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<mods:title id="53D4CB9E3316C1627186DF79894FD762">A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215</mods:title>
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<paragraph id="CE177823FFC0F93C5BB4FB35F65FFB2C" blockId="80.[151,512,1152,1212]" box="[151,512,1152,1179]" pageId="80" pageNumber="81">
<heading id="955FCF4FFFC0F93C5BB4FB35F65FFB2C" bold="true" box="[151,512,1152,1179]" fontSize="11" level="1" pageId="80" pageNumber="81" reason="1">
<emphasis id="FCDCA431FFC0F93C5BB4FB35F65FFB2C" bold="true" box="[151,512,1152,1179]" pageId="80" pageNumber="81">
<taxonomicName id="09A803A0FFC0F93C5BB4FB35F5E9FB2D" ID-CoL="3789ff54-f8d1-4f52-9cc4-a2fc34fd7011" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[151,438,1153,1179]" class="Malacostraca" family="Lyreididae" genus="Bournelyreidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="teodorii" status="sp. nov.">
<emphasis id="FCDCA431FFC0F93C5BB4FB35F5E9FB2D" bold="true" box="[151,438,1153,1179]" italics="true" pageId="80" pageNumber="81">Bournelyreidus teodorii</emphasis>
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<taxonomicNameLabel id="E7EF194AFFC0F93C5A9EFB34F65FFB2C" box="[445,512,1152,1178]" pageId="80" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="86B22BA8FFC0F93C5BB4FB10F552FB0A" box="[151,269,1188,1212]" pageId="80" pageNumber="81" type="description">
<paragraph id="CE177823FFC0F93C5BB4FB10F552FB0A" blockId="80.[151,512,1152,1212]" box="[151,269,1188,1212]" pageId="80" pageNumber="81">
(
<figureCitation id="569364A6FFC0F93C5BBCFB10F55AFB0A" box="[159,261,1188,1212]" captionStart="FIGURE 24" captionStartId="81.[151,250,1482,1505]" captionTargetBox="[151,1435,339,1458]" captionTargetId="figure-276@81.[151,1435,339,1458]" captionTargetPageId="81" captionText="FIGURE 24. A, Bournelyreidus teodorii n. sp. (Lyreididae, Lyreidinae), MNHN.F A38533 (holotype; ex D. Téodori Collection); middle/upper Maastrichtian, Saint-Loup-en-Comminges (Haute-Garonne, southern France); dorsal view of carapace and chelipeds; B, Lyreidus tridentatus De Haan, 1841 (Lyreididae, Lyreidinae), MAB k. 2883 (male), Recent, Philippines; dorsal view of carapace. Scale bars: A: 10mm; B: 5mm." figureDoi="http://doi.org/10.5281/zenodo.5248760" httpUri="https://zenodo.org/record/5248760/files/figure.png" pageId="80" pageNumber="81">Fig. 24A</figureCitation>
)
</paragraph>
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<subSubSection id="86B22BA8FFC0F93C5BB4FB5FF5CEFA01" pageId="80" pageNumber="81" type="diagnosis">
<paragraph id="CE177823FFC0F93C5BB4FB5FF10BFAD9" blockId="80.[151,1436,1259,2004]" pageId="80" pageNumber="81">
<emphasis id="FCDCA431FFC0F93C5BB4FB5FF54EFAB2" bold="true" box="[151,273,1259,1284]" inLineHeading="true" pageId="80" pageNumber="81" reason="1">Diagnosis.</emphasis>
Carapace large, oval, elongated, widest at one-third of total carapace length from front. Posterolateral margin with divergent posterior portion; anterolateral margin shorter, with 2 small teeth. Orbitofrontal margin tridentate, occupying about half of maximum carapace width. Outer orbital corners prominent, supraorbital borders with 2 orbital notches. Rostrum triangular, bifid, excavated. Carapace surface with minute pits and granules.
</paragraph>
<paragraph id="CE177823FFC0F93C5BE4FACFF5CEFA01" blockId="80.[151,1436,1259,2004]" pageId="80" pageNumber="81">
<emphasis id="FCDCA431FFC0F93C5BE4FACFF5E8FA22" bold="true" box="[199,439,1403,1428]" inLineHeading="true" pageId="80" pageNumber="81" reason="1">Derivation of name.</emphasis>
Named in honour of Mr Dominique Téodori (Pechbonnieu,
<collectingCountry id="B6BF38B3FFC0F93C5F4AFAC8F0E5FA22" box="[1129,1210,1404,1428]" name="France" pageId="80" pageNumber="81">France</collectingCountry>
), who collected and donated the specimen.
</paragraph>
</subSubSection>
<subSubSection id="86B22BA8FFC0F93C5BE4FA77F576F896" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="CE177823FFC0F93C5BE4FA77F576F896" blockId="80.[151,1436,1259,2004]" pageId="80" pageNumber="81">
<emphasis id="FCDCA431FFC0F93C5BE4FA77F5EBFA6A" bold="true" box="[199,436,1475,1500]" inLineHeading="true" pageId="80" pageNumber="81" reason="1">Material examined.</emphasis>
<typeStatus id="1113C681FFC0F93C5A9CFA70F678FA6A" box="[447,551,1476,1500]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
(
<collectionCode id="A8B9E0E6FFC0F93C591AFA70F6CCFA6A" box="[569,659,1476,1500]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="80" pageNumber="81">MNHN</collectionCode>
.F
<accessionNumber id="D1FBE5C0FFC0F93C5991FA70F751FA6A" box="[690,782,1476,1500]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A38533" pageId="80" pageNumber="81" type="EnaNcbi">A38533</accessionNumber>
) collected at working quarry southeast of the village of Saint-Loup-en-Comminges (Haute-Garonne,
<collectingRegion id="0C6CB6C1FFC0F93C59BEFA53F71AFA49" box="[669,837,1511,1535]" country="France" name="Midi-Pyrenees" pageId="80" pageNumber="81">Midi-Pyrénées</collectingRegion>
, southwest
<collectingCountry id="B6BF38B3FFC0F93C58F2FA5CF07EFA49" box="[977,1057,1512,1535]" name="France" pageId="80" pageNumber="81">France</collectingCountry>
;
<date id="BA165EE3FFC0F93C5F12FA53F0AFFA49" box="[1073,1264,1511,1535]" pageId="80" pageNumber="81" value="2009-09">September 2009</date>
), from a bluegrey marly limestone bed at the base of a series assigned to the Marnes bleues de Saint-Loup. A second specimen in the private collection of D. Teodori has also been examined; this retains a better-preserved anterolateral and frontal margin, so that features of the
<typeStatus id="1113C681FFC0F93C591EF9E0F6C0F9DA" box="[573,671,1620,1644]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
could be checked and confirmed. Co-occurring macrofaunal assemblages are rich in holasteroid echinoids, ostreid bivalves, fishes, cephalopods (e.g., the scaphitid ammonite
<taxonomicName id="09A803A0FFC0F93C5E41F9CDF607F902" authority="(Sowerby, 1817)" baseAuthorityName="Sowerby" baseAuthorityYear="1817" class="Cephalopoda" family="Scaphitidae" genus="Hoploscaphites" kingdom="Animalia" order="Ammonoidea" pageId="80" pageNumber="81" phylum="Mollusca" rank="species" species="constrictus">
<emphasis id="FCDCA431FFC0F93C5E41F9CDF5C8F902" italics="true" pageId="80" pageNumber="81">Hoploscaphites constrictus</emphasis>
(
<bibRefCitation id="AA3905D2FFC0F93C5A86F928F60FF902" author="Sowerby, J." box="[421,592,1692,1716]" pageId="80" pageNumber="81" refId="ref144726" refString="Sowerby, J. (1817) The Mineral Conchology of Great Britain, 2, pls. 151 - 184, A, 185 - 186. The author, London." type="book" year="1817">Sowerby, 1817</bibRefCitation>
)
</taxonomicName>
and other unidentified crustaceans. The thickness of such limestone levels varies between 0.1 and 0.3 m; they alternate with softer marls. As noted by
<bibRefCitation id="AA3905D2FFC0F93C58CFF974F17BF96E" author="Kennedy, W. J. &amp; Bilotte, M. &amp; Lepicard, B. &amp; Segura, F." box="[1004,1316,1728,1752]" pageId="80" pageNumber="81" pagination="1001 - 1037" refId="ref133417" refString="Kennedy, W. J., Bilotte, M., Lepicard, B. &amp; Segura, F. (1986) Upper Campanian and Maastrichtian ammonites from the Petites- Pyrenees, southern France. Eclogae geologicae Helvetiae, 79, 1001 - 1037." type="journal article" year="1986">
Kennedy
<emphasis id="FCDCA431FFC0F93C5F7BF975F0D5F96E" box="[1112,1162,1728,1752]" italics="true" pageId="80" pageNumber="81">et al</emphasis>
. (1986: 1004
</bibRefCitation>
, 1005, fig. 2) and
<bibRefCitation id="AA3905D2FFC0F93C5BC6F950F5B3F94A" author="Peybernes, B. &amp; Fondecave-Wallez, M. J. &amp; Eichene, P. &amp; Robin, E. &amp; Rocchia, R." box="[229,492,1764,1788]" pageId="80" pageNumber="81" pagination="647 - 653" refId="ref140349" refString="Peybernes, B., Fondecave-Wallez, M. J., Eichene, P., Robin, E. &amp; Rocchia, R. (1998) La limite Cretace-Tertiaire en domaine marin dans les Pyrenees centrales (zone sous-pyreneenne, France). Comptes rendus de l'Academie des Sciences (Paris), 326, 647 - 653." type="journal article" year="1998">
Peybernès
<emphasis id="FCDCA431FFC0F93C5A43F951F5CEF94A" box="[352,401,1764,1788]" italics="true" pageId="80" pageNumber="81">et al</emphasis>
. (1998)
</bibRefCitation>
, the Marnes bleues de St. Loup in the Blajan-Bazordan anticline are of late Maastrichtian age.
</paragraph>
</subSubSection>
<subSubSection id="86B22BA8FFC0F93D5BE4F89FF640FEAA" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81" type="description">
<paragraph id="CE177823FFC0F93D5BE4F89FF640FEAA" blockId="80.[151,1436,1259,2004]" lastBlockId="81.[151,1436,151,284]" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
<emphasis id="FCDCA431FFC0F93C5BE4F89FF507F8F2" bold="true" box="[199,344,1835,1860]" inLineHeading="true" pageId="80" pageNumber="81" reason="1">Description.</emphasis>
Carapace large (maximum length
<quantity id="0950D5C6FFC0F93C59FCF898F76EF8F5" box="[735,817,1836,1860]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="80" pageNumber="81" unit="mm" value="66.0">66 mm</quantity>
, width
<quantity id="0950D5C6FFC0F93C58A6F898F786F8F2" box="[901,985,1836,1860]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="80" pageNumber="81" unit="mm" value="40.0">40 mm</quantity>
), oval, elongated, widest behind the lateral spines at about at one-third of total carapace length from front. Orbitofrontal margin tridentate, occupying about half of maximum carapace width. Base of rostrum about one-third of orbitofrontal region. Rostrum triangular, bifid, excavated. Large subcircular orbits with prominent outer orbital spines. Supraorbital borders with 2 relatively deep orbital fissures. Posterolateral margin with diverging posterior portion; anterolateral margin shorter, with 2 rather small, forwardly directed spines. Branchiocardiac grooves, impressions of epimeral adductor muscle scars faint. Carapace surface with minute pits, granules. Granules significantly coarser along lateral margins. Both chelipeds similar in size, form; merus, carpus, propodus of comparable length, width, covered with granules; propodus distinctly more flattened.
</paragraph>
</subSubSection>
<caption id="9AD728ABFFC1F93D5BB4FA7EF6D4F98A" ID-DOI="http://doi.org/10.5281/zenodo.5248760" ID-Zenodo-Dep="5248760" httpUri="https://zenodo.org/record/5248760/files/figure.png" pageId="81" pageNumber="82" startId="81.[151,250,1482,1505]" targetBox="[151,1435,339,1458]" targetPageId="81">
<paragraph id="CE177823FFC1F93D5BB4FA7EF6D4F98A" blockId="81.[151,1436,1482,1596]" pageId="81" pageNumber="82">
<emphasis id="FCDCA431FFC1F93D5BB4FA7EF57EFA57" bold="true" box="[151,289,1482,1505]" pageId="81" pageNumber="82">FIGURE 24.</emphasis>
A,
<taxonomicName id="09A803A0FFC1F93D5A64FA7EF665FA56" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[327,570,1482,1504]" class="Malacostraca" family="Lyreididae" genus="Bournelyreidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="teodorii" status="sp. nov.">
<emphasis id="FCDCA431FFC1F93D5A64FA7EF665FA56" box="[327,570,1482,1504]" italics="true" pageId="81" pageNumber="82">Bournelyreidus teodorii</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D5962FA7FF624FA56" bold="true" box="[577,635,1483,1504]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D5962FA7FF624FA56" box="[577,635,1483,1504]" pageId="81" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="09A803A0FFC1F93D59AAFA7EF6A5FA56" authorityName="Guinot" authorityYear="1993" box="[649,762,1482,1504]" class="Malacostraca" family="Lyreididae" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="family">Lyreididae</taxonomicName>
,
<taxonomicName id="09A803A0FFC1F93D5827FA7EF72AFA56" authorityName="Guinot, 1993 b:" authorityYear="1326" box="[772,885,1482,1504]" class="Malacostraca" family="Lyreididae" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="subFamily" subFamily="Lyreidinae">Lyreidinae</taxonomicName>
), MNHN.F
<accessionNumber id="D1FBE5C0FFC1F93D58D3FA7EF01EFA56" box="[1008,1089,1482,1504]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A38533" pageId="81" pageNumber="82" type="EnaNcbi">A38533</accessionNumber>
(
<emphasis id="FCDCA431FFC1F93D5F72FA7FF0F1FA56" bold="true" box="[1105,1198,1483,1504]" pageId="81" pageNumber="82">holotype</emphasis>
;
<emphasis id="FCDCA431FFC1F93D5F9FFA7FF08CFA56" box="[1212,1235,1483,1504]" italics="true" pageId="81" pageNumber="82">ex</emphasis>
D. Téodori Collection); middle/upper Maastrichtian, Saint-Loup-en-Comminges (Haute-Garonne, southern France); dorsal view of carapace and chelipeds; B,
<taxonomicName id="09A803A0FFC1F93D5A07F9BCF6CAF9A8" authority="De Haan, 1841" authorityName="De Haan" authorityYear="1841" box="[292,661,1544,1566]" class="Malacostraca" family="Lyreididae" genus="Lyreidus" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tridentatus">
<emphasis id="FCDCA431FFC1F93D5A07F9BCF5AFF9A8" box="[292,496,1544,1566]" italics="true" pageId="81" pageNumber="82">Lyreidus tridentatus</emphasis>
De Haan, 1841
</taxonomicName>
(
<taxonomicName id="09A803A0FFC1F93D5980F9BCF74BF9A8" authorityName="Guinot" authorityYear="1993" box="[675,788,1544,1566]" class="Malacostraca" family="Lyreididae" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="family">Lyreididae</taxonomicName>
,
<taxonomicName id="09A803A0FFC1F93D583CF9BCF7CFF9A8" authorityName="Guinot, 1993 b:" authorityYear="1326" box="[799,912,1544,1566]" class="Malacostraca" family="Lyreididae" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="subFamily" subFamily="Lyreidinae">Lyreidinae</taxonomicName>
), MAB k. 2883 (male), Recent, Philippines; dorsal view of carapace. Scale bars: A: 10mm; B: 5mm.
</paragraph>
</caption>
<subSubSection id="86B22BA8FFC1F93D5BE4F939F0D3F873" pageId="81" pageNumber="82" type="discussion">
<paragraph id="CE177823FFC1F93D5BE4F939F0D3F873" blockId="81.[151,1436,1677,1989]" pageId="81" pageNumber="82">
<emphasis id="FCDCA431FFC1F93D5BE4F939F564F913" bold="true" box="[199,315,1677,1701]" inLineHeading="true" pageId="81" pageNumber="82" reason="1">Remarks.</emphasis>
Despite the fact that the carapace has suffered from sediment compaction, the morphology of the orbitofrontal and anterolateral margins allows assignment of this form to
<taxonomicName id="09A803A0FFC1F93D58EFF905F028F97F" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[972,1143,1713,1737]" class="Malacostraca" family="Lyreididae" genus="Bournelyreidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FCDCA431FFC1F93D58EFF905F028F97F" box="[972,1143,1713,1737]" italics="true" pageId="81" pageNumber="82">Bournelyreidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7EF194AFFC1F93D5F5CF905F092F97F" box="[1151,1229,1713,1737]" pageId="81" pageNumber="82" rank="genus">
<emphasis id="FCDCA431FFC1F93D5F5CF905F099F97F" bold="true" box="[1151,1222,1713,1737]" pageId="81" pageNumber="82">n. gen</emphasis>
.
</taxonomicNameLabel>
It is distinguished from
<taxonomicName id="09A803A0FFC1F93D5BF5F962F534F95B" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[214,363,1749,1773]" class="Malacostraca" family="Raninidae" genus="Raninella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="carlilensis" status="comb. nov.">
<emphasis id="FCDCA431FFC1F93D5BF5F962F534F95B" box="[214,363,1749,1773]" italics="true" pageId="81" pageNumber="82">B. carlilensis</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D5A57F961F584F95B" bold="true" box="[372,475,1749,1773]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D5A57F961F584F95B" box="[372,475,1749,1773]" pageId="81" pageNumber="82" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="FCDCA431FFC1F93D5936F962F671F95B" box="[533,558,1750,1773]" italics="true" pageId="81" pageNumber="82">B.</emphasis>
?
<taxonomicName id="09A803A0FFC1F93D5960F961F6EEF95B" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" baseAuthorityName="Bishop" baseAuthorityYear="1978" box="[579,689,1749,1773]" class="Malacostraca" family="Raninidae" genus="Raninella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="oaheensis" status="comb. nov.">
<emphasis id="FCDCA431FFC1F93D5960F961F6EEF95B" box="[579,689,1749,1773]" italics="true" pageId="81" pageNumber="82">oaheensis</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D5999F961F77EF95B" bold="true" box="[698,801,1749,1773]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D5999F961F77EF95B" box="[698,801,1749,1773]" pageId="81" pageNumber="82" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
in attaining a markedly larger size and in having a less broad orbitofrontal region and a much larger carpus.
<taxonomicName id="09A803A0FFC1F93D59CAF94DF7B5F8A7" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" baseAuthorityName="Roberts" baseAuthorityYear="1962" box="[745,1002,1785,1809]" class="Malacostraca" family="Lyreididae" genus="Bournelyreidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tridens" status="comb. nov.">
<emphasis id="FCDCA431FFC1F93D59CAF94DF7B5F8A7" box="[745,1002,1785,1809]" italics="true" pageId="81" pageNumber="82">Bournelyreidus tridens</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D58D2F94DF008F8A7" bold="true" box="[1009,1111,1785,1809]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D58D2F94DF008F8A7" box="[1009,1111,1785,1809]" pageId="81" pageNumber="82" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
has a much shorter anterolateral margin, and a more projected orbitofrontal region; its posterolateral margins are more concave and less divergent than the new species.
<taxonomicName id="09A803A0FFC1F93D5AE2F8F5F6B6F8EF" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[449,745,1857,1881]" class="Malacostraca" family="Lyreididae" genus="Bourneylyreidus" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="manningi" status="comb. nov.">
<emphasis id="FCDCA431FFC1F93D5AE2F8F5F6B6F8EF" box="[449,745,1857,1881]" italics="true" pageId="81" pageNumber="82">Bourneylyreidus manningi</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D59D3F8F5F709F8EF" bold="true" box="[752,854,1857,1881]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D59D3F8F5F709F8EF" box="[752,854,1857,1881]" pageId="81" pageNumber="82" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
has much coarser ornamentation and a more clearly areolated carapace. The new species differs from
<taxonomicName id="09A803A0FFC1F93D59EEF8D2F770F8CB" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[717,815,1893,1917]" class="Malacostraca" family="Lyreididae" genus="Palaeocorystes" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="laevis" status="comb. nov.">
<emphasis id="FCDCA431FFC1F93D59EEF8D2F770F8CB" box="[717,815,1893,1917]" italics="true" pageId="81" pageNumber="82">B. laevis</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D581BF8D1F7FFF8CB" bold="true" box="[824,928,1893,1917]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D581BF8D1F7FFF8CB" box="[824,928,1893,1917]" pageId="81" pageNumber="82" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
in having a broader base to the rostrum and subcircular orbits rather than near-rectangular ones; it can also be distinguished from
<taxonomicName id="09A803A0FFC1F93D5F78F83EF155F817" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[1115,1290,1930,1953]" class="Malacostraca" family="Raninidae" genus="Hemioon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="eysunesensis" status="comb. nov.">
<emphasis id="FCDCA431FFC1F93D5F78F83EF155F817" box="[1115,1290,1930,1953]" italics="true" pageId="81" pageNumber="82">B. eysunesensis</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FFC1F93D5E31F83DF127F817" bold="true" box="[1298,1400,1929,1953]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E7EF194AFFC1F93D5E31F83DF127F817" box="[1298,1400,1929,1953]" pageId="81" pageNumber="82" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
by its less concave anterolateral margin between the outer orbital and first anterolateral spine.
</paragraph>
</subSubSection>
</treatment>
</document>