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<document id="AEB1553C08E4EE6F4E93DFD2C7893C7A" ID-CLB-Dataset="5601" ID-DOI="10.11646/zootaxa.3796.2.8" ID-GBIF-Dataset="72f6fdb3-a9cb-4823-ada5-233a4510b0db" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915243" ID-ZooBank="9F9DA323-128B-44A7-9414-798B50DAAC44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623199009682" checkinUser="felipe" docAuthor="Maruyama, Munetoshi, Komatsu, Takashi, Katayama, Yuji, Song, Xiao-Bin &amp; Sakchoowong, Watana" docDate="2014" docId="47028781714FFFBA7E91FAE5AF19FB30" docLanguage="en" docName="zt03796p373.pdf" docOrigin="Zootaxa 3796 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aenictosymbia Maruyama 2014, gen. n." docType="treatment" docVersion="6" lastPageNumber="365" masterDocId="BB3BFFF9714CFFBE7E06FFECAA48FFF5" masterDocTitle="Myrmecophilous rove beetles (Coleoptera: Staphylinidae) associated with Aenictus hodgsoni (Hymenoptera: Formicidae) from Thailand, with description of two new genera and three new species" masterLastPageNumber="373" masterPageNumber="361" pageNumber="364" updateTime="1699006430599" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="23B2D3268CF0F570D2282B46C1741D01">Myrmecophilous rove beetles (Coleoptera: Staphylinidae) associated with Aenictus hodgsoni (Hymenoptera: Formicidae) from Thailand, with description of two new genera and three new species</mods:title>
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<mods:namePart id="F96ACED2C7E9097144A184F856648680">Maruyama, Munetoshi</mods:namePart>
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<mods:namePart id="47E15F6D41D6A5C1EC9DD5F89C418B3D">Komatsu, Takashi</mods:namePart>
<mods:affiliation id="3E3D5657C0A74DABA2592C6F53E5E33C">Department of Biology, Faculty of Science, Shinshu University, 3 - 1 - 1 Asahi, Matsumoto, Nagano 390 - 8621, Japan</mods:affiliation>
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<mods:namePart id="CC3163121FA987A561AA7C9759B261F9">Katayama, Yuji</mods:namePart>
<mods:affiliation id="00FE5ED0805B61EB3C28ECFC140FA4F2">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi, Matsuyama 790 - 8566, Japan</mods:affiliation>
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<mods:namePart id="13D5F9BCBC57286A018E10143F51B889">Song, Xiao-Bin</mods:namePart>
<mods:affiliation id="84449C764F61C19515A32A55140CEBA0">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China</mods:affiliation>
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<mods:namePart id="2541A6B2BE54EF1BA8D0D8FC8F2883B2">Sakchoowong, Watana</mods:namePart>
<mods:affiliation id="6B2F913DD2EFB4E53A02440DFD6E6993">Forest Entomology and Microbiology Group, Department of National Parks, Wildlife and Plant Conservation, 61 Paholyothin Road, Jatujak, Bangkok 10900, Thailand</mods:affiliation>
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<treatment id="47028781714FFFBA7E91FAE5AF19FB30" ID-DOI="http://doi.org/10.5281/zenodo.4915257" ID-GBIF-Taxon="183403948" ID-Zenodo-Dep="4915257" LSID="urn:lsid:plazi:treatment:47028781714FFFBA7E91FAE5AF19FB30" httpUri="http://treatment.plazi.org/id/47028781714FFFBA7E91FAE5AF19FB30" lastPageId="4" lastPageNumber="365" pageId="3" pageNumber="364">
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<taxonomicName id="08AB4D14714FFFBD7E91FAE5AB96FAD1" ID-CoL="8X6ZW" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[151,478,1289,1316]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FDDFEA85714FFFBD7E91FAE5AB01FAD6" bold="true" box="[151,329,1289,1315]" italics="true" pageId="3" pageNumber="364">Aenictosymbia</emphasis>
Maruyama
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,
<taxonomicNameLabel id="E6EC57FE714FFFBD7FECFAE6A875FAD1" box="[490,573,1290,1316]" pageId="3" pageNumber="364" rank="genus">gen. n.</taxonomicNameLabel>
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<subSubSection id="87B1651C714FFFBD7E91FAC0AB66FAB0" box="[151,302,1324,1349]" pageId="3" pageNumber="364" type="description">
<paragraph id="CF143697714FFFBD7E91FAC0AB66FAB0" blockId="3.[151,573,1289,1349]" box="[151,302,1324,1349]" pageId="3" pageNumber="364">
(
<figureCitation id="57902A12714FFFBD7E99FAC0AB6EFAB0" box="[159,294,1324,1349]" captionStart-0="FIGURES 1012" captionStart-1="FIGURES 1323" captionStart-2="FIGURES 2432" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionStartId-2="7.[151,263,1952,1975]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetBox-2="[161,1430,212,1917]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetId-2="figure-65@7.[151,1436,193,1932]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 1012. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." captionText-2="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" figureDoi-2="http://doi.org/10.5281/zenodo.4915259" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" httpUri-2="https://zenodo.org/record/4915259/files/figure.png" pageId="3" pageNumber="364">Figs. 1032</figureCitation>
)
</paragraph>
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<subSubSection id="87B1651C714FFFBD7E91FA98A944FA78" box="[151,780,1396,1421]" pageId="3" pageNumber="364" type="type_taxon">
<paragraph id="CF143697714FFFBD7E91FA98A944FA78" blockId="3.[151,1437,1396,2033]" box="[151,780,1396,1421]" pageId="3" pageNumber="364">
<emphasis id="FDDFEA85714FFFBD7E91FA98AB7CFA78" bold="true" box="[151,308,1396,1421]" pageId="3" pageNumber="364">
<typeStatus id="10108835714FFFBD7E91FA98AA9AFA78" box="[151,210,1396,1421]" pageId="3" pageNumber="364">Type</typeStatus>
species:
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<taxonomicName id="08AB4D14714FFFBD7F3DFA99A889FA78" ID-CoL="659TB" authority="Maruyama" authorityName="Maruyama" authorityYear="2014" box="[315,705,1397,1421]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">
<emphasis id="FDDFEA85714FFFBD7F3DFA99A876FA79" box="[315,574,1397,1421]" italics="true" pageId="3" pageNumber="364">Aenictosymbia cornuta</emphasis>
Maruyama
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,
<emphasis id="FDDFEA85714FFFBD7CCAFA98A944FA78" bold="true" box="[716,780,1396,1421]" pageId="3" pageNumber="364">
<taxonomicNameLabel id="E6EC57FE714FFFBD7CCAFA98A944FA78" box="[716,780,1396,1421]" pageId="3" pageNumber="364" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="87B1651C714FFFBA7EC1FA74A98CFD05" lastPageId="4" lastPageNumber="365" pageId="3" pageNumber="364" type="description">
<paragraph id="CF143697714FFFBD7EC1FA74A931FA44" blockId="3.[151,1437,1396,2033]" box="[199,889,1432,1457]" pageId="3" pageNumber="364">
<emphasis id="FDDFEA85714FFFBD7EC1FA74AB10FA44" bold="true" box="[199,344,1432,1457]" inLineHeading="true" pageId="3" pageNumber="364" reason="1">Description.</emphasis>
Body (
<figureCitation id="57902A12714FFFBD7FADFA74A879FA44" box="[427,561,1432,1457]" captionStart="FIGURES 1012" captionStartId="5.[151,263,1891,1914]" captionTargetBox="[189,1408,208,1848]" captionTargetId="figure-39@5.[151,1436,193,1870]" captionTargetPageId="5" captionText="FIGURES 1012. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915251" httpUri="https://zenodo.org/record/4915251/files/figure.png" pageId="3" pageNumber="364">Figs. 1012</figureCitation>
) myrmecoid, slender, glossy.
</paragraph>
<paragraph id="CF143697714FFFBD7EC1FA50A8BBF9E8" blockId="3.[151,1437,1396,2033]" pageId="3" pageNumber="364">
Head (
<figureCitation id="57902A12714FFFBD7F17FA50ABDEFA20" box="[273,406,1468,1493]" captionStart-0="FIGURES 1012" captionStart-1="FIGURES 1323" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1012. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Figs. 1013</figureCitation>
) almost spherical, with horn on anterior part of frons, gently narrowed posteriad, constricted at base to form “neck”; medial area of head slightly depressed; eyes moderate in size, prominent; antennae (
<figureCitation id="57902A12714FFFBD7B62FA0CAA96F9E8" captionStart-0="FIGURES 1012" captionStart-1="FIGURES 1323" captionStartId-0="5.[151,263,1891,1914]" captionStartId-1="6.[151,263,1837,1860]" captionTargetBox-0="[189,1408,208,1848]" captionTargetBox-1="[151,1422,204,1800]" captionTargetId-0="figure-39@5.[151,1436,193,1870]" captionTargetId-1="figure-66@6.[151,1436,193,1817]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1012. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." captionText-1="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.4915251" figureDoi-1="http://doi.org/10.5281/zenodo.4915255" httpUri-0="https://zenodo.org/record/4915251/files/figure.png" httpUri-1="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Figs. 1013</figureCitation>
) generalized, but segment I strongly thickened.
</paragraph>
<paragraph id="CF143697714FFFBD7EC1F9C4A88DF871" blockId="3.[151,1437,1396,2033]" pageId="3" pageNumber="364">
Mouthparts: Labrum (
<figureCitation id="57902A12714FFFBD7FC2F9C4A850F9B4" box="[452,536,1576,1601]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 14</figureCitation>
) generalized, rather irregularly with setae, without pseudopores; hypopharynx (
<figureCitation id="57902A12714FFFBD7E99F9A0AAB8F990" box="[159,240,1612,1637]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 14</figureCitation>
) with mesal area bearing 3 campanifom sensillae, 3 setulae antero-medially, 4 setulae along lateral margin. Mandibles (
<figureCitation id="57902A12714FFFBD7F19F99CAB3AF97C" box="[287,370,1648,1673]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 15</figureCitation>
) almost symmetric, but left mandible slightly thicker, with 2 patches of minute pseudopores around mesal area of dorsal disc, and 2 setae near base of outer margin. Mentum (
<figureCitation id="57902A12714FFFBD7A4FF978AED5F958" box="[1097,1181,1684,1709]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 16</figureCitation>
) sub-trapezoidal, with 24 pseudopores, several setae around lateral margin. Labium (
<figureCitation id="57902A12714FFFBD7D69F954A98CF924" box="[879,964,1720,1745]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 17</figureCitation>
): apodeme with lateral lobe short, gently curved, rounded at apex, without medial projection; prementum with 2 real pores and 1 setal pore antero-laterally, strongly sclerotized along anterior margin to connect both bases of palpi; palpus with segment I long, more than 2 time as long as II, with patch of some minute setae near base (
<figureCitation id="57902A12714FFFBD7D71F8C8A985F8C8" box="[887,973,1828,1853]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 17</figureCitation>
: arrow; probably sensillae); segment III narrow, elongate; maxilla (
<figureCitation id="57902A12714FFFBD7FCDF8A4A856F894" box="[459,542,1864,1889]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 18</figureCitation>
) generalized, no or few pseudopores overall; lacinia with 5 spines near apex, densely with pubescence; galea curved near apex.
</paragraph>
<paragraph id="CF143697714FFFBD7EC1F87DAB70F804" blockId="3.[151,1437,1396,2033]" pageId="3" pageNumber="364">
Pronotum (
<figureCitation id="57902A12714FFFBD7F42F87CAB83F85C" box="[324,459,1936,1961]" captionStart="FIGURES 1012" captionStartId="5.[151,263,1891,1914]" captionTargetBox="[189,1408,208,1848]" captionTargetId="figure-39@5.[151,1436,193,1870]" captionTargetPageId="5" captionText="FIGURES 1012. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915251" httpUri="https://zenodo.org/record/4915251/files/figure.png" pageId="3" pageNumber="364">Figs. 1012</figureCitation>
,
<figureCitation id="57902A12714FFFBD7FD0F87CA855F85C" box="[470,541,1936,1961]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">19, 20</figureCitation>
) elongate, constricted around basal 1/3, shallowly sulcate at midline; prosternite (
<figureCitation id="57902A12714FFFBD7E99F858AAB8F838" box="[159,240,1972,1997]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 20</figureCitation>
) subtriangular, elongate, convex at midline; triangular sclerite present (
<figureCitation id="57902A12714FFFBD7A08F858AE2AF838" box="[1038,1122,1972,1997]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="3" pageNumber="364">Fig. 20</figureCitation>
: arrow) from middle to near base of cavity.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1FF7BA9DBFF0D" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
Mesoventrite (
<figureCitation id="57902A127148FFBA7F6DFF7BABF5FF45" box="[363,445,151,176]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Fig. 21</figureCitation>
) short, completely fused with metaventrite, weakly carinate between processes of both; metaventrite long, more than 2 times as long as mesoventrite; mesocoxal cavity slightly margined postero-laterally. Elytra (
<figureCitation id="57902A127148FFBA7EEDFF33AB74FF0D" box="[235,316,223,248]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Fig. 22</figureCitation>
) rounded, well convex above; hind wings developed.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1FEE8A9B6FE72" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
Legs (
<figureCitation id="57902A127148FFBA7F16FEE8ABD2FEE8" box="[272,410,260,285]" captionStart-0="FIGURES 1323" captionStart-1="FIGURES 2432" captionStartId-0="6.[151,263,1837,1860]" captionStartId-1="7.[151,263,1952,1975]" captionTargetBox-0="[151,1422,204,1800]" captionTargetBox-1="[161,1430,212,1917]" captionTargetId-0="figure-66@6.[151,1436,193,1817]" captionTargetId-1="figure-65@7.[151,1436,193,1932]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." captionText-1="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.4915255" figureDoi-1="http://doi.org/10.5281/zenodo.4915259" httpUri-0="https://zenodo.org/record/4915255/files/figure.png" httpUri-1="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Figs. 2325</figureCitation>
) long, slender; claws developed; fore leg (
<figureCitation id="57902A127148FFBA7D89FEE8A9ACFEE8" box="[911,996,260,285]" captionStart="FIGURES 1323" captionStartId="6.[151,263,1837,1860]" captionTargetBox="[151,1422,204,1800]" captionTargetId="figure-66@6.[151,1436,193,1817]" captionTargetPageId="6" captionText="FIGURES 1323. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 13―head, dorsal view; 14―labrum (right, epipharynx); 15―left mandible; 16―memtum; 17―labium; 18, right maxilla; 19―pronotum, dorsal view; 20―ditto, ventral view; 21―meso- and meta traces; 22―right elytra; 23―left fore leg." figureDoi="http://doi.org/10.5281/zenodo.4915255" httpUri="https://zenodo.org/record/4915255/files/figure.png" pageId="4" pageNumber="365">Fig. 23</figureCitation>
) with coxa projected at base; base of femur projected ventrad; tarsal segments subequal in length; mid leg (
<figureCitation id="57902A127148FFBA7DA2FECBA9BDFEB5" box="[932,1013,295,320]" captionStart="FIGURES 2432" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Fig. 24</figureCitation>
) with coxa small, slightly longer than trochanter; tarsal segments becoming smaller from base to apex; hind leg (
<figureCitation id="57902A127148FFBA7DF5FEA0AE0FFE90" box="[1011,1095,332,357]" captionStart="FIGURES 2432" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Fig. 25</figureCitation>
) with coxa deeply notched at middle (Fig, 25: arrow); tarsal segments becoming smaller from base to apex.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1FE78ABF6FDAA" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
Abdomen strongly constricted at base (segments IIVI); tergite II rather long, shallowly emarginate on anterior margin; tergite III elongate, dilated apicad; sternite III with posterior area strongly triangularly convex (
