256 lines
34 KiB
XML
256 lines
34 KiB
XML
<document id="D149EC917949F6B370535E0832A77E85" ID-DOI="10.11646/zootaxa.3215.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5248640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629765289367" checkinUser="felipe" docAuthor="Van Bakel, Barry W. M., Guinot, Danièle, Artal, Pedro, Fraaije, René H. B. & Jagt, John W. M." docDate="2012" docId="4601C935FFA7F9545BB4FCAAF07FFD39" docLanguage="en" docName="zt03215p216.pdf" docOrigin="Zootaxa 3215 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3215.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Cenomanocarcinus vanstraeleni Stenzel 1945" docType="treatment" docVersion="6" lastPageNumber="57" masterDocId="BA38B14DFF90F96C5B23FFB4F45FFFB6" masterDocTitle="A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215" masterLastPageNumber="216" masterPageNumber="1" pageNumber="56" updateTime="1699109082209" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A137D87E901C483FC76DB5C35469DF30">A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215</mods:title>
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<mods:namePart id="575EDD50D90341BE603AEF8A7761154F">Van Bakel, Barry W. M.</mods:namePart>
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<mods:namePart id="11F773049850C9F1FA4E98D5A7639649">Guinot, Danièle</mods:namePart>
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<mods:namePart id="1084B0F1F61A6A05844017F89C495F2D">Artal, Pedro</mods:namePart>
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<mods:namePart id="C35688EAA00F9EE228D17F1DCC74C54C">Fraaije, René H. B.</mods:namePart>
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<mods:namePart id="1913C700FB1253BBAB138626DE01C7E3">Jagt, John W. M.</mods:namePart>
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<mods:date id="AB552ADBCF8B3F1182BC11FE0B272EA3">2012</mods:date>
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<mods:number id="0BD9A6846858480882871A4B56DAA243">2012-02-29</mods:number>
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<mods:number id="9F1D6AF94CC9A54F36B417B38CE40FC7">3215</mods:number>
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<treatment id="4601C935FFA7F9545BB4FCAAF07FFD39" ID-DOI="http://doi.org/10.5281/zenodo.5250366" ID-Zenodo-Dep="5250366" LSID="urn:lsid:plazi:treatment:4601C935FFA7F9545BB4FCAAF07FFD39" httpUri="http://treatment.plazi.org/id/4601C935FFA7F9545BB4FCAAF07FFD39" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
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<subSubSection id="86B22BA8FFA7F95B5BB4FCAAF6F1FC81" box="[151,686,798,823]" pageId="55" pageNumber="56" type="nomenclature">
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<paragraph id="CE177823FFA7F95B5BB4FCAAF6F1FC81" blockId="55.[151,686,798,858]" box="[151,686,798,823]" pageId="55" pageNumber="56">
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<heading id="955FCF4FFFA7F95B5BB4FCAAF6F1FC81" bold="true" box="[151,686,798,823]" fontSize="10" level="2" pageId="55" pageNumber="56" reason="2">
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<taxonomicName id="09A803A0FFA7F95B5BB4FCAAF6F1FC81" ID-CoL="52f8103e-6bf8-4ed9-b13f-40d38e9a6d07" authority="Stenzel, 1945" authorityName="Stenzel" authorityYear="1945" box="[151,686,798,823]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">
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<emphasis id="FCDCA431FFA7F95B5BB4FCAAF6F1FC81" bold="true" box="[151,686,798,823]" pageId="55" pageNumber="56">
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<emphasis id="FCDCA431FFA7F95B5BB4FCAAF653FC81" bold="true" box="[151,524,798,823]" italics="true" pageId="55" pageNumber="56">Cenomanocarcinus vanstraeleni</emphasis>
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<bibRefCitation id="AA3905D2FFA7F95B5937FCAAF6F1FC81" author="Stenzel, H. B." box="[532,686,798,823]" pageId="55" pageNumber="56" pagination="401 - 476" refId="ref145060" refString="Stenzel, H. B. (1945) Decapod crustaceans from the Cretaceous of Texas. The University of Texas Publication, 4401, 401 - 476, pls. 34 - 45." type="journal article" year="1945">Stenzel, 1945</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="86B22BA8FFA7F95B5BB4FCF6F514FCEC" box="[151,331,834,858]" pageId="55" pageNumber="56" type="description">
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<paragraph id="CE177823FFA7F95B5BB4FCF6F514FCEC" blockId="55.[151,686,798,858]" box="[151,331,834,858]" pageId="55" pageNumber="56">
