189 lines
20 KiB
XML
189 lines
20 KiB
XML
<document id="105C8DB27A16196E753C029065F2DEDA" ID-DOI="10.11646/zootaxa.3215.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5248640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629765289367" checkinUser="felipe" docAuthor="Van Bakel, Barry W. M., Guinot, Danièle, Artal, Pedro, Fraaije, René H. B. & Jagt, John W. M." docDate="2012" docId="4601C935FFF7F90B5BB4FC17F123F8D5" docLanguage="en" docName="zt03215p216.pdf" docOrigin="Zootaxa 3215 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3215.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eosymethis aragonensis Van Bakel & Guinot & Artal & Fraaije & Jagt 2012, n. sp." docType="treatment" docVersion="6" lastPageNumber="104" masterDocId="BA38B14DFF90F96C5B23FFB4F45FFFB6" masterDocTitle="A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215" masterLastPageNumber="216" masterPageNumber="1" pageNumber="104" updateTime="1699109082209" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="67A0D26846BFE6213631A76BAC8068F4">A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215</mods:title>
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<mods:namePart id="9A74C11AE405D62A791E941B7348B87C">Van Bakel, Barry W. M.</mods:namePart>
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<mods:namePart id="227C734D93AF8A911A7F31060F3C7DD9">Guinot, Danièle</mods:namePart>
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<mods:namePart id="8D2D4ABD9F851B8F5BB319E78761C955">Artal, Pedro</mods:namePart>
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<mods:namePart id="86FF348F8D036D7FA82B789669DC88FA">Fraaije, René H. B.</mods:namePart>
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<mods:namePart id="C8B42DD354BC9E32CD550FA6DEE53902">Jagt, John W. M.</mods:namePart>
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<mods:title id="71A331054EAB833D6F8A3E0BC0C7B548">Zootaxa</mods:title>
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<mods:date id="F886FCDD8C1C715ED0BFC30CF7C8C8EB">2012</mods:date>
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<mods:number id="4EFF0C935392B94E3BF17BBB38D7ABBD">2012-02-29</mods:number>
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<mods:number id="F394D51BE08EC5ABFE7961E9ED2AB122">3215</mods:number>
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<treatment id="4601C935FFF7F90B5BB4FC17F123F8D5" ID-DOI="http://doi.org/10.5281/zenodo.5250478" ID-Zenodo-Dep="5250478" LSID="urn:lsid:plazi:treatment:4601C935FFF7F90B5BB4FC17F123F8D5" httpUri="http://treatment.plazi.org/id/4601C935FFF7F90B5BB4FC17F123F8D5" lastPageNumber="104" pageId="103" pageNumber="104">
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<subSubSection id="86B22BA8FFF7F90B5BB4FC17F5ABFC0D" box="[151,500,931,956]" pageId="103" pageNumber="104" type="nomenclature">
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<paragraph id="CE177823FFF7F90B5BB4FC17F5ABFC0D" blockId="103.[151,500,931,991]" box="[151,500,931,956]" pageId="103" pageNumber="104">
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<heading id="955FCF4FFFF7F90B5BB4FC17F5ABFC0D" bold="true" box="[151,500,931,956]" fontSize="10" level="2" pageId="103" pageNumber="104" reason="2">
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<taxonomicName id="09A803A0FFF7F90B5BB4FC17F5F1FC0A" ID-CoL="8a25e9c6-d4cc-44a7-b147-dff5beb821b9" authority="Van Bakel & Guinot & Artal & Fraaije & Jagt, 2012" authorityName="Van Bakel & Guinot & Artal & Fraaije & Jagt" authorityYear="2012" box="[151,430,931,956]" class="Malacostraca" family="Raninidae" genus="Eosymethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="aragonensis" status="sp. nov.">Eosymethis aragonensis</taxonomicName>
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<taxonomicNameLabel id="E7EF194AFFF7F90B5A96FC10F5ABFC0D" box="[437,500,932,956]" pageId="103" pageNumber="104" rank="species">
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<emphasis id="FCDCA431FFF7F90B5A96FC10F5B2FC0A" bold="true" box="[437,493,932,956]" pageId="103" pageNumber="104">
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<emphasis id="FCDCA431FFF7F90B5A96FC10F593FC0A" box="[437,460,932,956]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">n.</emphasis>
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sp
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</emphasis>
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.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="86B22BA8FFF7F90B5BB4FC73F571FC69" box="[151,302,967,991]" pageId="103" pageNumber="104" type="description">
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<paragraph id="CE177823FFF7F90B5BB4FC73F571FC69" blockId="103.[151,500,931,991]" box="[151,302,967,991]" pageId="103" pageNumber="104">
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(
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<figureCitation id="569364A6FFF7F90B5BBCFC73F579FC69" box="[159,294,967,991]" captionStart="FIGURE 32" captionStartId="104.[151,250,1712,1735]" captionTargetBox="[151,1435,406,1691]" captionTargetId="figure-106@104.[151,1435,406,1691]" captionTargetPageId="104" captionText="FIGURE 32. A–C. Eosymethis aragonensis n. gen., n. sp. (Raninidae, Symethinae), lower Eocene (Ypresian), Isabena (Huesca, northern Spain), MGSB75293 (holotype; ex José Enrique Ortega Collection; indeterminate sex); overview of specimen, detail showing carapace, and detail showing falciform dactylus of right P2/P3. Scale bars: 5mm." figureDoi="http://doi.org/10.5281/zenodo.5248680" httpUri="https://zenodo.org/record/5248680/files/figure.png" pageId="103" pageNumber="104">Fig. 32A–C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="86B22BA8FFF7F90B5BB4FBBBF7FBFBFD" pageId="103" pageNumber="104" type="diagnosis">
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<paragraph id="CE177823FFF7F90B5BB4FBBBF5F0FB9E" blockId="103.[151,1436,1039,1892]" box="[151,431,1039,1064]" pageId="103" pageNumber="104">
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<emphasis id="FCDCA431FFF7F90B5BB4FBBBF54EFB9E" bold="true" box="[151,273,1039,1064]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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<paragraph id="CE177823FFF7F90B5BE4FB87F7FBFBFD" blockId="103.[151,1436,1039,1892]" box="[199,932,1075,1100]" pageId="103" pageNumber="104">
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<emphasis id="FCDCA431FFF7F90B5BE4FB87F5EBFBFA" bold="true" box="[199,436,1075,1100]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Derivation of name.</emphasis>
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From the region of
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<collectingRegion id="0C6CB6C1FFF7F90B59B6FB87F6B3FBFD" box="[661,748,1075,1099]" country="Spain" name="Aragon" pageId="103" pageNumber="104">Aragón</collectingRegion>
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, northern
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<collectingCountry id="B6BF38B3FFF7F90B587FFB87F7C0FBFD" box="[860,927,1075,1099]" name="Spain" pageId="103" pageNumber="104">Spain</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="86B22BA8FFF7F90B5BE4FBE3F767FB6D" pageId="103" pageNumber="104" type="materials_examined">
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<paragraph id="CE177823FFF7F90B5BE4FBE3F767FB6D" blockId="103.[151,1436,1039,1892]" pageId="103" pageNumber="104">
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||
<emphasis id="FCDCA431FFF7F90B5BE4FBE3F5EFFBC6" bold="true" box="[199,432,1111,1136]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Material examined.</emphasis>
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<materialsCitation id="7EC0727EFFF7F90B5A95FBE3F767FB6D" collectionCode="MGSB" county="upper Ilerdian'" location="Eocene" municipality="Ypresian" pageId="103" pageNumber="104" specimenCode="MGSB75293" specimenCount="1" stateProvince="Aragon" typeStatus="holotype">
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<typeStatus id="1113C681FFF7F90B5A95FBE3F67EFBD9" box="[438,545,1111,1135]" pageId="103" pageNumber="104" type="holotype">Holotype</typeStatus>
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, and only specimen so far known, a near-complete carapace with remains of the pereiopods, preserved as a natural mould, from the marls of the ‘
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<collectingCounty id="277600AFFFF7F90B58AAFBCFF067FB25" box="[905,1080,1147,1171]" pageId="103" pageNumber="104">upper Ilerdian’</collectingCounty>
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(lowermost
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<collectingMunicipality id="2E73E259FFF7F90B5FEFFBCFF16DFB25" box="[1228,1330,1147,1171]" pageId="103" pageNumber="104">Ypresian</collectingMunicipality>
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,
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||
<location id="CB772EF8FFF7F90B5E63FBCFF1CBFB25" LSID="urn:lsid:plazi:treatment:4601C935FFF7F90B5BB4FC17F123F8D5:CB772EF8FFF7F90B5E63FBCFF1CBFB25" box="[1344,1428,1147,1171]" county="upper Ilerdian'" municipality="Ypresian" name="Eocene" pageId="103" pageNumber="104" stateProvince="Aragon">Eocene</location>
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)
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<location id="CB772EF8FFF7F90B5BB4FB2BF50FFB01" LSID="urn:lsid:plazi:treatment:4601C935FFF7F90B5BB4FC17F123F8D5:CB772EF8FFF7F90B5BB4FB2BF50FFB01" box="[151,336,1183,1207]" county="upper Ilerdian'" municipality="Ypresian" name="Roda Formation" pageId="103" pageNumber="104" stateProvince="Aragon">Roda Formation</location>
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(
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<bibRefCitation id="AA3905D2FFF7F90B5A42FB2BF6C7FB01" author="Cuevas-Gozalo, M. & Donselaar, M. E. & Nio, S. D." box="[353,664,1183,1207]" pageId="103" pageNumber="104" pagination="215 - 266" refId="ref123572" refString="Cuevas-Gozalo, M., Donselaar, M. E. & Nio, S. D. (1985) Eocene clastic tidal deposits in the Tremp-Graus Basin (Prov. of Lerida and Huesca). 6 th European Regional Meeting IAS, Lerida, Guidebook Excursion no. 6, 215 - 266." type="journal article" year="1985">
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Cuevas-Gozalo
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<emphasis id="FCDCA431FFF7F90B593AFB14F612FB01" box="[537,589,1183,1207]" italics="true" pageId="103" pageNumber="104">et al</emphasis>
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||
. 1985
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</bibRefCitation>
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), approximately
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<quantity id="0950D5C6FFF7F90B5879FB14F7C9FB01" box="[858,918,1183,1207]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="103" pageNumber="104" unit="km" value="7.0">7 km</quantity>
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northwest of the village of
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<location id="CB772EF8FFF7F90B5FF9FB2BF170FB01" LSID="urn:lsid:plazi:treatment:4601C935FFF7F90B5BB4FC17F123F8D5:CB772EF8FFF7F90B5FF9FB2BF170FB01" box="[1242,1327,1183,1207]" county="upper Ilerdian'" municipality="Ypresian" name="Isabena" pageId="103" pageNumber="104" stateProvince="Aragon">Isabena</location>
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||
(
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<location id="CB772EF8FFF7F90B5E62FB2BF1C7FB01" LSID="urn:lsid:plazi:treatment:4601C935FFF7F90B5BB4FC17F123F8D5:CB772EF8FFF7F90B5E62FB2BF1C7FB01" box="[1345,1432,1183,1207]" county="upper Ilerdian'" municipality="Ypresian" name="Huesca" pageId="103" pageNumber="104" stateProvince="Aragon">Huesca</location>
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,
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<collectingRegion id="0C6CB6C1FFF7F90B5BB4FB77F4B2FB6D" box="[151,237,1219,1243]" country="Spain" name="Aragon" pageId="103" pageNumber="104">Aragón</collectingRegion>
|
||
) (
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||
<specimenCode id="9E0ED058FFF7F90B5A20FB77F5C1FB6D" box="[259,414,1219,1243]" collectionCode="MGSB" pageId="103" pageNumber="104">MGSB75293</specimenCode>
|
||
;
|
||
<emphasis id="FCDCA431FFF7F90B5A8BFB70F59EFB6D" box="[424,449,1220,1243]" italics="true" pageId="103" pageNumber="104">ex</emphasis>
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||
José Enrique Ortega Collection).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="86B22BA8FFF7F90B5BE4FB53F178F9D1" pageId="103" pageNumber="104" type="description">
|
||
<paragraph id="CE177823FFF7F90B5BE4FB53F178F9D1" blockId="103.[151,1436,1039,1892]" pageId="103" pageNumber="104">
|
||
<emphasis id="FCDCA431FFF7F90B5BE4FB53F507FAB6" bold="true" box="[199,344,1255,1280]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Description.</emphasis>
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||
Carapace small, subovally elongated in outline, longer than wide; maximum width at confluence of lateral margins, posterior of mid-length. Dorsal surface slightly concave in both directions. Orbitofrontal margin narrow, markedly advanced; orbits small, directed anteriorly; front strongly projected with triangular tip. Anterolateral margins long, with notable angular hepatic node after concave first portion; posterior portion nearly straight, diverging posteriorly. Posterolateral margins shorter, from straight to slightly concave, rimmed, convergent to the rear. Posterior margin fairly concave, much broader than orbitofrontal margin. Dorsal regions poorly defined; postfrontal depression present; gastric regions weakly defined; branchial regions with notable oblique branchial scars, lateratelly disposed; cardiac region large, elongated, bounded by marked branchiocardiac grooves, with angular node at lateral upper portion; anterior portion separated from gastric regions by subtle groove. Dorsal surface covered by large, dense pits, uniformly distributed over entire carapace; some granules in posterior portion of posterolateral margins. Pereiopods partially preserved, showing short, flattened articles with falciform dactylus.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="86B22BA8FFF7F90B5BE4F9C0F123F8D5" pageId="103" pageNumber="104" type="discussion">
|
||
<paragraph id="CE177823FFF7F90B5BE4F9C0F123F8D5" blockId="103.[151,1436,1039,1892]" pageId="103" pageNumber="104">
|
||
<emphasis id="FCDCA431FFF7F90B5BE4F9C0F564F93A" bold="true" box="[199,315,1652,1676]" inLineHeading="true" pageId="103" pageNumber="104" reason="1">Remarks.</emphasis>
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The species exhibits features that permit placement in
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<taxonomicName id="09A803A0FFF7F90B588AF9C7F06DF93D" authority="Van Bakel & Guinot & Artal & Fraaije & Jagt, 2012" authorityName="Van Bakel & Guinot & Artal & Fraaije & Jagt" authorityYear="2012" box="[937,1074,1651,1675]" class="Malacostraca" family="Raninidae" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="185" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Symethinae">Symethinae</taxonomicName>
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: the carapace has a narrow, projected anterior portion of the carapace and a post-frontal depression; the dorsal surface shows an isolated, lateral, acute groove; the anterolateral margin has only a diminutive node; and falciform dactyli of the pereiopods.
|
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<taxonomicName id="09A803A0FFF7F90B5E70F90FF484F941" authority="Van Bakel & Guinot & Artal & Fraaije & Jagt, 2012" authorityName="Van Bakel & Guinot & Artal & Fraaije & Jagt" authorityYear="2012" class="Malacostraca" family="Raninidae" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="185" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Symethinae">Symethinae</taxonomicName>
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||
so far consists of a single genus,
|
||
<taxonomicName id="09A803A0FFF7F90B5972F96BF6EBF941" authorityName="Weber" authorityYear="1795" box="[593,692,1759,1783]" class="Malacostraca" family="Raninidae" genus="Symethis" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="104" phylum="Arthropoda" rank="genus">
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||
<emphasis id="FCDCA431FFF7F90B5972F96BF6EBF941" box="[593,692,1759,1783]" italics="true" pageId="103" pageNumber="104">Symethis</emphasis>
|
||
</taxonomicName>
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, with only three extant species (
|
||
<bibRefCitation id="AA3905D2FFF7F90B5F3FF96BF0B3F941" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1052,1260,1759,1783]" pageId="103" pageNumber="104" pagination="1 - 286" refId="ref139027" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement no. 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="FCDCA431FFF7F90B5F64F954F020F941" box="[1095,1151,1759,1783]" italics="true" pageId="103" pageNumber="104">et al.</emphasis>
|
||
2008: 43
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||
</bibRefCitation>
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||
).
|
||
<taxonomicName id="09A803A0FFF7F90B5FDCF96BF121F941" authority="Van Bakel & Guinot & Artal & Fraaije & Jagt, 2012" authorityName="Van Bakel & Guinot & Artal & Fraaije & Jagt" authorityYear="2012" box="[1279,1406,1759,1783]" class="Malacostraca" family="Raninidae" genus="Eosymethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="104" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FCDCA431FFF7F90B5FDCF96BF121F941" box="[1279,1406,1759,1783]" italics="true" pageId="103" pageNumber="104">Eosymethis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FCDCA431FFF7F90B5EA6F954F497F8AA" bold="true" pageId="103" pageNumber="104">
|
||
<taxonomicNameLabel id="E7EF194AFFF7F90B5EA6F954F497F8AA" pageId="103" pageNumber="104" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="09A803A0FFF7F90B5ADFF8B0F600F8AA" authorityName="Weber" authorityYear="1795" box="[508,607,1796,1820]" class="Malacostraca" family="Raninidae" genus="Symethis" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FCDCA431FFF7F90B5ADFF8B0F600F8AA" box="[508,607,1796,1820]" italics="true" pageId="103" pageNumber="104">Symethis</emphasis>
|
||
</taxonomicName>
|
||
on the basis of a smoother, less ‘eroded’ anterior carapace and in having the cuticle microstructure consisting of large perforations or setal pits rather than fungiform nodes (
|
||
<bibRefCitation id="AA3905D2FFF7F90B5E2AF893F48BF8D5" author="Waugh, D. A. & Feldmann, R. M. & Schweitzer, C. E." pageId="103" pageNumber="104" pagination="15 - 41" refId="ref148784" refString="Waugh, D. A., Feldmann, R. M. & Schweitzer, C. E. (2009) Systematic evaluation of raninid cuticle microstructure. Bulletin of the Mizunami Fossil Museum, 35, 15 - 41." type="journal article" year="2009">
|
||
Waugh
|
||
<emphasis id="FCDCA431FFF7F90B5E41F89CF1C3F889" box="[1378,1436,1831,1855]" italics="true" pageId="103" pageNumber="104">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="09A803A0FFF7F90B5BC5F8F8F5A9F8D5" authority="Van Bakel & Guinot & Artal & Fraaije & Jagt, 2012" authorityName="Van Bakel & Guinot & Artal & Fraaije & Jagt" authorityYear="2012" box="[230,502,1868,1892]" class="Malacostraca" family="Raninidae" genus="Eosymethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="aragonensis" status="sp. nov.">
|
||
<emphasis id="FCDCA431FFF7F90B5BC5F8F8F5A9F8D5" box="[230,502,1868,1892]" italics="true" pageId="103" pageNumber="104">Eosymethis aragonensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FCDCA431FFF7F90B5ADFF8F8F664F8D2" bold="true" box="[508,571,1868,1892]" pageId="103" pageNumber="104">
|
||
<taxonomicNameLabel id="E7EF194AFFF7F90B5ADFF8F8F664F8D2" box="[508,571,1868,1892]" pageId="103" pageNumber="104" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
originates from what appears to have been a fairly muddy-bottom habitat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |