitation id="64CCC17FD82692B943A37E95737BE0B0" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">40E</figureCitation>
|
||
) 4
|
||
<normalizedToken id="FA25E0B6F6476E81D69D4EED66905A8B" originalValue="×">x</normalizedToken>
|
||
longer than wide; dorsal protuberance rounded. Gonoplac (Fig.
|
||
<figureCitation id="ACB1BF4E90F512D637C02BAAD4F3AAF2" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Effossana intrinseca sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure40" httpUri="https://binary.pensoft.net/fig/798193" pageId="0" pageNumber="79">40G</figureCitation>
|
||
) 3.7
|
||
<normalizedToken id="33BB946E21F8DA213E497BE110804BAE" originalValue="×">x</normalizedToken>
|
||
longer than wide. Other characteristics as in the diagnosis of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C5029A48D57410F1523B13718D3B5A39" pageId="0" pageNumber="79" type="materials_examined">
|
||
<paragraph id="39730E572A605B892A7F111A72F80F89" pageId="0" pageNumber="79">Material examined.</paragraph>
|
||
<paragraph id="5449CE49BEC4A51333AE4CF964ECBB7F" pageId="0" pageNumber="79">
|
||
<materialsCitation id="BF89D81D70F92CDAD508F03370A73993" ID-GBIF-Occurrence="4017211314" collectingDate="1985-03-27" collectorName="C. Ellias" country="Brazil" location="Caceres" specimenCount="1" stateProvince="Mato Grosso" typeStatus="Holotype">
|
||
<emphasis id="89F6CC8CF0A62444E6DEF02934339738" bold="true" pageId="0" pageNumber="79">
|
||
<typeStatus id="C45726EF5C825A6EF82F1E6973189715">Holotype</typeStatus>
|
||
male:
|
||
<collectingCountry id="D7154472801ECE4296BB3B3029A74FA8" name="Brazil">Brazil</collectingCountry>
|
||
:
|
||
<collectingRegion id="1D2D858DF735AAAAEC37FDB4EF745391" country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<normalizedToken id="D0B67D03B310E5B5DC5310049362BDB2" originalValue="“Cáceres">
|
||
"
|
||
<location id="35BE7C6DD7F7543BF046BB43682956E9" LSID="urn:lsid:plazi:treatment:8163D5A361105F87B85132B47B068843:35BE7C6DD7F7543BF046BB43682956E9" country="Brazil" name="Caceres" stateProvince="Mato Grosso">Caceres</location>
|
||
</normalizedToken>
|
||
, MT\
|
||
<collectingDate id="C51D4D9641278F9FBAC6A313668E864A" value="1985-03-27">27/III/1985</collectingDate>
|
||
\
|
||
<collectorName id="E8FB159C47572857A9FBFC3FCF01F01E">C. Ellias</collectorName>
|
||
leg.\ Polonoroeste", "Dpt° Zool\
|
||
<normalizedToken id="C1B5384ED5C8FB931DD698ED1AED96AB" originalValue="UF-Paraná”">UF-Parana"</normalizedToken>
|
||
(DZUP)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="C17AA14BD2A51ED9D71E9EEE863EBEEF" ID-GBIF-Occurrence="4017211336" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="C. Ellias" country="Brazil" location="Paratypes" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Paratypes" typeStatus="Paratypes">
|
||
<emphasis id="AFDF68B4A630FCE67A593C655EF89814" bold="true" pageId="0" pageNumber="79">
|
||
<collectingRegion id="63B9E9AA0DD2D510E70C83322954165C" country="Brazil" name="Paratypes">
|
||
<typeStatus id="F2EFFE0F0389D920F795D4465429A0BF">Paratypes</typeStatus>
|
||
</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="2CAB63147D3D4F150AB4E111E99C9101" type="male">2♂</specimenCount>
|
||
,
|
||
<specimenCount id="167248CA6875946BCD5AF9BC3F8E2B0D" type="female">1♀</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="630E7F5211F5C6CE1437BF2B7566A84C" ID-GBIF-Occurrence="4017211353" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="C. Ellias" country="Brazil" location="Caceres" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
|
||
<specimenCount id="3FC1666C3CE3DDAB5B1FB0E5259D19DC" type="male">1♂</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
except
|
||
<date id="950F1C3F65D6E74A17DC784C0B531536" value="1984-12-20">20.XII.1984</date>
|
||
(DZUP);
|
||
<materialsCitation id="4C06004DD0EEDD17098A9D6201F71EF8" ID-GBIF-Occurrence="4017211411" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="C. Ellias" country="Brazil" location="Caceres" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
|
||
<specimenCount id="64EA9E3D9A6090EB8A3B86C530F127A1" type="female">1♀</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
except
|
||
<date id="46A752D1B44ED5754BC0947F88BF5004" value="1985-01-14">14.I.1985</date>
|
||
(DZUP).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="268BF1EE92596FD1503A1FBED1C3DDCA" pageId="0" pageNumber="79" type="etymology">
|
||
<paragraph id="25B917A58C633C08710CD93FB9EAF931" pageId="0" pageNumber="79">Etymology.</paragraph>
|
||
<paragraph id="E726F30BF9BB2E0C995A28B08FC2719B" pageId="0" pageNumber="79">the new species name refers to the process of pygofer directed inward.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82FE4D53EA1F95E49B33D0EAE3891DD2" pageId="0" pageNumber="79" type="remarks">
|
||
<paragraph id="EF45A7B6EDFD3F75172EE9862446BCE8" pageId="0" pageNumber="79">Remarks.</paragraph>
|
||
<paragraph id="B9A7539EDF83941DDA6699CE8BF7BA86" pageId="0" pageNumber="79">
|
||
<taxonomicName id="6E438EE102160E6F5A61D70BFFD40DF6" authorityName="Domahovski & Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Effossana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Effossana intrinseca" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="intrinseca">
|
||
<emphasis id="C1889A99CECBBFB781715C4198E23160" italics="true" pageId="0" pageNumber="79">Effossana intrinseca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8D80C717F424BB05B540822C94BD3E58" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
|
||
can be distinguish from
|
||
<taxonomicName id="F72C1FEBADEDB93EF872C8176556EA15" authorityName="Domahovski & Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="E. circumnota" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="circumnota">
|
||
<emphasis id="D416BEACA3846E247AC94D0020C9BA1F" italics="true" pageId="0" pageNumber="79">E. circumnota</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D8A103B12E806368DE00CBEFA094A551" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
|
||
by the pygofer (Fig.
|
||
<figureCitation id="03D623B027A8890690FF49A3BC6842FF" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39g</figureCitation>
|
||
) with a subapical process directed inward; the style (Fig.
|
||
<figureCitation id="893FE5CD4062538C10E712206C099E36" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Effossana intrinseca sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure39" httpUri="https://binary.pensoft.net/fig/798192" pageId="0" pageNumber="79">39K</figureCitation>
|
||
) with ventral margin of blade without process, and the apex truncated and very wide, forming a thin and acute process directed dorsally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |