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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.418.7629" ID-PMC="PMC4104466" ID-Pensoft-Pub="1313-2970-418-1" ID-Pensoft-UUID="F034FFA3FFAFFFBEB578C42DD65AFFA2" ID-PubMed="25061358" ID-Zenodo-Dep="578347" ID-ZooBank="D71C8A3DA6CA40A5B3A034A1FD1C16A0" ModsDocID="1313-2970-418-1" checkinTime="1451245725588" checkinUser="pensoft" docAuthor="Skvarla, Michael J., Fisher, J. Ray &amp; Dowling, Ashley P. G." docDate="2014" docId="5ECE2AAC03E555B65956D1FD82D8C5BF" docLanguage="en" docName="ZooKeys 418: 1-103" docOrigin="ZooKeys 418" docPubDate="2014-06-20" docSource="http://dx.doi.org/10.3897/zookeys.418.7629" docTitle="Lupaeus Castro &amp; Den Heyer 2009" docType="treatment" docVersion="5" id="F034FFA3FFAFFFBEB578C42DD65AFFA2" lastPageNumber="26" masterDocId="F034FFA3FFAFFFBEB578C42DD65AFFA2" masterDocTitle="A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records" masterLastPageNumber="103" masterPageNumber="1" pageNumber="25" updateTime="1668158708043" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skvarla, Michael J.</mods:namePart>
<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, J. Ray</mods:namePart>
<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
<mods:affiliation>Department of Entomology, 319 AGRI Building, University of Arkansas, Fayetteville, Arkansas, 72701, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>418</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>103</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.418.7629</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.418.7629</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-418-1</mods:identifier>
<mods:identifier type="ZooBank">D71C8A3DA6CA40A5B3A034A1FD1C16A0</mods:identifier>
<mods:identifier type="Pensoft-UUID">F034FFA3FFAFFFBEB578C42DD65AFFA2</mods:identifier>
<mods:identifier type="Zenodo-Dep">578347</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152053720" LSID="urn:lsid:plazi:treatment:5ECE2AAC03E555B65956D1FD82D8C5BF" httpUri="http://treatment.plazi.org/id/5ECE2AAC03E555B65956D1FD82D8C5BF" lastPageId="26" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection pageId="24" pageNumber="25" type="nomenclature">
<paragraph pageId="24" pageNumber="25">
<taxonomicName LSID="5ECE2AAC-03E5-55B6-5956-D1FD82D8C5BF" authority="Castro &amp; Den Heyer, 2009" authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lupaeus" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Lupaeus Castro &amp; Den Heyer, 2009</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="historical review">
<paragraph pageId="24" pageNumber="25">Historical review.</paragraph>
<paragraph pageId="24" pageNumber="25">
<bibRefCitation author="Berlese, A" journalOrPublisher="Redia" pageId="83" pageNumber="84" pagination="125 - 177" refId="B40" refString="Berlese, A, 1916. Centuria secunda di Acari nuovi. Redia 12 (1): 125 - 177" title="Centuria secunda di Acari nuovi." volume="12" year="1916">Berlese (1916)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis italics="true" pageId="24" pageNumber="25">Eupalus subterraneus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Thor, S" editor="Thor, S" journalOrPublisher="Walter De Gruyter and Co, Leipzig" pageId="94" pageNumber="95" pagination="164 - 175" refId="B246" refString="Thor, S, Willmann, C, 1941. Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. In: Thor, S, Willmann, C, Eds., Das Tierreich, Eine Zusammenstellung und Kennzeichung der rezenten Tierformen. Walter De Gruyter and Co, Leipzig: 164 - 175" title="Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae." volumeTitle="Das Tierreich, Eine Zusammenstellung und Kennzeichung der rezenten Tierformen." year="1941">Thor and Willmann (1941)</bibRefCitation>
redescribed
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis italics="true" pageId="24" pageNumber="25">Eupalus subterraneus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Baker, EW" journalOrPublisher="Anales de la Escuela Nacional de Ciencias Biologicas Mexico" pageId="82" pageNumber="83" pagination="229 - 273" refId="B11" refString="Baker, EW, Hoffmann, A, 1948. Acaros de la familia Cunaxidae. Anales de la Escuela Nacional de Ciencias Biologicas Mexico 5 (3-4): 229 - 273" title="Acaros de la familia Cunaxidae." volume="5" year="1948">Baker and Hoffmann (1948)</bibRefCitation>
erected
<taxonomicName authorityName="Baker &amp; Hoffmann" authorityYear="1948" class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Cunaxoides</emphasis>
</taxonomicName>
in place of
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Eupalus</emphasis>
</taxonomicName>
as
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Eupalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Eupalus</emphasis>
</taxonomicName>
was preoccupied; they also described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minutus">
<emphasis italics="true" pageId="24" pageNumber="25">Cunaxoides minutus</emphasis>
</taxonomicName>
and redescribed and illustrated
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis italics="true" pageId="24" pageNumber="25">Cunaxoides subterraneus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Den Heyer, J" journalOrPublisher="Acarologia" pageId="86" pageNumber="87" pagination="18 - 31" refId="B80" refString="Den Heyer, J, 1979b. Pulaeus, a new cunaxid genus (Prostigmata: Acari). Acarologia 21 (1): 18 - 31" title="Pulaeus, a new cunaxid genus (Prostigmata: Acari)." volume="21" year="1979 b">Den Heyer (1979b)</bibRefCitation>
erected
<taxonomicName authorityName="Den Heyer" authorityYear="1978" class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus</emphasis>
</taxonomicName>
, moving those species with
<emphasis italics="true" pageId="24" pageNumber="25">f2</emphasis>
present and setae present on basifemora IV to the new genus from
<taxonomicName authorityName="Baker &amp; Hoffmann" authorityYear="1948" class="Arachnida" family="Cunaxidae" genus="Cunaxoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Cunaxoides</emphasis>
</taxonomicName>
; he also described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus martini</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="clarae">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus clarae</emphasis>
</taxonomicName>
and placed
<taxonomicName authorityName="Den Heyer" authorityYear="1978" class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus</emphasis>
</taxonomicName>
into the subfamily
<taxonomicName authorityName="Den Heyer" authorityYear="1978" lsidName="" pageId="24" pageNumber="25" rank="subFamily" subFamily="Cunaxoidinae">Cunaxoidinae</taxonomicName>
.
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platygnathus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus platygnathus</emphasis>
</taxonomicName>
was described by
<bibRefCitation author="Bu, G" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="84" pageNumber="85" pagination="70 - 73" refId="B44" refString="Bu, G, Li, L, 1991. A new species of the genus Pulaeus from China (Acari: Cunaxidae). Acta Zootaxonomica Sinica 16 (1): 70 - 73" title="A new species of the genus Pulaeus from China (Acari: Cunaxidae)." volume="16" year="1991">Bu and Li (1991)</bibRefCitation>
.
<bibRefCitation author="Corpuz-Raros, LA" journalOrPublisher="Philippine Entomologist" pageId="85" pageNumber="86" pagination="119 - 138" refId="B59" refString="Corpuz-Raros, LA, 1996b. Philippine predatory mites of the family Cunaxidae (Acari). Genus Pulaeus Den Heyer with records of two species from Central Kalimantan, Borneo and Java, Indonesia. Philippine Entomologist 10 (2): 119 - 138" title="Philippine predatory mites of the family Cunaxidae (Acari). Genus Pulaeus Den Heyer with records of two species from Central Kalimantan, Borneo and Java, Indonesia." volume="10" year="1996 b">Corpuz-Raros (1996b)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus dentatus</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lenis">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus lenis</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="longisetus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus longisetus</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="villacarlosae">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus villacarlosae</emphasis>
</taxonomicName>
, and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="filipinus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus filipinus</emphasis>
</taxonomicName>
from the Philippines.
<bibRefCitation author="Hu, SJ" journalOrPublisher="Journal of Ninbo Teachers College" pageId="88" pageNumber="89" pagination="56 - 59" refId="B144" refString="Hu, SJ, 1997. Cunaxid mites recorded in China. Journal of Ninbo Teachers College 15 (1): 56 - 59" title="Cunaxid mites recorded in China." volume="15" year="1997">Hu (1997)</bibRefCitation>
reported
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platygnathus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus platygnathus</emphasis>
</taxonomicName>
from China.
<bibRefCitation author="Lin, LZ" journalOrPublisher="Fujian Sciences and Technology Press, Fuzhou" pageId="90" pageNumber="91" pagination="109 - 121" refId="B176" refString="Lin, LZ, Zhang, YZ, 2000. Fauna of Insects in Fuijan Province of China. Fujian Sciences and Technology Press, Fuzhou: 109 - 121" title="Fauna of Insects in Fuijan Province of China" year="2000">Lin and Zhang (2000)</bibRefCitation>
reported
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platygnathus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus platygnathus</emphasis>
</taxonomicName>
from China. Lin and Zhang (2003) reported
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minutus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus minutus</emphasis>
</taxonomicName>
from China.
<bibRefCitation author="Corpuz-Raros, LA" journalOrPublisher="Asia Life Science" pageId="85" pageNumber="86" pagination="153 - 173" refId="B63" refString="Corpuz-Raros, LA, 2007. Additional species of Bonziinae and Cunaxoidinae and description of the male of Coleoscirus horidula (Tseng) (Coleoscirinae) from the Philippines (Cunaxidae, Acari). Asia Life Science 16 (2): 153 - 173" title="Additional species of Bonziinae and Cunaxoidinae and description of the male of Coleoscirus horidula (Tseng) (Coleoscirinae) from the Philippines (Cunaxidae, Acari)." volume="16" year="2007">Corpuz-Raros (2007)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="polilloensis">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus polilloensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus philippinensis</emphasis>
</taxonomicName>
from the Philippines. Castro and
<bibRefCitation author="Den Heyer, J" journalOrPublisher="Arthropod fauna of the UAE" pageId="86" pageNumber="87" pagination="17 - 25" refId="B92" refString="Den Heyer, J, 2009. . Arthropod fauna of the UAE Vol. 2: 17 - 25" volume="Vol. 2" year="2009">Den Heyer (2009)</bibRefCitation>
erected
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus</emphasis>
</taxonomicName>
and moved into it those species of
<taxonomicName authorityName="Den Heyer" authorityYear="1978" class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Pulaeus</emphasis>
</taxonomicName>
that possess two pointed processes on the pedipalp tibiotarsus and 1 simple seta on basifemora IV; they also described
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lectus">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus lectus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lobidorsalis">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus lobidorsalis</emphasis>
</taxonomicName>
and provided a key to the Brazilian and South African species.
<bibRefCitation author="Sergeyenko, AL" journalOrPublisher="Journal of Entomological Research" pageId="93" pageNumber="94" pagination="239 - 248" refId="B231" refString="Sergeyenko, AL, 2011b. Mites of the genera Pulaeus and Lupaeus (Acari: Prostigmata: Cunaxidae) of Crimea, Ukraine. Zootaxa 2088: 54-68.Shakhsi Zare F, Arbabi M, Kmali H, Ghasemzadeh M (2012) Study on faunestic, distribution and host range of Tetranychoidea (Arachnida: Acari) on fruit trees in Mashhad region, Iran. Journal of Entomological Research 4 (3): 239 - 248" title="Mites of the genera Pulaeus and Lupaeus (Acari: Prostigmata: Cunaxidae) of Crimea, Ukraine. Zootaxa 2088: 54 - 68. Shakhsi Zare F, Arbabi M, Kmali H, Ghasemzadeh M (2012) Study on faunestic, distribution and host range of Tetranychoidea (Arachnida: Acari) on fruit trees in Mashhad region, Iran." volume="4" year="2011 b">Sergeyenko (2011b)</bibRefCitation>
described
<taxonomicName authorityName="Sergeyenko" authorityYear="2011" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="valentinae">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus valentinae</emphasis>
</taxonomicName>
.
<bibRefCitation author="Den Heyer, J" journalOrPublisher="Journal of Natural History" pageId="86" pageNumber="87" pagination="2049 - 2070" refId="B106" refString="Den Heyer, J, Ueckermann, EA, Khanjani, M, 2013. Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae. Journal of Natural History 47 (31-32): 2049 - 2070" title="Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae." volume="47" year="2013">Den Heyer et al. (2013)</bibRefCitation>
described
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="iranensis">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus iranensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sativae">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus sativae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
<paragraph pageId="24" pageNumber="25">
<emphasis italics="true" pageId="24" pageNumber="25">Gnathosoma</emphasis>
.
<emphasis bold="true" pageId="24" pageNumber="25">Pedipalps</emphasis>
3-segmented. Femurogenua at least twice as long as wide, complemented with 6 setae. Tibiotarsi at least twice as long as wide, usually complemented with 6 setae; they possess 2 or 3 pointed processes and may possess a bladder- or knob-like apophysis (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="24" pageNumber="25">Fig. 24a</figureCitation>
).
<emphasis bold="true" pageId="24" pageNumber="25">Subcapitulum</emphasis>
with 6 pairs of setae (
<emphasis italics="true" pageId="24" pageNumber="25">hg1</emphasis>
-
<emphasis italics="true" pageId="24" pageNumber="25">4</emphasis>
and 2 pairs of adoral setae); setae
<emphasis italics="true" pageId="24" pageNumber="25">hg4</emphasis>
often the longest.
<emphasis bold="true" pageId="24" pageNumber="25">Chelicera</emphasis>
with seta present.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/29884" pageId="24" pageNumber="25" start="Figures 24–27" startId="F10">
<paragraph pageId="24" pageNumber="25">
<emphasis bold="true" pageId="24" pageNumber="25">Figures 24-27.</emphasis>
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus</emphasis>
</taxonomicName>
illustrations.
<emphasis bold="true" pageId="24" pageNumber="25">24a</emphasis>
Pedipalp tibiotarsus
<emphasis bold="true" pageId="24" pageNumber="25">24b</emphasis>
Genital setae not in a row,
<emphasis italics="true" pageId="24" pageNumber="25">g3</emphasis>
out of line
<emphasis bold="true" pageId="24" pageNumber="25">25-27</emphasis>
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus</emphasis>
</taxonomicName>
key illustrations. Setae and cupules removed from figures
<emphasis bold="true" pageId="24" pageNumber="25">25a, b</emphasis>
to increase clairity
<emphasis bold="true" pageId="24" pageNumber="25">25a</emphasis>
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="longisetus">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus longisetus</emphasis>
</taxonomicName>
, dorsal
<emphasis bold="true" pageId="24" pageNumber="25">25b</emphasis>
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="2007" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="polilloensis">
<emphasis italics="true" pageId="24" pageNumber="25">Lupaeus polilloensis</emphasis>
</taxonomicName>
, dorsal
<emphasis bold="true" pageId="24" pageNumber="25">26a</emphasis>
Ventral, small platelet present
<emphasis bold="true" pageId="24" pageNumber="25">26b</emphasis>
Ventral, small platelet absent
<emphasis bold="true" pageId="24" pageNumber="25">27a</emphasis>
Setae
<emphasis italics="true" pageId="24" pageNumber="25">f1</emphasis>
,
<emphasis italics="true" pageId="24" pageNumber="25">f2</emphasis>
born on small platelets
<emphasis bold="true" pageId="24" pageNumber="25">27b</emphasis>
Setae
<emphasis italics="true" pageId="24" pageNumber="25">f1</emphasis>
,
<emphasis italics="true" pageId="24" pageNumber="25">f2</emphasis>
born on integument.
</paragraph>
</caption>
<paragraph pageId="24" pageNumber="25">
<emphasis italics="true" pageId="24" pageNumber="25">Idiosoma, dorsal</emphasis>
. Proterosoma bears a well-sclerotized shield complemented with 2 pairs of setae (
<emphasis italics="true" pageId="24" pageNumber="25">lps</emphasis>
and
<emphasis italics="true" pageId="24" pageNumber="25">mps</emphasis>
) and 2 pairs of setose sensillae (
<emphasis italics="true" pageId="24" pageNumber="25">at</emphasis>
and
<emphasis italics="true" pageId="24" pageNumber="25">pt</emphasis>
). Dorsal hysterosoma bears a sclerotized plate that is variable in size and fused with the proterosomal shield; it may be complemented with a variable number of setae depending on the size of the plate. Setae
<emphasis italics="true" pageId="24" pageNumber="25">c1</emphasis>
-
<emphasis italics="true" pageId="24" pageNumber="25">h1</emphasis>
,
<emphasis italics="true" pageId="24" pageNumber="25">c2</emphasis>
,
<emphasis italics="true" pageId="24" pageNumber="25">f2</emphasis>
, and
<emphasis italics="true" pageId="24" pageNumber="25">h2</emphasis>
present. Cupule
<emphasis italics="true" pageId="24" pageNumber="25">im</emphasis>
present laterad and posterior of
<emphasis italics="true" pageId="24" pageNumber="25">e1</emphasis>
. Integument not covered in shields or plates is striated.
</paragraph>
<paragraph pageId="25" pageNumber="26">
<emphasis italics="true" pageId="25" pageNumber="26">
<pageBreakToken pageId="25" pageNumber="26" start="start">Idiosoma</pageBreakToken>
, ventral
</emphasis>
.
<emphasis bold="true" pageId="25" pageNumber="26">Coxae</emphasis>
sclerotized and well-defined. Coxae I-II may be fused and may coalesce medially to form a sternal shield. Coxae III-IV may be fused. Each coxa complemented with 2-4 setae. Genital plates each bear 4 setae (
<emphasis italics="true" pageId="25" pageNumber="26">g1</emphasis>
-
<emphasis italics="true" pageId="25" pageNumber="26">4</emphasis>
). Setae
<emphasis italics="true" pageId="25" pageNumber="26">g1,2,4</emphasis>
usually occur in a straight line near the midline and setae
<emphasis italics="true" pageId="25" pageNumber="26">g3</emphasis>
occur near the edge of the genital plates (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="25" pageNumber="26">Fig. 24b</figureCitation>
). 2 pairs of genital papillae visible underneath the plates. Anal plates bear 1 pair of setae; 1 pair of setae present ventrally on the integument near the anal plates. Cupule
<emphasis italics="true" pageId="25" pageNumber="26">ih</emphasis>
present ventrally laterad; the integumental setae associated with the anal plates. Integument not covered in shields or plates striated.
<emphasis bold="true" pageId="25" pageNumber="26">Legs.</emphasis>
Tarsi never constricted apically so as to end in lobes. Trichobothrium on leg tibia IV present. Basifemora setal formula 4-6-3-1. Depression of the famulus occurs on distal half of tarsus I. Tibiae I-II possess striated blunt solenidia. Ambulacral claws rippled and occur on either side of a 4-rayed empodium.
</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="key to adult female lupaeus">
<paragraph pageId="25" pageNumber="26">
Key to adult female
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="25" pageNumber="26">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="longisetus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus longisetus</emphasis>
</taxonomicName>
is known only from the male and is not included in the key. It can be recognized by the following characters: small platelet between the edges of a divided sternal shield absent, basifemora I with 3 sts, and setae
<emphasis italics="true" pageId="25" pageNumber="26">e1</emphasis>
elongate and barbed (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="25" pageNumber="26">Fig. 25a</figureCitation>
).
</paragraph>
<paragraph pageId="25" pageNumber="26">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="2007" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="polilloensis">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus polilloensis</emphasis>
</taxonomicName>
is only known from the male and is not included in the key. It can be recoginized by the following characters: small platelet between the edges of a divided sternal shield absent; basifemora I-II setal formula 4-6; platelets complemented with setae
<emphasis italics="true" pageId="25" pageNumber="26">f1</emphasis>
,
<emphasis italics="true" pageId="25" pageNumber="26">f2</emphasis>
with fused medially into one plate; and the dorsal shield densely granulate (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="25" pageNumber="26">Fig. 25b</figureCitation>
).
</paragraph>
<paragraph pageId="25" pageNumber="26">
As suggested by
<bibRefCitation author="Den Heyer, J" editor="Roskov, Y" journalOrPublisher="Reading, UK" pageId="86" pageNumber="87" publicationUrl="10.1080/01647954.2010.495953" refId="B94" refString="Den Heyer, J, 2011b. BdelloideaBase: Bdellid &amp; Cunaxid Databases (version Sep 2011). In: Roskov, Y, Kunze, T, Paglinawan, L, Orrell, T, Nicolson, D, Culham, A, Bailly, N, Kirk, P, Bourgoin, T, Baillargeon, G, Hernandez, F, De Wever, A, Eds., Species 2000 &amp; ITIS Catalogue of Life, 2013 Annual Checklist. Reading, UK, DOI: 10.1080/01647954.2010.495953" title="BdelloideaBase: Bdellid &amp; Cunaxid Databases (version Sep 2011)." url="10.1080/01647954.2010.495953" volumeTitle="Species 2000 &amp; ITIS Catalogue of Life, 2013 Annual Checklist." year="2011 b">Den Heyer (2011b)</bibRefCitation>
the following species are moved to
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus</emphasis>
</taxonomicName>
from
<taxonomicName authorityName="Den Heyer" authorityYear="1978" class="Arachnida" family="Cunaxidae" genus="Pulaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="25" pageNumber="26">Pulaeus</emphasis>
</taxonomicName>
:
<taxonomicName baseAuthorityName="Baker &amp; Hoffmann" baseAuthorityYear="1948" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="minutus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus minutus</emphasis>
</taxonomicName>
(Baker and Hoffmann) and
<taxonomicName baseAuthorityName="Berlese" baseAuthorityYear="1916" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus subterraneus</emphasis>
</taxonomicName>
(Berlese).
</paragraph>
<paragraph lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<table inLine="true" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">1</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
Small platelet ventromedially between edges of divided sternal plate present (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="25" pageNumber="26">Fig. 26a</figureCitation>
); South Africa, Brazil
</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
<taxonomicName baseAuthorityName="Den Heyer" baseAuthorityYear="1981" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="martini">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus martini</emphasis>
</taxonomicName>
(Den Heyer, 1979)
</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
Small platelet ventromedially between edges of divided sternal plate absent (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="25" pageNumber="26">Fig. 26b</figureCitation>
)
</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">2</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">2 (1)</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Basifemora I with 4 sts</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">3</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Basifemora I with 5 sts; Philippines</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="filipinus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus filipinus</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">3 (2)</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Basifemora II with 4 sts; USA</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
<taxonomicName baseAuthorityName="Baker &amp; Hoffmann" baseAuthorityYear="1948" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="minutus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus minutus</emphasis>
</taxonomicName>
(Baker &amp; Hoffmann, 1948)
</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Basifemora II with 5 sts</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">4</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Basifemora II with 6 sts</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">7</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">4 (3)</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
Setae
<emphasis italics="true" pageId="25" pageNumber="26">f1</emphasis>
shorter than
<emphasis italics="true" pageId="25" pageNumber="26">c1</emphasis>
; Philippines
</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lenis">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus lenis</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
Setae
<emphasis italics="true" pageId="25" pageNumber="26">f1</emphasis>
the same length as
<emphasis italics="true" pageId="25" pageNumber="26">c1</emphasis>
</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lectus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus lectus</emphasis>
</taxonomicName>
Castro &amp; Den Heyer, 2009
</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
Setae
<emphasis italics="true" pageId="25" pageNumber="26">f1</emphasis>
longer than
<emphasis italics="true" pageId="25" pageNumber="26">c1</emphasis>
, usually by at least 1.5 times
</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">5</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">5 (4)</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Genua I with 9 total simple setae and solenidia; Philippines</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis italics="true" pageId="25" pageNumber="26">Lupaeus dentatus</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="25" pageNumber="26">
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">-</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">Genua I with 7 total simple setae and solenidia</td>
<td colspan="1" pageId="25" pageNumber="26" rowspan="1">6</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<pageBreakToken pageId="26" pageNumber="27" start="start">6</pageBreakToken>
(5)
</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
Setae
<emphasis italics="true" pageId="26" pageNumber="27">c1</emphasis>
-
<emphasis italics="true" pageId="26" pageNumber="27">e1</emphasis>
equal in length; Brazil
</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName authorityName="Castro &amp; Den Heyer" authorityYear="2009" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lobidorsalis">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus lobidorsalis</emphasis>
</taxonomicName>
Castro &amp; Den Heyer, 2009
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
Setae
<emphasis italics="true" pageId="26" pageNumber="27">e1</emphasis>
one-fourth longer than
<emphasis italics="true" pageId="26" pageNumber="27">c1</emphasis>
,
<emphasis italics="true" pageId="26" pageNumber="27">d1</emphasis>
; Italy, USA
</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName baseAuthorityName="Berlese" baseAuthorityYear="1916" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus subterraneus</emphasis>
</taxonomicName>
(Berlese, 1916)
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">7 (3)</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
Setae
<emphasis italics="true" pageId="26" pageNumber="27">f1</emphasis>
,
<emphasis italics="true" pageId="26" pageNumber="27">f2</emphasis>
on platelets, which may be separate or fused medially (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="26" pageNumber="27">Fig. 27a</figureCitation>
)
</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">8</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
Setae
<emphasis italics="true" pageId="26" pageNumber="27">f1</emphasis>
,
<emphasis italics="true" pageId="26" pageNumber="27">f2</emphasis>
on integeument (
<figureCitation captionStart="Figures 24–27" captionStartId="F10" captionText="Figures 24 - 27. Lupaeus illustrations. 24 a Pedipalp tibiotarsus 24 b Genital setae not in a row, g 3 out of line 25 - 27 Lupaeus key illustrations. Setae and cupules removed from figures 25 a, b to increase clairity 25 a Lupaeus longisetus, dorsal 25 b Lupaeus polilloensis, dorsal 26 a Ventral, small platelet present 26 b Ventral, small platelet absent 27 a Setae f 1, f 2 born on small platelets 27 b Setae f 1, f 2 born on integument." httpUri="https://binary.pensoft.net/fig/29884" pageId="26" pageNumber="27">Fig. 27b</figureCitation>
)
</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">11</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">8 (7)</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Tibia II with 1 s, 5 sts</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">9</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Tibia II with 2 s (1 asl, 1 bsl), 5 sts; Ukraine</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName authorityName="Sergeyenko" authorityYear="2011" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="valentinae">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus valentinae</emphasis>
</taxonomicName>
Sergeyenko, 2011
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">9 (8)</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Pedipalp tibiotarsus with 4 sts; Philippines</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName baseAuthorityName="Corpuz-Raros" baseAuthorityYear="1996" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="villacarlosae">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus villacarlosae</emphasis>
</taxonomicName>
(Corpuz-Raros, 1996)
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Pedipalp tibiotarsus with 5 sts</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">10</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">10 (9)</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Tarsus I with 3 asl, 2 terminal solenidion, 1 fam, 20 or 21 sts; tarsus IV with 14 sts</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="iranensis">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus iranensis</emphasis>
</taxonomicName>
Den Heyer, 2013
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Tarsus I with 3 asl, 1 dorsodistal solenidion, 1 terminal solenidion, 1 fam, 22 sts; tarsus IV with 16 sts</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sativae">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus sativae</emphasis>
</taxonomicName>
Den Heyer, 2013
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">11 (7)</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Cheliceral seta not as long as width of cheliceral digit; China</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName baseAuthorityName="Bu &amp; Li" baseAuthorityYear="1991" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platygnathus">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus platygnathus</emphasis>
</taxonomicName>
(Bu &amp; Li, 1991)
</td>
</tr>
<tr pageId="26" pageNumber="27">
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">-</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">Cheliceral seta longer than width of cheliceral digit; South Africa, Brazil</td>
<td colspan="1" pageId="26" pageNumber="27" rowspan="1">
<taxonomicName baseAuthorityName="Den Heyer" baseAuthorityYear="1981" class="Arachnida" family="Cunaxidae" genus="Lupaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Trombidiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="clarae">
<emphasis italics="true" pageId="26" pageNumber="27">Lupaeus clarae</emphasis>
</taxonomicName>
(Den Heyer, 1979)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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