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(
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<figureCitation id="569364A6FFA7F95B5BBCFCF6F51CFCEC" box="[159,323,834,858]" captionStart="FIGURE 18" captionStartId="52.[151,250,1789,1812]" captionTargetBox="[151,1436,193,1768]" captionTargetId="figure-154@52.[151,1436,193,1768]" captionTargetPageId="52" captionText="FIGURE 18. A, D. Cenomanocarcinus inflatus Van Straelen, 1936 (Cenomanocarcinidae), MNHN J08587 (lectotype), photograph of MAB k. 2968 (cast), upper Cenomanian, Butte de Gazonfier, Le Mans (northwestern France); dorsal and left lateral views (direction of photography in D indicated by arrow in A); B, C, E, Cenomanocarcinus vanstraeleni Stenzel, 1945, MAB k. 2909 (B, C; indeterminate sex), upper Albian-lower Cenomanian (Pawpaw Formation), Tarrant County, Texas (U.S.A); dorsal and left lateral views (direction of photography in C indicated by arrow in B); MAB k. 2885 (E; indeterminate sex), dorsal view of carapace; F, Cenomanocarcinus beardi Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross, 2003a, MAB k. 2880 (indeterminate sex), Cenomanian (Woodbine Formation), Wagner Park, Texas (U.S.A), dorsal view of carapace. ebs, epibranchial spine; pbs, posterobranchial spine. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.5248664" httpUri="https://zenodo.org/record/5248664/files/figure.png" pageId="55" pageNumber="56">Fig. 18B, C, E</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="86B22BA8FFA7F95B5BB4FC3EF5C0FBB8" pageId="55" pageNumber="56" type="materials_examined">
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<paragraph id="CE177823FFA7F95B5BB4FC3EF5C0FBB8" blockId="55.[151,1436,906,2011]" pageId="55" pageNumber="56">
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<emphasis id="FCDCA431FFA7F95B5BB4FC3EF5DCFC15" bold="true" box="[151,387,906,931]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Material examined.</emphasis>
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<collectionCode id="A8B9E0E6FFA7F95B5AAFFC3EF593FC14" box="[396,460,906,930]" pageId="55" pageNumber="56">MAB</collectionCode>
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k. 2885 (indeterminate sex), carapace,
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<collectionCode id="A8B9E0E6FFA7F95B58B7FC3EF78BFC14" box="[916,980,906,930]" pageId="55" pageNumber="56">MAB</collectionCode>
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k. 2895 (female), partial carapace with remains of abdomen, thoracic sternum and base of mxp3,
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<collectionCode id="A8B9E0E6FFA7F95B5801FC1AF73DFC70" box="[802,866,942,966]" pageId="55" pageNumber="56">MAB</collectionCode>
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k. 2909, (indeterminate sex), partial carapace with remains of P1 and thoracic sternum; all from the upper Albian‒lower Cenomanian (Pawpaw Formation), Tarrant County (
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<collectingRegion id="0C6CB6C1FFA7F95B5BDBFC42F562FBB8" box="[248,317,1014,1038]" country="United States of America" name="Texas" pageId="55" pageNumber="56">Texas</collectingRegion>
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,
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<collectingCountry id="B6BF38B3FFA7F95B5A6BFC42F526FBB8" box="[328,377,1014,1038]" name="United States of America" pageId="55" pageNumber="56">U.S.</collectingCountry>
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A).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="86B22BA8FFA7F9545BE4FBAFF07FFD39" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="discussion">
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<paragraph id="CE177823FFA7F95B5BE4FBAFF6C9FB50" blockId="55.[151,1436,906,2011]" pageId="55" pageNumber="56">
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<emphasis id="FCDCA431FFA7F95B5BE4FBAFF564FB85" bold="true" box="[199,315,1051,1075]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Remarks.</emphasis>
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The morphology of this species has been described by