<figureCitation id="57902A127148FFBA7B44FE5BAFDFFE25" box="[1346,1431,439,464]" captionStart="FIGURES 1012" captionStartId="5.[151,263,1891,1914]" captionTargetBox="[189,1408,208,1848]" captionTargetId="figure-39@5.[151,1436,193,1870]" captionTargetPageId="5" captionText="FIGURES 1012. Habitus of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 10―dorsal view; 11―ventral view; 12―lateral view." figureDoi="http://doi.org/10.5281/zenodo.4915251" httpUri="https://zenodo.org/record/4915251/files/figure.png" pageId="4" pageNumber="365">Fig. 11</figureCitation>
: arrow; no distinct gland openings); tergites IVVII short, transverse; stermites IVVII large, strongly convex; tergite VIII with anterior margin rounded, without pseudopores or minute setae; sternite VIII with anterior margin slightly produced, rounded; tergite and sternite IX fused at base of tergite X of which anterior margin submembranous, truncate.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1FD80ABE2FD38" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
Male: median lobe of aedeagus (
<figureCitation id="57902A127148FFBA7C3EFD80A888FD70" box="[568,704,620,645]" captionStart="FIGURES 2432" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Figs. 29, 30</figureCitation>
) urceolate, strongly produced paramerally at base, with a pair of exposed sclerites of internal sac; paramere (
<figureCitation id="57902A127148FFBA7C8DFD63A896FD5D" box="[651,734,655,680]" captionStart="FIGURES 2432" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Fig. 31</figureCitation>
) with basal part of paramerite short, narrow; velum extended near apex of apical lobe.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1FD3BA98CFD05" blockId="4.[151,1437,151,1221]" box="[199,964,727,752]" pageId="4" pageNumber="365">
Female: spermatheca (
<figureCitation id="57902A127148FFBA7FC5FD3BA85DFD05" box="[451,533,727,752]" captionStart="FIGURES 2432" captionStartId="7.[151,263,1952,1975]" captionTargetBox="[161,1430,212,1917]" captionTargetId="figure-65@7.[151,1436,193,1932]" captionTargetPageId="7" captionText="FIGURES 2432. Body parts of Aenictosymbia cornuta Maruyama, gen. &amp; sp. n. 24―left mid leg; 25―left hind leg; 26―male tergite VIII; 27―male sternite VIII; 28―male abdominal tergites IX and X; 29―median lobe of aedeagus, lateral view; 30―ditto, apical part, ventra view; 31―paramere; 32―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4915259" httpUri="https://zenodo.org/record/4915259/files/figure.png" pageId="4" pageNumber="365">Fig. 32</figureCitation>
) coiled at base, with apical part small.
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7148FFBA7EC1FD10AF02FC8A" pageId="4" pageNumber="365" type="diagnosis">
<paragraph id="CF1436977148FFBA7EC1FD10AF02FC8A" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
<emphasis id="FDDFEA857148FFBA7EC1FD10AB87FCE0" bold="true" box="[199,463,764,789]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">Differential diagnosis.</emphasis>
This genus is similar to
<taxonomicName id="08AB4D147148FFBA7CEFFD10AE69FCE0" authority="Seevers, 1965" authorityName="Seevers" authorityYear="1965" box="[745,1057,764,789]" class="Insecta" family="Staphylinidae" genus="Aenictophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7CEFFD10A930FCE1" box="[745,888,764,788]" italics="true" pageId="4" pageNumber="365">Aenictophila</emphasis>
<bibRefCitation id="AB3A4B667148FFBA7D86FD10AE69FCE0" author="Seevers, C. H." box="[896,1057,764,789]" pageId="4" pageNumber="365" pagination="138 - 351" refId="ref5738" refString="Seevers, C. H. (1965) The systematics, evolution and zoogeography of staphylinid beetles associated with army ants (Coleoptera: Staphylinidae). Fieldiana: Zoology, 47, 138 - 351." type="journal article" year="1965">Seevers, 1965</bibRefCitation>
</taxonomicName>
, in general appearance, the states of meso- and metaventrites, and abdomen, but the convexity of the abdominal sternites is simpler in
<taxonomicName id="08AB4D147148FFBA7E91FCA8AB74FCA9" authorityName="Maruyama" authorityYear="2014" box="[151,316,836,860]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7E91FCA8AB74FCA9" box="[151,316,836,860]" italics="true" pageId="4" pageNumber="365">Aenictosymbia</emphasis>
</taxonomicName>
. In
<taxonomicName id="08AB4D147148FFBA7F76FCA8ABB7FCA9" authorityName="Seevers" authorityYear="1965" box="[368,511,836,860]" class="Insecta" family="Staphylinidae" genus="Aenictophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7F76FCA8ABB7FCA9" box="[368,511,836,860]" italics="true" pageId="4" pageNumber="365">Aenictophila</emphasis>
</taxonomicName>
, the posterior areas of both sternites III and IV are strongly convex, and their surfaces have distinct gland opening patches, including trichome-like bundles of pubescence on sternite III.
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7148FFBA7EC1FC60AF51FC3D" pageId="4" pageNumber="365" type="etymology">
<paragraph id="CF1436977148FFBA7EC1FC60AF51FC3D" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
<emphasis id="FDDFEA857148FFBA7EC1FC60AB06FC50" bold="true" box="[199,334,908,933]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">Etymology.</emphasis>
A combination of the host genus name
<taxonomicName id="08AB4D147148FFBA7D37FC60A9D9FC56" authorityName="Shuckard" authorityYear="1840" box="[817,913,908,931]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7D37FC60A9D9FC56" box="[817,913,908,931]" italics="true" pageId="4" pageNumber="365">Aenictus</emphasis>
</taxonomicName>
, and the Greek words
<emphasis id="FDDFEA857148FFBA7AA0FC61AE83FC51" box="[1190,1227,909,932]" italics="true" pageId="4" pageNumber="365">syn</emphasis>
(meaning &quot;with&quot;, &quot;together&quot;) and
<emphasis id="FDDFEA857148FFBA7F4FFC5CAB30FC3D" box="[329,376,944,968]" italics="true" pageId="4" pageNumber="365">bios</emphasis>
(meaning &quot;life&quot;, &quot;living&quot;), meaning a symbiont of
<taxonomicName id="08AB4D147148FFBA7DB4FC5CAE5AFC32" authorityName="Shuckard" authorityYear="1840" box="[946,1042,944,967]" class="Insecta" family="Formicidae" genus="Aenictus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7DB4FC5CAE5AFC32" box="[946,1042,944,967]" italics="true" pageId="4" pageNumber="365">Aenictus</emphasis>
</taxonomicName>
ants. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="87B1651C7148FFBA7EC1FC38AF19FB30" pageId="4" pageNumber="365" type="distribution">
<paragraph id="CF1436977148FFBA7EC1FC38A864FC18" blockId="4.[151,1437,151,1221]" box="[199,556,980,1005]" pageId="4" pageNumber="365">
<emphasis id="FDDFEA857148FFBA7EC1FC38AB28FC18" bold="true" box="[199,352,980,1005]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">Distribution.</emphasis>
Central
<collectingCountry id="B7BC76077148FFBA7FC6FC38A86FFC18" box="[448,551,980,1005]" name="Thailand" pageId="4" pageNumber="365">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="CF1436977148FFBA7EC1FC1BAF19FB30" blockId="4.[151,1437,151,1221]" pageId="4" pageNumber="365">
<emphasis id="FDDFEA857148FFBA7EC1FC1BABDFFBE5" bold="true" box="[199,407,1015,1040]" inLineHeading="true" pageId="4" pageNumber="365" reason="1">Systematic notes.</emphasis>
This genus apparently belongs to the
<taxonomicName id="08AB4D147148FFBA7D50FC14A9ADFBE5" authorityName="Seevers" authorityYear="1965" box="[854,997,1016,1040]" class="Insecta" family="Staphylinidae" genus="Aenictophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7D50FC14A9ADFBE5" box="[854,997,1016,1040]" italics="true" pageId="4" pageNumber="365">Aenictophila</emphasis>
</taxonomicName>
generic group, which is composed of myrmecoid genera of
<taxonomicName id="08AB4D147148FFBA7F8BFBF0A86AFBC0" authorityName="Fleming" authorityYear="1821" box="[397,546,1052,1077]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
, namely,
<taxonomicName id="08AB4D147148FFBA7C8DFBF0A952FBC1" authorityName="Seevers" authorityYear="1965" box="[651,794,1052,1076]" class="Insecta" family="Staphylinidae" genus="Aenictophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7C8DFBF0A952FBC1" box="[651,794,1052,1076]" italics="true" pageId="4" pageNumber="365">Aenictophila</emphasis>
</taxonomicName>
,
<taxonomicName id="08AB4D147148FFBA7D21FBF1AEACFBC0" authority="Kistner &amp; Jacobson, 1975" authorityName="Kistner &amp; Jacobson" authorityYear="1975" box="[807,1252,1052,1077]" class="Insecta" family="Staphylinidae" genus="Mimaenictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7D21FBF1A9F0FBC1" box="[807,952,1053,1076]" italics="true" pageId="4" pageNumber="365">Mimaenictus</emphasis>
<bibRefCitation id="AB3A4B667148FFBA7DB9FBF0AEACFBC0" author="Jacobson, H. R. &amp; Kistner, D. H." box="[959,1252,1052,1077]" pageId="4" pageNumber="365" pagination="163 - 200" refId="ref5319" refString="Jacobson, H. R. &amp; Kistner, D. H. (1975) The natural history of Pygostenini. Section 1. New species and new records of Pygostenini. Sociobiology, 1, 163 - 200." type="journal article" year="1975">Kistner &amp; Jacobson, 1975</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="08AB4D147148FFBA7AE9FBF1AB88FBAD" authority="Kistner &amp; Jacobson, 1975" authorityName="Kistner &amp; Jacobson" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Procantonnetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7AE9FBF1AFD4FBC1" box="[1263,1436,1053,1076]" italics="true" pageId="4" pageNumber="365">Procantonnetia</emphasis>
<bibRefCitation id="AB3A4B667148FFBA7E91FBD3AB88FBAD" author="Jacobson, H. R. &amp; Kistner, D. H." box="[151,448,1087,1112]" pageId="4" pageNumber="365" pagination="163 - 200" refId="ref5319" refString="Jacobson, H. R. &amp; Kistner, D. H. (1975) The natural history of Pygostenini. Section 1. New species and new records of Pygostenini. Sociobiology, 1, 163 - 200." type="journal article" year="1975">Kistner &amp; Jacobson, 1975</bibRefCitation>
</taxonomicName>
,
<emphasis id="FDDFEA857148FFBA7FCAFBACA804FBAD" box="[460,588,1088,1112]" italics="true" pageId="4" pageNumber="365">Weissflogia</emphasis>
<bibRefCitation id="AB3A4B667148FFBA7C52FBD3A8A7FBAD" author="Kistner, D. H. &amp; Weissflog, A. &amp; Rosciszewski, K. &amp; Maschwitz, U." box="[596,751,1087,1112]" pageId="4" pageNumber="365" pagination="123 - 221" refId="ref5456" refString="Kistner, D. H., Weissflog, A., Rosciszewski, K. &amp; Maschwitz, U. (1997) New species, new genera, and new records of myrmecophiles associated with army ants (Aenictus sp.) with the description of a new subtribe of Staphylinidae (Coleoptera; Hymenoptera, Formicidae). Sociobiology, 29, 123 - 221." type="journal article" year="1997">Kistner, 1997</bibRefCitation>
. These genera including
<taxonomicName id="08AB4D147148FFBA7A0DFBACAEF8FBAD" authorityName="Maruyama" authorityYear="2014" box="[1035,1200,1088,1112]" class="Insecta" family="Staphylinidae" genus="Aenictosymbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="FDDFEA857148FFBA7A0DFBACAEF8FBAD" box="[1035,1200,1088,1112]" italics="true" pageId="4" pageNumber="365">Aenictosymbia</emphasis>
</taxonomicName>
share an exoskeletal modification of a gland on abdominal sternite III (IIIV in some genera). This characteristic, including the presence of a gland on sternite III, is not observed in other
<taxonomicName id="08AB4D147148FFBA7D41FB6BA991FB55" authorityName="Fleming" authorityYear="1821" box="[839,985,1159,1184]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
genera and suggests monophyly of the generic group. Phylogenetics and a detailed morphological study are in progress by MM and his colleagues.
</paragraph>
</subSubSection>
</treatment>
</document>