203 lines
27 KiB
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203 lines
27 KiB
XML
<document id="245B65D32F00A91090EC94A3778CA7AE" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="7BD8AF6479FDD0D8C2BB4DCA38A84082" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus septicola Guignot 1956" docType="treatment" docVersion="5" lastPageNumber="136" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="135" updateTime="1701314941532" updateUser="plazi">
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<mods:titleInfo id="B5CE92A5E00E7F302DD18D71C87C4350">
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<mods:title id="656763B96E97177A0A0CED6374613CDF">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="045B1D15E26C9CE9040F1C8D7C61327E">Bistroem, Olof</mods:namePart>
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<mods:namePart id="47652096B32F8470155218D920D13787">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="FA96E473E45D11D74348B6187F3EFA3D">Bergsten, Johannes</mods:namePart>
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<mods:title id="98CE744EDF256544B70D671B7A19B329">ZooKeys</mods:title>
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<mods:date id="3F17B95BC9918850CC315E1C06193BD9">2015</mods:date>
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<mods:number id="8BF0B277BAE926998EBA0994C75BEC12">542</mods:number>
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<mods:classification id="E9BB83EDDDC5B4239656710921398EDB">journal article</mods:classification>
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<mods:identifier id="16500C2BBA00235548D4084F6DE0DB72" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="7BD8AF6479FDD0D8C2BB4DCA38A84082" ID-GBIF-Taxon="127862741" LSID="urn:lsid:plazi:treatment:7BD8AF6479FDD0D8C2BB4DCA38A84082" httpUri="http://treatment.plazi.org/id/7BD8AF6479FDD0D8C2BB4DCA38A84082" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135">
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<subSubSection id="94971A9A1E16A590847D6AB2BD0638B4" pageId="134" pageNumber="135" type="multiple">
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<paragraph id="7EAC282BB64F8505E22BC8BD8455F30A" pageId="134" pageNumber="135">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="F6041C0199D8D6E94FDC9EDACB6BF993" pageId="134" pageNumber="135" type="nomenclature">
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<paragraph id="FED7A09CCC3FE82DF311CDAA4008D00E" pageId="134" pageNumber="135">
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<taxonomicName id="CC823B1E660CAE8191CB1E76DBA884F9" ID-CoL="8LZVT" authority="Guignot, 1956" authorityName="Guignot" authorityYear="1956" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola Guignot, 1956</taxonomicName>
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Figs 101-102, 289-290, 444, 557
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</paragraph>
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</subSubSection>
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<subSubSection id="B571EA0CB68D49F0E7B5830CD59FB72D" pageId="134" pageNumber="135" type="reference_group">
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<paragraph id="5AE696646D2A99AB8CCC3280DD713DC1" pageId="134" pageNumber="135">
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<taxonomicName id="05E4F81C110978BFBC6E8BB2151C12E9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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<bibRefCitation id="4479858CE3AC7A16E1B22325F7E48E1C" author="Guignot, F" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="258" pageNumber="259" pagination="211 - 223" title="Coleopteres hydrocanthares. Le Parc National du Niokolo-Koba. 13." volume="48" year="1956 b">Guignot 1956b</bibRefCitation>
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: 220, 221 (original description, faunistics);
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<bibRefCitation id="F24E482BAAB2B0E53506684B3C306A9B" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="48 - 63" title="Dytiscidae (Coleoptera) of Nyasaland and Southern Rhodesia. X. Cybisterini and addenda." volume="29" year="1967">Omer-Cooper 1967</bibRefCitation>
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: 60 (discussion,
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<taxonomicName id="EF50F3C3C5068B69E8216896911AAC50" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pullatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="pullatus">Laccophilus pullatus</taxonomicName>
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Omer-Cooper
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<taxonomicName id="FF682C9566BD951490EE614FBAB8CCB7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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Guignot,
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<taxonomicName id="2A4EF033E49BF3ACC6F0FF43982BE118" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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Guignot and
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<taxonomicName id="D92A45C28775AB36802C83FB61FB5F1F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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Gschwendtner are synonyms);
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<bibRefCitation id="9E5034F1962A686040EA829E064A29CD" author="Medler, JT" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="259" pageNumber="260" pagination="1 - 919" title="Insects of Nigeria - Check list and bibliography." volume="30" year="1980">Medler 1980</bibRefCitation>
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: 155 (faunistics, list);
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<bibRefCitation id="1330E18BF98FFCFF340EAA346B6B5233" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 250 (catalogue, faunistics, list, synonymy,
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<taxonomicName id="BA88BD4F8678C46B5755FE870E0428AE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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Gschwendtner);
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<bibRefCitation id="B2F7BD58FBF105C0414B08591365D79A" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 213 (catalogue, faunistics, list, synonymy,
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<taxonomicName id="0C0A88BA3A5FC8BAC163A102124F9BD3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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Gschwendtner). Restored species.
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</paragraph>
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<paragraph id="A0CE867AAD0312213171BC8E01180DA2" pageId="134" pageNumber="135">
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<taxonomicName id="6FCE6D9DAAE054DEAF51D9B9A82AB2EA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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<bibRefCitation id="4C8EB4649DC86C2808F994C21034A819" author="Guignot, F" journalOrPublisher="Resultats Scientifiques de l'Exploration Hydrobiologique des Lacs Kivu, Edouard et Albert (1952 - 1954)" pageId="258" pageNumber="259" pagination="133 - 169" title="Haliplidae et Dytiscidae (Coleoptera Hydrocanthares)." volume="3" year="1959 d">Guignot 1959d</bibRefCitation>
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: 163 (original description, faunistics, discussion);
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<bibRefCitation id="312D0D6A7766AEC490365642E06843F7" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="48 - 63" title="Dytiscidae (Coleoptera) of Nyasaland and Southern Rhodesia. X. Cybisterini and addenda." volume="29" year="1967">Omer-Cooper 1967</bibRefCitation>
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: 60 (discussion,
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<taxonomicName id="2DBFE7672399980566F5C68A3C08D8FC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pullatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="pullatus">Laccophilus pullatus</taxonomicName>
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O-mer-Cooper,
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<taxonomicName id="8617154B3605E44603255794F66B470C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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Guignot.,
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<taxonomicName id="6E38E6B122F1F38102369F7D5B6B049F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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Guignot and
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<taxonomicName id="6BA07EB8793568D2FF7F0F30D43BF1CE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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Gschwendtner are synonyms);
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<bibRefCitation id="77B93DF9F4A83A31EBA7AE1CE1C9D013" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 246 (catalogue, faunistics, list, synonymy,
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<taxonomicName id="F89982B6C1632555F82B1CB635CE92D4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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Gschwendtner;
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<bibRefCitation id="6029F5A34FEFC35AAC2C35D7A14BA17A" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 213 (catalogus, list, synonymy,
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<taxonomicName id="9A658824F350D8125040CF7C59A8E7D1" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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Gschwendtner). Confirmed synonymy.
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</paragraph>
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</subSubSection>
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<subSubSection id="958624779625A79F06FD18351DC101E8" pageId="134" pageNumber="135" type="type localities">
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<paragraph id="0E4A2B5DA642FC7FA57C911D94F13326" pageId="134" pageNumber="135">Type localities.</paragraph>
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<paragraph id="348C087CEEFFD5DC0F8CE9C7C18FDAD1" pageId="134" pageNumber="135">
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<taxonomicName id="89C683FFDF12026590F9D1259EED16A3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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: Senegal: Niokolo-Koba, Badi.
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</paragraph>
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<paragraph id="F74F142AEEB328621067FADFA6B1DC7E" pageId="134" pageNumber="135">
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<taxonomicName id="C2B18E42D86D614D5DB87DB31006E95D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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: Zaire: Lake Albert, Mwita.
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</paragraph>
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</subSubSection>
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<subSubSection id="EB6B03D519B96F890BCD6EA5F2EA5D86" pageId="134" pageNumber="135" type="type material">
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<paragraph id="52030DC189F1737816C2BD31778269CD" pageId="134" pageNumber="135">Type material studied</paragraph>
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<paragraph id="C64DF41955B2C6054F258E09765D331D" pageId="134" pageNumber="135">
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(8 exs.).
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<taxonomicName id="5BB25DA47D7971C803147067D52A3AE2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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: Holotype: male; "Mission IFAN au Parc National du Niokolo Koba Badi (
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<normalizedToken id="F1B450CC00791748C24D488A16768A3D" originalValue="Sénégal">Senegal</normalizedToken>
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) 15.VIII.-25.IX-1955 / Type / F. Guignot det. 1954 / F. Guignot det.1954
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<taxonomicName id="8CF99A6252D3A653C49644C8418E9ADB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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sp. n. Type (MNHN). - Paratypes: "Mission IFAN au Parc National du Niokolo Koba Badi (
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<normalizedToken id="F466432C7F203AEA0DE0D78F333B2170" originalValue="Sénégal">Senegal</normalizedToken>
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) 15.VIII.-25.IX-1955 / Paratype / Dr. F. Guignot det.
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<taxonomicName id="0E15CAF7D2CE0995E6EEC75B15283BDE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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Guign." (4 exs. IRSNB); same but "Ouassadou 12.VIII. 55" (1 ex. IRSNB).
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</paragraph>
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<paragraph id="28E3842A23F4F813005015A6EF31F4C1" pageId="134" pageNumber="135">
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<taxonomicName id="E4C69E9809E0ACAE41040105AFE2D216" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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: Holotype: male: "Congo Belge. Lac Albert: Mwita (
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<normalizedToken id="FF550F85843789263B29264A858887E2" originalValue="prét">pret</normalizedToken>
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foret galerie) U.V. 22.XII. 1953 J. Verbeke - KEA 4083 / Type / F. Guignot det., 1956
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<taxonomicName id="888D23757DFB8755A7124248BAE0F249" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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sp. n. Type male" (IRSNB). - Paratype: "Congo Belge Lac Albert: Bezaha U.V. 19.12. 1953, 4070 / female symbol / Paratype" (1 ex. MNHN).
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</paragraph>
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</subSubSection>
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<subSubSection id="56ABF48605D8BE34BBE84639DD9A5B6F" pageId="134" pageNumber="135" type="materials_examined">
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<paragraph id="8321B539A00D22ABBC797F95009A6028" pageId="134" pageNumber="135">Additional material studied</paragraph>
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<paragraph id="A6F6ABD375CC0D7B2BBC598B6F082D31" pageId="134" pageNumber="135">(29 exs.): Sudan: "Upper Nile Malakal 5-20.1. 1963 Linnavuori" (22 exs. MZH; habitus in Fig. 444); same data and "ad lucem" (2 exs., MZH); "Sudan / Linnavuori" (1 ex. MZH); "L. Shambe 21.1. 1954 Omer-Cooper" (2 exs. AMGS). - Nigeria: "Marsk, Katsina-Dawura rd. 6.10.1963 JOC." (1 ex. AMGS). - Zaire: "Lac Albert Kasenyi UV 21.VI. 1953 4012" (1 ex. IRSNB).</paragraph>
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</subSubSection>
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<subSubSection id="B79BE4175B3E16C1AEA29AA07E0120B0" pageId="134" pageNumber="135" type="comments on synonymy">
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<paragraph id="B2BF300F21912A1B5B900CA22DC4C011" pageId="134" pageNumber="135">Comments on synonymy.</paragraph>
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<paragraph id="CAFD5A4F8CCFFE58BF37FC151061D699" pageId="134" pageNumber="135">
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Earlier synonymy of
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<taxonomicName id="60259FE4BA951C1A0B82C1DF67AC8D5F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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and
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<taxonomicName id="26C9387C523E72B80DAD68DC5F77C21D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alberticus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="alberticus">Laccophilus alberticus</taxonomicName>
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is verified by examination of their holotypes.
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<taxonomicName id="119558357DACC0F843E42B0CC5502D4A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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, being the older name is the valid name of the species. Furthermore, examination revealed that
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<taxonomicName id="6B7510BC8B2383713736AB8CE33B15CF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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is not synonymous with
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<taxonomicName id="EFF966350B37642A029A0A4FC889AF3F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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and
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<taxonomicName id="5C85FEEAE8F4EB950A88FC60D0466E41" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pullatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="pullatus">Laccophilus pullatus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6009E9AC9C4A26C437F0E3731D5945B" pageId="134" pageNumber="135" type="diagnosis">
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<paragraph id="3DA710D879B2F84F86EB74130C10280A" pageId="134" pageNumber="135">Diagnosis.</paragraph>
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<paragraph id="2D53AE6E10EC428054598539BCD39924" pageId="134" pageNumber="135">
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<taxonomicName id="D1F154507400FB5AB682D7233AC42A08" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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is characterized by pale ferrugineous to ferrugineous elytra in combination with peculiarly shaped penis (extreme apex slightly extended). Morphologically, in case of penis and body shape,
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<taxonomicName id="F21281D43B3F9B7A328724CA10D7DE24" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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resembles much of
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<taxonomicName id="815623B0B14FD8A859E165B91B3F81B0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pullatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="pullatus">Laccophilus pullatus</taxonomicName>
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and
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<taxonomicName id="C16300C61AE684AC7A8FDA09720A16E5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luteosignatus" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="luteosignatus">Laccophilus luteosignatus</taxonomicName>
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. These two species have, however, blackish to dark ferrugineous elytra, which separate them clearly from
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<taxonomicName id="7781A87F670C5C31E092511CF2CAAA9E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus septicola" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="septicola">Laccophilus septicola</taxonomicName>
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, with its much paler main colour of elytra.
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</paragraph>
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</subSubSection>
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<subSubSection id="E7FC6689A831B09691287B68D9696F13" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135" type="description">
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<paragraph id="C97BDE81D2ECF4BB19542E658A26F9C8" pageId="134" pageNumber="135">Description.</paragraph>
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<paragraph id="CB3684710E80824FE12B5D9D0D543CD9" pageId="134" pageNumber="135">Body length 2.9-3.3 mm, width 1.6-1.8 mm. Dorsal, aspect of body almost without colour pattern (Fig. 444). Rarely humeral regions may be slightly paler forming minor pale spots.</paragraph>
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<paragraph id="AEFC93284DFA23919E01175CFC36CEEA" pageId="134" pageNumber="135">Head: Pale ferrugineous. Submat, finely reticulated. Reticulation indistinctly double. Large meshes indistinct; only in part discernible and slightly more strongly developed than small meshes. Almost impunctate; fine punctures discernible at eyes.</paragraph>
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<paragraph id="BC76AA0C48A81BFC3F9EDF7D801923E6" pageId="134" pageNumber="135">Pronotum: Pale ferrugineous, mediobasally sometimes slightly darker, but lacks well delimited darkened area. Submat, finely microsculptured. Reticulation double; large meshes contain two to five small meshes. Small meshes sometimes weakly developed and hardly discernible. Punctation sparse, very fine, in part absent; discernible at frontal margin and laterally.</paragraph>
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<paragraph id="0D1EA3DB3BBC04FDB55EDC6CB5BFA995" pageId="134" pageNumber="135">Elytra: Pale ferrugineous to ferrugineous. In comparsion with head and pronotum, elytra extensively slightly darker (Fig. 444). Rarely with somewhat vague, paler humeral spots. Submat, finely microsculptured. Double reticulation only in part discernible (at basal region). Punctation very fine and irregularly distributed; large areas lack punctures. Irregular, somewhat sparse punctures form a rather indistinct, discal row. Lateral, pre-apical furrow fine, finely pubescent.</paragraph>
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<paragraph id="05CDE7249EC58BA47F6980520A246FA1" pageId="135" pageNumber="136">
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<pageBreakToken id="8ADA057427FE344338FD23838D962014" pageId="135" pageNumber="136" start="start">Ventral</pageBreakToken>
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aspect: Pale ferrugineous. Submat, very finely and partly indistinctly microsculptured. Basal ventrirtes with fine, curved striae. Almost impunctate. Metacoxal lines-process-area not distinctly modified. Metacoxal plates in anterior half with fine, in part rather indistinct, transversely located furrows. In posterior half furrows almost absent. Prosternal process slightly extended, rather slender, pointed. Apical ventrirte asymmetric; with a sublateral, sharp knob (Fig. 101).
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</paragraph>
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<paragraph id="BBCABE35BD6E890288C945107A1D4D46" pageId="135" pageNumber="136">Legs: Protarsus slender, claws moderately extensive and slightly curved. Pro- and mesotarsus with suckers.</paragraph>
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<paragraph id="43A0BA422FDB586C2463669D6A3C5A40" pageId="135" pageNumber="136">Male genitalia: Penis in lateral aspect medially bent and from there penis narrows towards extreme apex, which is extended to a small tip (Figs 289-290).</paragraph>
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<paragraph id="8AB19532DC95E8EAD3C9B85EFD038292" pageId="135" pageNumber="136">Female: Apical ventrirte simple (Fig. 102). Pro- and mesotarsus almost as in male but lack suckers.</paragraph>
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</subSubSection>
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<subSubSection id="5719095193124635A1FF0A51E476C6A2" pageId="135" pageNumber="136" type="distribution">
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<paragraph id="A8BA53535B5E0DBAA7BBB7A39B47C168" pageId="135" pageNumber="136">Distribution.</paragraph>
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<paragraph id="DDD922B5FA625EF61D2581563EE7ACA4" pageId="135" pageNumber="136">Senegal, Sudan, Nigeria, Zaire (Fig. 557). Only verified records accepted in map because of different earlier taxonomic opinion regarding species delimitations.</paragraph>
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</subSubSection>
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<subSubSection id="6D47ABDC50F9D4BF9FB28D0D4AE51883" pageId="135" pageNumber="136" type="collecting circumstances">
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<paragraph id="C31FB0D7F2E2AC8CD907FD1432A91BB9" pageId="135" pageNumber="136">Collecting circumstances.</paragraph>
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<paragraph id="0D1E95EA927D80A2F3FED69B167D0116" pageId="135" pageNumber="136">Not documented. In Sudan sampled at light collection.</paragraph>
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</subSubSection>
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</treatment>
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</document> |