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<bibRefCitation id="AA3905D2FFA7F95B5886FBAEF0DCFB84" author="Stenzel, H. B." box="[933,1155,1050,1074]" pageId="55" pageNumber="56" pagination="401 - 476" refId="ref145060" refString="Stenzel, H. B. (1945) Decapod crustaceans from the Cretaceous of Texas. The University of Texas Publication, 4401, 401 - 476, pls. 34 - 45." type="journal article" year="1945">Stenzel (1945: 447)</bibRefCitation>
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,
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<bibRefCitation id="AA3905D2FFA7F95B5FACFBAFF1C3FB84" author="Vega, F. J. & Nyborg, T. & Patarroyo, P. & Luque, J. & Stinnesbeck, W." box="[1167,1436,1050,1075]" pageId="55" pageNumber="56" pagination="403 - 422" refId="ref148139" refString="Vega, F. J., Nyborg, T., Rojas-Briceno, Patarroyo, P., Luque, J., Porras-Muzquiz & Stinnesbeck, W. (2007 a) Upper Cretaceous Crustacea from Mexico and Colombia: similar faunas and environments during Turonian times. Revista Mexicana de Ciencias Geologicas, 24, 403 - 422." type="journal article" year="2007">
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Vega
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<emphasis id="FCDCA431FFA7F95B5FEEFBAFF0A1FB85" box="[1229,1278,1051,1075]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
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. (2007a: 414)
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</bibRefCitation>
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and
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<bibRefCitation id="AA3905D2FFA7F95B5BE4FB8AF589FBE0" author="Guinot, D. & Vega, F. J. & Van Bakel, B. W. M." box="[199,470,1086,1110]" pageId="55" pageNumber="56" pagination="681 - 719" refId="ref130419" refString="Guinot, D., Vega, F. J. & Van Bakel, B. W. M. (2008) Cenomanocarcinidae n. fam., a new Cretaceous podotreme family (Crustacea, Brachyura, Raninoidia), with comments on related families. Geodiversitas, 30, 681 - 719." type="journal article" year="2008">
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Guinot
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<emphasis id="FCDCA431FFA7F95B5A3FFB8BF510FBE0" box="[284,335,1086,1110]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
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. (2008: 14)
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</bibRefCitation>
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. A large specimen of
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<taxonomicName id="09A803A0FFA7F95B59EDFB8AF7F8FBE0" authorityName="Stenzel" authorityYear="1945" box="[718,935,1086,1110]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">
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<emphasis id="FCDCA431FFA7F95B59EDFB8AF6B7FBE0" box="[718,744,1086,1110]" italics="true" pageId="55" pageNumber="56">C.</emphasis>
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aff.
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<emphasis id="FCDCA431FFA7F95B583EFB8AF7F8FBE0" box="[797,935,1086,1110]" italics="true" pageId="55" pageNumber="56">vanstraeleni</emphasis>
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</taxonomicName>
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was described (
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<bibRefCitation id="AA3905D2FFA7F95B5F7EFB8AF106FBE0" author="Guinot, D. & Vega, F. J. & Van Bakel, B. W. M." box="[1117,1369,1086,1110]" pageId="55" pageNumber="56" pagination="681 - 719" refId="ref130419" refString="Guinot, D., Vega, F. J. & Van Bakel, B. W. M. (2008) Cenomanocarcinidae n. fam., a new Cretaceous podotreme family (Crustacea, Brachyura, Raninoidia), with comments on related families. Geodiversitas, 30, 681 - 719." type="journal article" year="2008">
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||
Guinot
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<emphasis id="FCDCA431FFA7F95B5F96FB8BF0B8FBE0" box="[1205,1255,1086,1110]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
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||
. 2008: 18
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</bibRefCitation>
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||
) with episternite 5 showing ‘a pair of prominences that is blunt, weathered, but well-preserved and recognisable on left side (crab seen by ventral view). This pair of prominences is assumed to be a part of the abdominal holding system’. Two new specimens, smaller in size but with well-preserved but partial thoracic sterna, retain such prominences and allow a more detailed description.
