treatments-xml/data/5B/15/B3/5B15B336501E5758861C7B093B37C528.xml

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<document ID-CLB-Dataset="24349" ID-DOI="http://dx.doi.org/10.3897/zookeys.902.38967" ID-GBIF-Dataset="dfd124b1-4a06-4a50-9d92-7a12f8f8b361" ID-PMC="PMC6971156" ID-Pensoft-Pub="1313-2970-902-61" ID-Pensoft-UUID="5D3BBF83D2435220A5E5F8E8C9ED791D" ID-PubMed="31988598" ID-ZooBank="2DB048800FB848BFAAF35EC2176F8D6E" ModsDocID="1313-2970-902-61" checkinTime="1578966145580" checkinUser="pensoft" docAuthor="Chang, Wan-Jin &amp; Li, Shuqiang" docDate="2020" docId="5B15B336501E5758861C7B093B37C528" docLanguage="en" docName="ZooKeys 902: 61-105" docOrigin="ZooKeys 902" docSource="http://dx.doi.org/10.3897/zookeys.902.38967" docTitle="Psiloderces gawanaensis Li &amp; Chang 2020, sp. nov." docType="treatment" docUuid="15281939-437A-4A72-B5E3-ABF4BA47B4A7" docUuidSource="ZooBank" docVersion="7" id="5D3BBF83D2435220A5E5F8E8C9ED791D" lastPageNumber="61" masterDocId="5D3BBF83D2435220A5E5F8E8C9ED791D" masterDocTitle="Fourteen new species of the spider genus Psiloderces Simon, 1892 from Southeast Asia (Araneae, Psilodercidae)" masterLastPageNumber="105" masterPageNumber="61" pageNumber="61" updateTime="1732937456495" updateUser="ExternalLinkService">
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<mods:title id="8196E081A94FF382F07E8395EEC040DF">Fourteen new species of the spider genus Psiloderces Simon, 1892 from Southeast Asia (Araneae, Psilodercidae)</mods:title>
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<mods:namePart id="715263377DF5FE0F3ADD897AFDC6E837">Chang, Wan-Jin</mods:namePart>
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<mods:namePart id="572941219E08B596B7C80605D868BF06">Li, Shuqiang</mods:namePart>
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<mods:date id="34D8B17213E5E374F055A935A2E22948">2020</mods:date>
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<mods:identifier id="53B3519B12CE1995690C7DC0CA186A8E" type="DOI">http://dx.doi.org/10.3897/zookeys.902.38967</mods:identifier>
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<treatment id="5B15B336501E5758861C7B093B37C528" ID-GBIF-Taxon="161677827" LSID="urn:lsid:zoobank.org:act:15281939-437A-4A72-B5E3-ABF4BA47B4A7" httpUri="http://treatment.plazi.org/id/5B15B336501E5758861C7B093B37C528" lastPageNumber="61" pageId="0" pageNumber="61" scope_family="Psilodercidae" scope_order="Araneae">
<subSubSection id="8597CBF11646C7C9579D1D91D8930643" pageId="0" pageNumber="61" type="nomenclature">
<paragraph id="A3C19135608B615395D81A7CE8DA6599" pageId="0" pageNumber="61">
<taxonomicName id="ACA848DBF1E2EC2E9B5241B4BFAB1F45" ID-CoL="98ZWP" LSID="5B15B336-501E-5758-861C-7B093B37C528" authority="Li &amp; Chang" authorityName="Li &amp; Chang" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Psiloderces" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psiloderces gawanaensis" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="gawanaensis" status="sp. nov.">Psiloderces gawanaensis Li &amp; Chang</taxonomicName>
<taxonomicNameLabel id="C554D9835CCACA4BDC02B012D3DEBC6D" pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
<figureCitation id="C2BFF6F9A47E5FBB9B986DEC888BD6B5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">Figs 3</figureCitation>
<figureCitation id="17B4891FF79B8ACDEE0B3D032D14DE84" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Psiloderces gawanaensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BT = bristle, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.902.38967.figure4" httpUri="https://binary.pensoft.net/fig/370848" pageId="0" pageNumber="61">, 4</figureCitation>
<figureCitation id="E0F4C562A39CF9B7EA5B1576C7281959" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Cheliceral retromargin, posterior view. A Psiloderces bangkiraiensis sp. nov. B P. wangou sp. nov. C P. heise sp. nov. D P. cuyapoensis sp. nov. E P. bontocensis sp. nov. F P. gawanaensis sp. nov. G P. xichang sp. nov. H P. cattienensis sp. nov. I P. pingguo sp. nov. Abbreviations: CL = cheliceral laminar, RT = retromargin teeth." figureDoi="10.3897/zookeys.902.38967.figure28" httpUri="https://binary.pensoft.net/fig/370872" pageId="0" pageNumber="61">, 28F</figureCitation>
<figureCitation id="C4131BABC6B40B5C2B7C8471792264A2" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Distribution of new Psiloderces species in Southeast Asia. 1 P. heise sp. nov. 2 P. gawanaensis sp. nov. 3 P. bontocensis sp. nov. 4 P. cuyapoensis sp. nov. 5 P. xichang sp. nov. 6 P. cattienensis sp. nov. 7 P. pingguo sp. nov. 8 P. palopoensis sp. nov. 9 P. penajamensis sp. nov. 10 P. grohotensis sp. nov. 11 P. bangkiraiensis sp. nov. 12 P. bolang sp. nov. 13 P. wangou sp. nov. 14 P. malinoensis sp. nov." figureDoi="10.3897/zookeys.902.38967.figure30" httpUri="https://binary.pensoft.net/fig/370874" pageId="0" pageNumber="61">, 30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="13ED8CB31E34EC6C3C1DF3839152066D" pageId="0" pageNumber="61" type="types">
<paragraph id="EE817CB1F8041D7CE568586886233445" pageId="0" pageNumber="61">Types.</paragraph>
<paragraph id="87EE57F015B48232CEA02289C4B408F0" pageId="0" pageNumber="61">
<emphasis id="CC9BB83BD00134F4F5BAE8B57BEC311A" bold="true" italics="true" pageId="0" pageNumber="61">Holotype:</emphasis>
♂ (IZCAS), Philippines, Luzon Island, Mountain Province, Bontoc Town, road to Gawana Town,
<geoCoordinate id="9B499E25EFCFEA8B99ED1E22D1EEACA5" degrees="17" direction="north" minutes="3.6396" orientation="latitude" precision="1" value="17.06066">17°3.6396'N</geoCoordinate>
,
<geoCoordinate id="E99DE2ECE029B4AEEF7B081496BA06D4" degrees="121" direction="east" minutes="3.0402" orientation="longitude" precision="1" value="121.05067">121°3.0402'E</geoCoordinate>
, 1674 m, 26.V.2015, F. Ballarin &amp; Y. Li.
<emphasis id="E4FDEA6E8CE6F7F1F980C0A747497E3B" bold="true" italics="true" pageId="0" pageNumber="61">Paratype</emphasis>
: 1♀ (IZCAS), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="51FBDC7FA0853F02BC6BBD07B13D808F" pageId="0" pageNumber="61" type="etymology">
<paragraph id="43A12119C85E2CF437FF63BDBC128DAE" pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph id="6F5FD667CDB07D6B009206741A6E844B" pageId="0" pageNumber="61">The species name is an adjective referring to the type locality.</paragraph>
</subSubSection>
<subSubSection id="C7A94710F727E82BC97D4A25C534D4AA" pageId="0" pageNumber="61" type="diagnosis">
<paragraph id="ED577CDDF74C6C2FB6E6CE57F370BE9E" pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph id="B6B9A9EB528DB74A76ADBFEC83D3F0CB" pageId="0" pageNumber="61">
Males of
<taxonomicName id="E4E7FF5CE79120476EAC9D511292FE00" class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. gawanaensis" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="gawanaensis">
<emphasis id="486CC87DE9469650C046A1DB389495C2" italics="true" pageId="0" pageNumber="61">P. gawanaensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from congeners by the human heart-shaped bulb (inverted pyriform) (Fig.
<figureCitation id="EC99EA3CEB6ED5F2DB0C858D440687DA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Psiloderces gawanaensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BT = bristle, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.902.38967.figure4" httpUri="https://binary.pensoft.net/fig/370848" pageId="0" pageNumber="61">4B</figureCitation>
) bearing a threadlike embolus, the presence of 4 distinct bristles on the cymbial protrusion, and the clypeal projection bearing a few setae (Fig.
<figureCitation id="36E01F53D7A4A43AEFB80CD5949C87AA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Psiloderces gawanaensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BT = bristle, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.902.38967.figure4" httpUri="https://binary.pensoft.net/fig/370848" pageId="0" pageNumber="61">4C, D</figureCitation>
) (vs. absence or less than 4 bristles on cymbial protrusion); females can be differentiated from congeners by the widely spaced, horseshoe-shaped spermathecae (Fig.
<figureCitation id="384816A33FE56BA937123C992AAD35F1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">3A</figureCitation>
).
</paragraph>
<caption id="8D7800FCD94A0AB07175DB7960D9C271" doi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph id="74DF16941AF0C52129282A0B4F74E5FE" pageId="0" pageNumber="61">
<emphasis id="B77CB436B2D9C784A85E8ED8C6F6B1E7" bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
<taxonomicName id="5A1038358C1C288F3F44A5771267C5A5" authorityName="Li &amp; Chang" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Psiloderces" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psiloderces gawanaensis" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="gawanaensis">
<emphasis id="09C5C5AEFDF11ACE182D4146049E5634" italics="true" pageId="0" pageNumber="61">Psiloderces gawanaensis</emphasis>
</taxonomicName>
sp. nov., male holotype and female paratype.
<emphasis id="913366BFDC39006C9BD1675FE9647A4E" bold="true" pageId="0" pageNumber="61">A</emphasis>
Endogyne, dorsal view
<emphasis id="1F642A00878D16D70E7085A555B55CD4" bold="true" pageId="0" pageNumber="61">B</emphasis>
female epigastric area, ventral view
<emphasis id="D03D99E49B6E23B7E9E12BC6AA97F90A" bold="true" pageId="0" pageNumber="61">C</emphasis>
male habitus, lateral view
<emphasis id="1DF0054249A33348E908E4861F84FE3D" bold="true" pageId="0" pageNumber="61">D</emphasis>
male habitus, dorsal view
<emphasis id="76CBBA6EB8B32FC1DC869392E2A2E630" bold="true" pageId="0" pageNumber="61">E</emphasis>
female habitus, dorsal view
<emphasis id="3B87D66A781060082F102074C1C32434" bold="true" pageId="0" pageNumber="61">F</emphasis>
female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca.
</paragraph>
</caption>
<caption id="F10EA2BC397732E9E04B403FEAF07ED6" doi="10.3897/zookeys.902.38967.figure4" httpUri="https://binary.pensoft.net/fig/370848" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph id="C6F22D4EBC3828B81ADADA83A027D058" pageId="0" pageNumber="61">
<emphasis id="FDCA8A73BC735F6A7049E9D6CA762AF8" bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
<taxonomicName id="B6FEC8F3965F261964A72C25934431CF" authorityName="Li &amp; Chang" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Psiloderces" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psiloderces gawanaensis" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="gawanaensis">
<emphasis id="EDF1D268AFCB585426CFD3D2A2F5FBFD" italics="true" pageId="0" pageNumber="61">Psiloderces gawanaensis</emphasis>
</taxonomicName>
sp. nov., male holotype.
<emphasis id="CFAC4C0F5BD3B935038403F84463B5C3" bold="true" pageId="0" pageNumber="61">A</emphasis>
Palp, ventral view
<emphasis id="BCA88ADC0F2EB3ACF37902F4392A321A" bold="true" pageId="0" pageNumber="61">B</emphasis>
bulb, ventral view
<emphasis id="C34E4B993B96B0BDFB3AE0198FF6D082" bold="true" pageId="0" pageNumber="61">C</emphasis>
palp, prolateral view
<emphasis id="7C17D9380A9FDB8FBF187BB7786A21A4" bold="true" pageId="0" pageNumber="61">D</emphasis>
palp, retrolateral view. Abbreviations: BT = bristle, CP = cymbial protrusion, EM = embolus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="ED114E26A8BB5FA10F10BEA28B824B40" pageId="0" pageNumber="61" type="description">
<paragraph id="A264FEF4A33BE9D16F6FD054D98B19C0" pageId="0" pageNumber="61">Description.</paragraph>
<paragraph id="01D678B56A6A65A0C3E157C079AC03F9" pageId="0" pageNumber="61">
<emphasis id="248C6A4769ADB45F9A293579DC2CCF5B" bold="true" pageId="0" pageNumber="61">Male</emphasis>
(Holotype). Total length 1.54; carapace 0.60 long, 0.63 wide; abdomen 0.94 long, 0.56 wide. Carapace wider than long, brown, with 3 longitudinal dark brown bands, median band half length of carapace, median band two times wider than lateral band (Fig.
<figureCitation id="E88ED8C89824672A1C43834654E1421C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">3D</figureCitation>
). Chelicerae pale brown (Fig.
<figureCitation id="D9D1F0E79C3CF7D643B56A3CCF83B834" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Cheliceral retromargin, posterior view. A Psiloderces bangkiraiensis sp. nov. B P. wangou sp. nov. C P. heise sp. nov. D P. cuyapoensis sp. nov. E P. bontocensis sp. nov. F P. gawanaensis sp. nov. G P. xichang sp. nov. H P. cattienensis sp. nov. I P. pingguo sp. nov. Abbreviations: CL = cheliceral laminar, RT = retromargin teeth." figureDoi="10.3897/zookeys.902.38967.figure28" httpUri="https://binary.pensoft.net/fig/370872" pageId="0" pageNumber="61">28F</figureCitation>
). Clypeus dark brown, with long and slightly bent medial projection, bearing few setae apically (Fig.
<figureCitation id="08C89FD39D1B9851824D8DF563CE7F1C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">3C, D</figureCitation>
). Endites dark brown. Labium dark brown delimiting pair of light brown circular spots. Sternum dark brown, delimiting light brown band medially. Abdomen elongated, dorsum with brown stripes, antero-ventrally brown with elliptical patch, posterior part with indistinct brown pattern. Legs uniformly brown; measurements: I 5.21 (1.33, 0.25, 1.63, 1.20, 0.80), II 4.03 (1.09, 0.20, 1.25, 0.94, 0.55), III 3.49 (0.94, 0.20, 0.94, 0.86, 0.55), IV 4.88 (1.25, 0.25, 1.38, 1.25, 0.75). Palp (Fig.
<figureCitation id="E2BADFFFA4E97303B698608B8BAC93B2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Psiloderces gawanaensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BT = bristle, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.902.38967.figure4" httpUri="https://binary.pensoft.net/fig/370848" pageId="0" pageNumber="61">4A-D</figureCitation>
): femur slender, 4 times longer than patella; patella not swollen; tibia 3 times shorter than femur; cymbium purplish distally, 2 times shorter than wide, and 2 times wider than femur, protrusion with 4 distinct bristles basally and anteriorly; bulb light brown, inverted pyriform with threadlike embolus arising distally (Fig.
<figureCitation id="A6694EF70754222DECBFB05711D3BCBB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Psiloderces gawanaensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BT = bristle, CP = cymbial protrusion, EM = embolus." figureDoi="10.3897/zookeys.902.38967.figure4" httpUri="https://binary.pensoft.net/fig/370848" pageId="0" pageNumber="61">4B</figureCitation>
).
</paragraph>
<paragraph id="1DBE32F897D137681EAA25B16B8903DD" pageId="0" pageNumber="61">
<emphasis id="DB8B90169B86E7B2E08D342C14843E06" bold="true" pageId="0" pageNumber="61">Female</emphasis>
(Paratype). General features and coloration similar to those of male (Fig.
<figureCitation id="5A964E134ACE53ACA93737F5B019AB9D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">3E, F</figureCitation>
). Measurements: total length 1.64; carapace 0.70 long, 0.60 wide; abdomen 0.94 long, 1.20 wide. Leg measurements: I 5.23 (1.25, 0.25, 1.60, 1.25, 0.88), II 4.30 (1.10, 0.20, 1.30, 1.00, 0.70), III missing, IV 5.24 (1.30, 0.23, 1.60, 1.33, 0.78). Endogyne (Fig.
<figureCitation id="017DB93B66C56EFC2FD515742724B802" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">3A</figureCitation>
): encircled spermathecae resemble a horseshoe, curving downwards (Fig.
<figureCitation id="3D29D9DEDF603CFB2AFEB640B661871F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Psiloderces gawanaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, lateral view D male habitus, dorsal view E female habitus, dorsal view F female habitus, ventral view. Abbreviations: PR = clypeal protrusion, SP = spermatheca." figureDoi="10.3897/zookeys.902.38967.figure3" httpUri="https://binary.pensoft.net/fig/370847" pageId="0" pageNumber="61">3A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="99E23751C6E6F40F97302C02E52B4F18" pageId="0" pageNumber="61" type="distribution">
<paragraph id="01289C75D7DD9C688D1C16E6A8CA7374" pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph id="08FF5D38A2B7EA089A6C2AAA8CCCF625" pageId="0" pageNumber="61">
Known only from the type locality (Fig.
<figureCitation id="8F436D9C10DE653381D6527795101A65" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Distribution of new Psiloderces species in Southeast Asia. 1 P. heise sp. nov. 2 P. gawanaensis sp. nov. 3 P. bontocensis sp. nov. 4 P. cuyapoensis sp. nov. 5 P. xichang sp. nov. 6 P. cattienensis sp. nov. 7 P. pingguo sp. nov. 8 P. palopoensis sp. nov. 9 P. penajamensis sp. nov. 10 P. grohotensis sp. nov. 11 P. bangkiraiensis sp. nov. 12 P. bolang sp. nov. 13 P. wangou sp. nov. 14 P. malinoensis sp. nov." figureDoi="10.3897/zookeys.902.38967.figure30" httpUri="https://binary.pensoft.net/fig/370874" pageId="0" pageNumber="61">30</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>