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</paragraph>
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<paragraph id="CE177823FFA7F95B5BE4FB46F586FA54" blockId="55.[151,1436,906,2011]" pageId="55" pageNumber="56">
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One specimen (MAB k. 2909;
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<figureCitation id="569364A6FFA7F95B5902FB46F6FCFABC" box="[545,675,1266,1290]" captionStart="FIGURE 40" captionStartId="120.[151,250,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure-159@120.[151,1436,193,1771]" captionTargetPageId="120" captionText="FIGURE 40. Double peg in Palaeocorystoidea; A, B, Silvacarcinus laurae Collins & Smith, 1993 (Orithopsidae), IRScNB TCCI 6115 (holotype), lower Eocene (Ypresian), Brussels (Belgium); ventral view showing thoracic sternum, and detail of thoracic sternum; C–F. Cenomanocarcinus vanstraeleni Stenzel, 1945 (Cenomanocarcinidae), MAB k. 2909 (C, E), upper Albianlower Cenomanian (Pawpaw Formation), Tarrant County, Texas (U.S.A); details of thoracic sternum; MAB k. 2895 (D, F); same provenance, detail of abdomen and thoracic sternum, and ventral view showing thoracic sternum, abdomen and base of mxp3. 4, 5, 6, thoracic sternites 4, 5, 6; 4/5, 5/6, thoracic sternal sutures 4/5, 5/6; a6, abdominal somite 6; cmxp3, coxa of mxp3; cx2, P2 coxa; dp, distal peg; e4, e5, episternites 4, 5; g2, gynglyme for P2 coxa; p, peg (undifferentiated); sp, subdistal peg; t, telson. Scale bars: A-E: 5mm; F: 10mm." figureDoi="http://doi.org/10.5281/zenodo.5248698" httpUri="https://zenodo.org/record/5248698/files/figure.png" pageId="55" pageNumber="56">Fig. 40C, E</figureCitation>
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) is an incomplete carapace with well-preserved proximal portion of mxp3, thoracic sternites 3‒5, P1 and P2 bases, abdominal somite 6 and telson. The relatively wide, flat thoracic sternum and wide abdomen suggest a female. Abdominal somite 6 and the telson are disarticulated and posteriorly displaced, exposing episternites 5. On either side, near the P2 gynglyme, a prominent double peg is present. The structure is raised, the distal (posterior) peg being slightly larger and slightly more admedially positioned than the subdistal (anterior) one. The anterior corners of abdominal somite 6 are rounded and swollen, which suggests a socket was present ventrally.
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</paragraph>
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<paragraph id="CE177823FFA7F95B5BE4FA5AF129F9C4" blockId="55.[151,1436,906,2011]" pageId="55" pageNumber="56">
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The second specimen (MAB k. 2895;
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<figureCitation id="569364A6FFA7F95B5951FA5AF6AAF9B0" box="[626,757,1518,1542]" captionStart="FIGURE 40" captionStartId="120.[151,250,1792,1815]" captionTargetBox="[151,1436,193,1771]" captionTargetId="figure-159@120.[151,1436,193,1771]" captionTargetPageId="120" captionText="FIGURE 40. Double peg in Palaeocorystoidea; A, B, Silvacarcinus laurae Collins & Smith, 1993 (Orithopsidae), IRScNB TCCI 6115 (holotype), lower Eocene (Ypresian), Brussels (Belgium); ventral view showing thoracic sternum, and detail of thoracic sternum; C–F. Cenomanocarcinus vanstraeleni Stenzel, 1945 (Cenomanocarcinidae), MAB k. 2909 (C, E), upper Albianlower Cenomanian (Pawpaw Formation), Tarrant County, Texas (U.S.A); details of thoracic sternum; MAB k. 2895 (D, F); same provenance, detail of abdomen and thoracic sternum, and ventral view showing thoracic sternum, abdomen and base of mxp3. 4, 5, 6, thoracic sternites 4, 5, 6; 4/5, 5/6, thoracic sternal sutures 4/5, 5/6; a6, abdominal somite 6; cmxp3, coxa of mxp3; cx2, P2 coxa; dp, distal peg; e4, e5, episternites 4, 5; g2, gynglyme for P2 coxa; p, peg (undifferentiated); sp, subdistal peg; t, telson. Scale bars: A-E: 5mm; F: 10mm." figureDoi="http://doi.org/10.5281/zenodo.5248698" httpUri="https://zenodo.org/record/5248698/files/figure.png" pageId="55" pageNumber="56">Fig. 40D, F</figureCitation>
|
||
), a left side of a well-preserved carapace, retaining elements of P1 and P2 and a small portion of the thoracic sternum. Sternites 4 and 5 are partially preserved, episternites 4 and 5 complete. Episternite 5 shows well the P2 gynglyme in ventral view. A well-preserved double peg, separated by a weak depression is present, the distal peg being larger, more posterolaterally placed than the subdistal one.
|
||
</paragraph>
|
||
<paragraph id="CE177823FFA7F95B5BE4F9CBF1C9F90D" blockId="55.[151,1436,906,2011]" pageId="55" pageNumber="56">In another, rather small, specimen (MAB k. 2922), it is seen that the blunt crest on the pterygostome becomes more acute anteriorly, terminating below the orbits in a prominent, pointed subantennary lobe of the pterygostome.</paragraph>
|
||
<paragraph id="CE177823FFA7F9545BE4F972F726FF41" blockId="55.[151,1436,906,2011]" lastBlockId="56.[151,1436,151,572]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
|
||
A comparison of the available material permits better differentiation of
|
||
<emphasis id="FCDCA431FFA7F95B58C4F972F4E6F8B4" italics="true" pageId="55" pageNumber="56">
|
||
<taxonomicName id="09A803A0FFA7F95B58C4F972F00CF968" authorityName="Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross" authorityYear="2003" box="[999,1107,1734,1758]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="beardi">C. beardi</taxonomicName>
|
||
,
|
||
<taxonomicName id="09A803A0FFA7F95B5F7DF972F08DF968" authorityName="Van Straelen" authorityYear="1936" box="[1118,1234,1734,1758]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">C. inflatus</taxonomicName>
|
||
and
|
||
<taxonomicName id="09A803A0FFA7F95B5E29F972F4E6F8B4" authorityName="Stenzel" authorityYear="1945" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">C. vanstraeleni</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="09A803A0FFA7F95B5BEAF95FF5A1F8B5" authorityName="Van Straelen" authorityYear="1936" box="[201,510,1771,1795]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA7F95B5BEAF95FF5A1F8B5" box="[201,510,1771,1795]" italics="true" pageId="55" pageNumber="56">Cenomanocarcinus inflatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="09A803A0FFA7F95B591BF95FF6BBF8B5" authorityName="Stenzel" authorityYear="1945" box="[568,740,1771,1795]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">
|
||
<emphasis id="FCDCA431FFA7F95B591BF95FF6BBF8B5" box="[568,740,1771,1795]" italics="true" pageId="55" pageNumber="56">C. vanstraeleni</emphasis>
|
||
</taxonomicName>
|
||
are closely similar, but both species are highly variable with regard to tuberculation and granulation. Both possess an anterolateral margin with five or six teeth anterior to the epibranchial spine. The cervical groove notches the anterolateral margin more strongly and acutely in
|
||
<taxonomicName id="09A803A0FFA7F95B5E29F887F4E6F8D8" authorityName="Stenzel" authorityYear="1945" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">
|
||
<emphasis id="FCDCA431FFA7F95B5E29F887F4E6F8D8" italics="true" pageId="55" pageNumber="56">C. vanstraeleni</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="09A803A0FFA7F95B5A34F8E2F5D3F8D8" authorityName="Van Straelen" authorityYear="1936" box="[279,396,1878,1902]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA7F95B5A34F8E2F5D3F8D8" box="[279,396,1878,1902]" italics="true" pageId="55" pageNumber="56">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
. The nature of the epibranchial spine differs between the two species, being strongly developed in
|
||
<taxonomicName id="09A803A0FFA7F95B5BD3F8CFF53BF825" authorityName="Van Straelen" authorityYear="1936" box="[240,356,1915,1939]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA7F95B5BD3F8CFF53BF825" box="[240,356,1915,1939]" italics="true" pageId="55" pageNumber="56">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
, but similar to or only marginally better developed than the anterolateral teeth in
|
||
<taxonomicName id="09A803A0FFA7F95B5FC9F8CFF1CBF825" authorityName="Stenzel" authorityYear="1945" box="[1258,1428,1915,1939]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">
|
||
<emphasis id="FCDCA431FFA7F95B5FC9F8CFF1CBF825" box="[1258,1428,1915,1939]" italics="true" pageId="55" pageNumber="56">C. vanstraeleni</emphasis>
|
||
</taxonomicName>
|
||
. There are also differences in the posterolateral margins: a conspicuously strong spiniform tooth halfway the otherwise smooth and rounded posterolateral margin in
|
||
<taxonomicName id="09A803A0FFA7F95B59F1F877F717F86D" authorityName="Van Straelen" authorityYear="1936" box="[722,840,1987,2011]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA7F95B59F1F877F717F86D" box="[722,840,1987,2011]" italics="true" pageId="55" pageNumber="56">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<taxonomicName id="09A803A0FFA7F95B589AF877F03AF86D" authorityName="Stenzel" authorityYear="1945" box="[953,1125,1987,2011]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="vanstraeleni">
|
||
<emphasis id="FCDCA431FFA7F95B589AF877F03AF86D" box="[953,1125,1987,2011]" italics="true" pageId="55" pageNumber="56">C. vanstraeleni</emphasis>
|
||
</taxonomicName>
|
||
has a posterolateral margin which is sharper in cross section and bears many small spinules, of which one may be slightly larger, but by no means similar to the spine of
|
||
<taxonomicName id="09A803A0FFA8F9545AC6FF08F604FF62" authorityName="Van Straelen" authorityYear="1936" box="[485,603,188,212]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA8F9545AC6FF08F604FF62" box="[485,603,188,212]" italics="true" pageId="56" pageNumber="57">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
. The most obvious difference between the two species is the width of the cardiac region, which is much greater in
|
||
<taxonomicName id="09A803A0FFA8F9545943FF54F68BFF4E" authorityName="Van Straelen" authorityYear="1936" box="[608,724,224,248]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA8F9545943FF54F68BFF4E" box="[608,724,224,248]" italics="true" pageId="56" pageNumber="57">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="569364A6FFA8F95459C7FF6BF734FF41" box="[740,875,223,247]" captionStart="FIGURE 18" captionStartId="52.[151,250,1789,1812]" captionTargetBox="[151,1436,193,1768]" captionTargetId="figure-154@52.[151,1436,193,1768]" captionTargetPageId="52" captionText="FIGURE 18. A, D. Cenomanocarcinus inflatus Van Straelen, 1936 (Cenomanocarcinidae), MNHN J08587 (lectotype), photograph of MAB k. 2968 (cast), upper Cenomanian, Butte de Gazonfier, Le Mans (northwestern France); dorsal and left lateral views (direction of photography in D indicated by arrow in A); B, C, E, Cenomanocarcinus vanstraeleni Stenzel, 1945, MAB k. 2909 (B, C; indeterminate sex), upper Albian-lower Cenomanian (Pawpaw Formation), Tarrant County, Texas (U.S.A); dorsal and left lateral views (direction of photography in C indicated by arrow in B); MAB k. 2885 (E; indeterminate sex), dorsal view of carapace; F, Cenomanocarcinus beardi Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross, 2003a, MAB k. 2880 (indeterminate sex), Cenomanian (Woodbine Formation), Wagner Park, Texas (U.S.A), dorsal view of carapace. ebs, epibranchial spine; pbs, posterobranchial spine. Scale bars: 10mm." figureDoi="http://doi.org/10.5281/zenodo.5248664" httpUri="https://zenodo.org/record/5248664/files/figure.png" pageId="56" pageNumber="57">Fig. 18A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CE177823FFA8F9545BE4FEB0F652FD8A" blockId="56.[151,1436,151,572]" pageId="56" pageNumber="57">
|
||
<taxonomicName id="09A803A0FFA8F9545BE4FEB0F5B0FEAA" authorityName="Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross" authorityYear="2003" box="[199,495,260,284]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="FCDCA431FFA8F9545BE4FEB0F5B0FEAA" box="[199,495,260,284]" italics="true" pageId="56" pageNumber="57">Cenomanocarcinus beardi</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from its congeners by the lack of a tooth between the strong fourth anterolateral tooth and the epibranchial spine. It therefore has five, instead of six or seven, anterolateral teeth (epibranchial included, outer orbital excluded). The epibranchial spine is strongly developed, and the posterolateral margin is smooth and rounded; these characters are similar to
|
||
<taxonomicName id="09A803A0FFA8F9545878FEC4F78FFE3E" authorityName="Van Straelen" authorityYear="1936" box="[859,976,368,392]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA8F9545878FEC4F78FFE3E" box="[859,976,368,392]" italics="true" pageId="56" pageNumber="57">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
and thus differentiate
|
||
<taxonomicName id="09A803A0FFA8F9545FF2FEC4F164FE3E" authorityName="Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross" authorityYear="2003" box="[1233,1339,368,392]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="FCDCA431FFA8F9545FF2FEC4F164FE3E" box="[1233,1339,368,392]" italics="true" pageId="56" pageNumber="57">C. beardi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="09A803A0FFA8F9545EA1FEC4F57EFE1A" authorityName="Stenzel" authorityYear="1945" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="vanstraeleni">
|
||
<emphasis id="FCDCA431FFA8F9545EA1FEC4F57EFE1A" italics="true" pageId="56" pageNumber="57">C. vanstraeleni</emphasis>
|
||
</taxonomicName>
|
||
. The strong spiniform tooth halfway the posterolateral margin, characteristic of
|
||
<taxonomicName id="09A803A0FFA8F9545F97FE20F175FE1A" authorityName="Van Straelen" authorityYear="1936" box="[1204,1322,404,428]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA8F9545F97FE20F175FE1A" box="[1204,1322,404,428]" italics="true" pageId="56" pageNumber="57">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
, is nevertheless absent in
|
||
<taxonomicName id="09A803A0FFA8F9545A7AFE0CF59DFE66" authorityName="Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross" authorityYear="2003" box="[345,450,440,464]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="FCDCA431FFA8F9545A7AFE0CF59DFE66" box="[345,450,440,464]" italics="true" pageId="56" pageNumber="57">C. beardi</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the posterior spine at the base of the branchial crest is stronger. The three crests have fewer but stronger tubercles and the transversely vaulted connection (referred to as ‘H’-shape by
|
||
<bibRefCitation id="AA3905D2FFA8F9545E4AFE68F5C0FDA1" author="Schweitzer, C. E. & Feldmann, R. M. & Fam, J. & Hessin, W. A. & Hetrick, S. W. & Nyborg, T. G. & Ross, R. L. M." pageId="56" pageNumber="57" refId="ref143653" refString="Schweitzer, C. E., Feldmann, R. M., Fam, J., Hessin, W. A., Hetrick, S. W., Nyborg, T. G. & Ross, R. L. M. (2003 a) Cretaceous and Eocene Decapod Crustaceans from Southern Vancouver Island, British Columbia, Canada, 66 pp. NRC Research Press, Ottawa." type="book" year="2003">
|
||
Schweitzer
|
||
<emphasis id="FCDCA431FFA8F9545BD3FDB5F57EFDAE" box="[240,289,512,536]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
|
||
. 2003a: 36
|
||
</bibRefCitation>
|
||
) is clearly seen, whereas it is indistinct or obsolete in
|
||
<taxonomicName id="09A803A0FFA8F95458D9FDB4F031FDAE" authorityName="Van Straelen" authorityYear="1936" box="[1018,1134,512,536]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA8F95458D9FDB4F031FDAE" box="[1018,1134,512,536]" italics="true" pageId="56" pageNumber="57">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="FCDCA431FFA8F9545F87FDB4F109FDAE" box="[1188,1366,512,536]" italics="true" pageId="56" pageNumber="57">
|
||
<taxonomicName id="09A803A0FFA8F9545F87FDB4F10DFDAE" authorityName="Stenzel" authorityYear="1945" box="[1188,1362,512,536]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="vanstraeleni">C. vanstraeleni</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Characters are summarised in Table 5.
|
||
</paragraph>
|
||
<paragraph id="CE177823FFA8F9545BB4FDC2F07FFD39" blockId="56.[151,1056,627,655]" box="[151,1056,627,655]" pageId="56" pageNumber="57">
|
||
<emphasis id="FCDCA431FFA8F9545BB4FDC2F554FD38" bold="true" box="[151,267,630,654]" pageId="56" pageNumber="57">TABLE 5.</emphasis>
|
||
Characters of
|
||
<taxonomicName id="09A803A0FFA8F9545A8BFDC3F69BFD38" authorityName="Schweitzer, Feldmann, Fam, Hessin, Hetrick, Nyborg & Ross" authorityYear="2003" box="[424,708,631,654]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="FCDCA431FFA8F9545A8BFDC3F69BFD38" box="[424,708,631,654]" italics="true" pageId="56" pageNumber="57">Cenomanocarcinus beardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="09A803A0FFA8F95459F2FDC3F71FFD38" authorityName="Van Straelen" authorityYear="1936" box="[721,832,631,654]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="inflatus">
|
||
<emphasis id="FCDCA431FFA8F95459F2FDC3F71FFD38" box="[721,832,631,654]" italics="true" pageId="56" pageNumber="57">C. inflatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="09A803A0FFA8F9545857FDC3F047FD39" authorityName="Stenzel" authorityYear="1945" box="[884,1048,627,655]" class="Malacostraca" family="Cenomanocarcinidae" genus="Cenomanocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="vanstraeleni">
|
||
<emphasis id="FCDCA431FFA8F9545857FDC3F047FD39" box="[884,1048,627,655]" italics="true" pageId="56" pageNumber="57">C. vanstraeleni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |