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<mods:title id="B7D6F3229FA3A4C53465F025F3E1D001">A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215</mods:title>
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<mods:namePart id="68055425FDEF314BE2E72283F2A2E032">Van Bakel, Barry W. M.</mods:namePart>
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<mods:namePart id="367904A46A523FC4BE71E9A058F62A4D">Guinot, Danièle</mods:namePart>
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<mods:namePart id="7D76A21174656D533EE9652A25413F64">Artal, Pedro</mods:namePart>
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<mods:namePart id="7E14EF7654463151A324C180FA1A5B01">Fraaije, René H. B.</mods:namePart>
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<subSubSection id="86B22BA8FF8AF9765BB4FF23F5A6FF07" box="[151,505,151,177]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="CE177823FF8AF9765BB4FF23F5A6FF07" blockId="26.[151,505,151,210]" box="[151,505,151,177]" pageId="26" pageNumber="27">
<heading id="955FCF4FFF8AF9765BB4FF23F5A6FF07" bold="true" box="[151,505,151,177]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="FCDCA431FF8AF9765BB4FF23F5A6FF07" bold="true" box="[151,505,151,177]" pageId="26" pageNumber="27">
<taxonomicName id="09A803A0FF8AF9765BB4FF23F5EFFF07" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[151,432,151,177]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="navarrensis" status="sp. nov.">
<emphasis id="FCDCA431FF8AF9765BB4FF23F5EFFF07" bold="true" box="[151,432,151,177]" italics="true" pageId="26" pageNumber="27">Eucorystes navarrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7EF194AFF8AF9765A94FF23F5A6FF07" box="[439,505,151,177]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="86B22BA8FF8AF9765BB4FF0EF57FFF64" box="[151,288,186,210]" pageId="26" pageNumber="27" type="description">
<paragraph id="CE177823FF8AF9765BB4FF0EF57FFF64" blockId="26.[151,505,151,210]" box="[151,288,186,210]" pageId="26" pageNumber="27">
(
<figureCitation id="569364A6FF8AF9765BBCFF0EF548FF64" box="[159,279,186,210]" captionStart="FIGURE 6" captionStartId="27.[151,250,1888,1911]" captionTargetBox="[151,1436,195,1864]" captionTargetId="figure-95@27.[151,1436,195,1864]" captionTargetPageId="27" captionText="FIGURE 6. A, C, Eucorystes carteri (McCoy, 1854) (Palaeocorystidae), IRScNB unregistered [Van Straelen Collection; female], upper Albian Cambridge Greensand, Cambridge (southern England), dorsal view of carapace, and ventral view showing thoracic sternum and appendages; B, Eucorystes navarrensis n. sp., MGSB75291 (holotype), Albian, Alloz (Navarra, northern Spain), dorsal view of carapace; D, Eucorystes navarrensis n. sp., MGSB75292a (paratype), silicone rubber cast of natural mould showing dorsal carapace, without cuticle. Scale bars: 5mm." figureDoi="http://doi.org/10.5281/zenodo.5248652" httpUri="https://zenodo.org/record/5248652/files/figure.png" pageId="26" pageNumber="27">Fig. 6B, D</figureCitation>
)
</paragraph>
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<subSubSection id="86B22BA8FF8AF9765BB4FEB6F7B7FE44" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="CE177823FF8AF9765BB4FEB6F513FE78" blockId="26.[151,1437,258,1686]" pageId="26" pageNumber="27">
<emphasis id="FCDCA431FF8AF9765BB4FEB6F54EFEAD" bold="true" box="[151,273,258,283]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Diagnosis.</emphasis>
Small-sized carapace, subhexagonally elongated, maximum width at level of epibranchial spine. Dorsal surface with numerous strap-like lobes, grooves, separating dorsal regions into numerous portions. Front narrow, bifid, with 2 long, divergent spines. Narrow, rounded, salient axial carina along entire carapace axis. Dorsal regions defined by narrow, clearly flat-topped lobes; hepatic region with 2 small protuberances; mesogastric region rhombic; urogastric region V-shaped; epibranchial region subdivided. Posterior half of carapace lacking welldefined regions.
</paragraph>
<paragraph id="CE177823FF8AF9765BE4FE6EF7B7FE44" blockId="26.[151,1437,258,1686]" box="[199,1000,474,499]" pageId="26" pageNumber="27">
<emphasis id="FCDCA431FF8AF9765BE4FE6EF5EBFE45" bold="true" box="[199,436,474,499]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Derivation of name.</emphasis>
<materialsCitation id="7EC0727EFF8AF9765A98FE6FF7BBFE44" box="[443,996,474,499]" country="Spain" location="From" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Navarra">
<location id="CB772EF8FF8AF9765A98FE6FF5A7FE44" LSID="urn:lsid:plazi:treatment:4601C935FF8AF9765BB4FF23F64DF920:CB772EF8FF8AF9765A98FE6FF5A7FE44" box="[443,504,475,498]" country="Spain" name="From" pageId="26" pageNumber="27" stateProvince="Navarra">From</location>
<collectingRegion id="0C6CB6C1FF8AF9765ADCFE6FF606FE44" box="[511,601,475,498]" country="Spain" name="Navarra" pageId="26" pageNumber="27">Navarra</collectingRegion>
(northern
<collectingCountry id="B6BF38B3FF8AF97659ECFE6EF74CFE44" box="[719,787,474,498]" name="Spain" pageId="26" pageNumber="27">Spain</collectingCountry>
), the
<typeStatus id="1113C681FF8AF976586CFE6FF720FE45" box="[847,895,475,499]" pageId="26" pageNumber="27">type</typeStatus>
location
</materialsCitation>
.
</paragraph>
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<subSubSection id="86B22BA8FF8AF9765BE4FE4AF480FD34" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="CE177823FF8AF9765BE4FE4AF480FD34" blockId="26.[151,1437,258,1686]" pageId="26" pageNumber="27">
<emphasis id="FCDCA431FF8AF9765BE4FE4AF5EFFDA1" bold="true" box="[199,432,510,535]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Material examined.</emphasis>
<materialsCitation id="7EC0727EFF8AF9765A95FE4AF480FD34" collectionCode="MGSB" county="Alloz" location="Instituto Geologico de Espana" municipality="Albian" pageId="26" pageNumber="27" specimenCode="MGSB75291, MGSB75292" specimenCount="2" stateProvince="Navarra" typeStatus="holotype">
<typeStatus id="1113C681FF8AF9765A95FE4AF67EFDA0" box="[438,545,510,534]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
, a dorsal carapace (
<specimenCode id="9E0ED058FF8AF97659D8FE4AF7C8FDA0" box="[763,919,510,534]" collectionCode="MGSB" pageId="26" pageNumber="27">MGSB75291</specimenCode>
);
<typeStatus id="1113C681FF8AF9765889FE4BF052FDA1" box="[938,1037,511,535]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
, a dorsal carapace preserved upside down (
<specimenCode id="9E0ED058FF8AF9765BC7FD96F5DFFD8C" box="[228,384,546,570]" collectionCode="MGSB" pageId="26" pageNumber="27">MGSB75292</specimenCode>
a); a cast with well-preserved front (
<specimenCode id="9E0ED058FF8AF9765831FD96F7F1FD8C" box="[786,942,546,570]" collectionCode="MGSB" pageId="26" pageNumber="27">MGSB75292</specimenCode>
b), all from approximately 1.5 km northwest of the village of
<collectingCounty id="277600AFFF8AF9765A76FDF2F5CCFDE8" box="[341,403,582,606]" pageId="26" pageNumber="27">Alloz</collectingCounty>
(
<collectingRegion id="0C6CB6C1FF8AF9765A80FDF2F65CFDE8" box="[419,515,582,606]" country="Spain" name="Navarra" pageId="26" pageNumber="27">Navarra</collectingRegion>
), from limestones considered to be of
<collectingMunicipality id="2E73E259FF8AF976589AFDF2F05AFDE8" box="[953,1029,582,606]" pageId="26" pageNumber="27">Albian</collectingMunicipality>
age (
<location id="CB772EF8FF8AF9765F67FDF2F1C3FDE8" LSID="urn:lsid:plazi:treatment:4601C935FF8AF9765BB4FF23F64DF920:CB772EF8FF8AF9765F67FDF2F1C3FDE8" box="[1092,1436,582,606]" county="Alloz" municipality="Albian" name="Instituto Geologico de Espana" pageId="26" pageNumber="27" stateProvince="Navarra">Instituto Geológico de España</location>
1978).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="86B22BA8FF8AF9765BE4FD3AF575FB28" pageId="26" pageNumber="27" type="description">
<paragraph id="CE177823FF8AF9765BE4FD3AF575FB28" blockId="26.[151,1437,258,1686]" pageId="26" pageNumber="27">
<emphasis id="FCDCA431FF8AF9765BE4FD3AF507FD11" bold="true" box="[199,344,654,679]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Description.</emphasis>
Carapace small, subhexagonally elongated in outline, maximum width at base of epibranchial region, dorsal surface weakly convex in both directions. Orbitofrontal margin wide, only partially preserved. Front narrow, bifid, with long, divergent spines. Lateral margins broadly arched, lateral spines incompletely preserved. Posterior margin strongly concave, notably shorter than orbitofrontal margin. Mesogastric region small, markedly rhombohedric in shape, well defined by furrows, anterior process narrow, ridged, not reaching base of front. Protogastric region clearly subdivided into 3 isolated, flat-topped lobes. Epigastric regions elongated, joining central protogastric ridge. Urogastric region V-shaped, with lateral portions extending obliquely forwards. Hepatic region with 2 notable small protuberances; anterior one conical, below orbital margin, posterior one larger, slightly elongated. Cervical groove well-defined, deeper at axial portion, interrupted by axial carina. Epibranchial region divided into 3 strap-like lobes; anterior portion subtriangular, median portion near straight, directed obliquely towards lateral margins, fairly well separated from urogastric lobe, posterior portion arched. Cardiac region relatively narrow, elongated, anterior portion laterally bounded by deep branchiocardiac grooves. Continuous axial carina along carapace fairly salient, rounded. Posterior half of carapace, strap-like lobes covered by fungiform granules, markedly enlarged on axial carina; grooves, furrows in anterior half of carapace densely covered by minute setal pits.
</paragraph>
</subSubSection>
<subSubSection id="86B22BA8FF8AF9765BE4FB1FF64DF920" pageId="26" pageNumber="27" type="discussion">
<paragraph id="CE177823FF8AF9765BE4FB1FF64DF920" blockId="26.[151,1437,258,1686]" pageId="26" pageNumber="27">
<emphasis id="FCDCA431FF8AF9765BE4FB1FF564FB75" bold="true" box="[199,315,1195,1219]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Remarks.</emphasis>
The new species is differentiated from all congeners in having an uninterrupted axial carina over the complete dorsal surface. In
<taxonomicName id="09A803A0FF8AF9765AF7FB7BF608FB50" authorityName="Wright &amp; Collins" authorityYear="1972" box="[468,599,1230,1254]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="FCDCA431FF8AF9765AF7FB7BF608FB50" box="[468,599,1230,1254]" italics="true" pageId="26" pageNumber="27">E. ligulatus</emphasis>
</taxonomicName>
, the only other species that shows a distinct axial carina on the posterior carapace (
<bibRefCitation id="AA3905D2FF8AF9765A2BFB46F64CFABC" author="Wright, C. W. &amp; Collins, J. S. H." box="[264,531,1266,1290]" pageId="26" pageNumber="27" pagination="1 - 114" refId="ref150164" refString="Wright, C. W. &amp; Collins, J. S. H. (1972) British Cretaceous crabs. Monograph of the Palaeontographical Society (London), 126 (533), 1 - 114, 22 pls." type="journal article" year="1972">Wright &amp; Collins 1972</bibRefCitation>
: pl. 16, fig. 4), the carina is interrupted at both sides of the urogastric region. In addition,
<taxonomicName id="09A803A0FF8AF9765A20FAA3F5D9FA98" authorityName="Wright &amp; Collins" authorityYear="1972" box="[259,390,1302,1326]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ligulatus">
<emphasis id="FCDCA431FF8AF9765A20FAA3F5D9FA98" box="[259,390,1302,1326]" italics="true" pageId="26" pageNumber="27">E. ligulatus</emphasis>
</taxonomicName>
bears additional branchial strap-like lobes, which are absent in
<taxonomicName id="09A803A0FF8AF9765F47FAA3F157FA98" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[1124,1288,1303,1326]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="navarrensis" status="sp. nov.">
<emphasis id="FCDCA431FF8AF9765F47FAA3F157FA98" box="[1124,1288,1303,1326]" italics="true" pageId="26" pageNumber="27">E. navarrensis</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FF8AF9765E31FAA3F10CFA99" bold="true" box="[1298,1363,1303,1327]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="E7EF194AFF8AF9765E31FAA3F10CFA99" box="[1298,1363,1303,1327]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Apart from the nature of the axial carina, the new species can be distinguished from
<taxonomicName id="09A803A0FF8AF97658DAFA8FF03BFAE4" baseAuthorityName="McCoy" baseAuthorityYear="1854" box="[1017,1124,1339,1362]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="FCDCA431FF8AF97658DAFA8FF03BFAE4" box="[1017,1124,1339,1362]" italics="true" pageId="26" pageNumber="27">E. carteri</emphasis>
</taxonomicName>
and
<taxonomicName id="09A803A0FF8AF9765FBAFA8FF153FAE5" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[1177,1292,1339,1363]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="iserbyti" status="sp. nov.">
<emphasis id="FCDCA431FF8AF9765FBAFA8FF153FAE5" box="[1177,1292,1339,1363]" italics="true" pageId="26" pageNumber="27">E. iserbyti</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FF8AF9765E31FA8FF10EFAE5" bold="true" box="[1298,1361,1339,1363]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="E7EF194AFF8AF9765E31FA8FF10EFAE5" box="[1298,1361,1339,1363]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the different nature and organisation of dorsal grooves and strap-like ridges. In
<taxonomicName id="09A803A0FF8AF97658D4FAEBF0C4FAC0" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[1015,1179,1375,1398]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="navarrensis" status="sp. nov.">
<emphasis id="FCDCA431FF8AF97658D4FAEBF0C4FAC0" box="[1015,1179,1375,1398]" italics="true" pageId="26" pageNumber="27">E. navarrensis</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FF8AF9765F87FAEBF0BBFAC1" bold="true" box="[1188,1252,1375,1399]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="E7EF194AFF8AF9765F87FAEBF0BBFAC1" box="[1188,1252,1375,1399]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the grooves are broader and more numerous, which lead to narrower, more acute lobes; in
<taxonomicName id="09A803A0FF8AF97658C3FA37F00BFA2D" authority="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt, 2012" authorityName="Van Bakel &amp; Guinot &amp; Artal &amp; Fraaije &amp; Jagt" authorityYear="2012" box="[992,1108,1411,1435]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="iserbyti" status="sp. nov.">
<emphasis id="FCDCA431FF8AF97658C3FA37F00BFA2D" box="[992,1108,1411,1435]" italics="true" pageId="26" pageNumber="27">E. iserbyti</emphasis>
</taxonomicName>
<emphasis id="FCDCA431FF8AF9765F7FFA37F0C4FA2D" bold="true" box="[1116,1179,1411,1435]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="E7EF194AFF8AF9765F7FFA37F0C4FA2D" box="[1116,1179,1411,1435]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the lobes are rounded, rather than flat-topped.
<taxonomicName id="09A803A0FF8AF9765A87FA13F6CCFA09" baseAuthorityName="Bishop" baseAuthorityYear="1983" box="[420,659,1447,1471]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="eichhorni">
<emphasis id="FCDCA431FF8AF9765A87FA13F6CCFA09" box="[420,659,1447,1471]" italics="true" pageId="26" pageNumber="27">Eucorystes eichhorni</emphasis>
</taxonomicName>
, a much younger (late Campanian) species from
<collectingRegion id="0C6CB6C1FF8AF9765FEAFA13F173FA08" box="[1225,1324,1447,1470]" country="United States of America" name="Montana" pageId="26" pageNumber="27">Montana</collectingRegion>
(
<collectingCountry id="B6BF38B3FF8AF9765E1DFA12F1D2FA08" box="[1342,1421,1446,1470]" name="United States of America" pageId="26" pageNumber="27">U.S.A.</collectingCountry>
), also has profuse dorsal regional ornament, presents distinct, long grooves bounding the anterior mesogastric process, as well as a different distribution of strap-like lobes.
<taxonomicName id="09A803A0FF8AF9765864FA5BF07DF9B0" baseAuthorityName="Glaessner" baseAuthorityYear="1980" box="[839,1058,1519,1542]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="FCDCA431FF8AF9765864FA5BF07DF9B0" box="[839,1058,1519,1542]" italics="true" pageId="26" pageNumber="27">Eucorystes exiguus</emphasis>
</taxonomicName>
, from the lower Cenomanian of Bathurst Island (
<collectingCountry id="B6BF38B3FF8AF9765A72F9A6F5E2F99C" box="[337,445,1554,1578]" name="Australia" pageId="26" pageNumber="27">Australia</collectingCountry>
), and
<taxonomicName id="09A803A0FF8AF9765923F9A7F6CCF99D" authorityName="Wright &amp; Collins" authorityYear="1972" box="[512,659,1555,1579]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oxtedensis">
<emphasis id="FCDCA431FF8AF9765923F9A7F6CCF99D" box="[512,659,1555,1579]" italics="true" pageId="26" pageNumber="27">E. oxtedensis</emphasis>
</taxonomicName>
exhibit less clearly subdivided anterior regions and deeper branchiocardiac grooves.
<taxonomicName id="09A803A0FF8AF9765A7CF983F636F9F9" authorityName="Nagao" authorityYear="1931" box="[351,617,1591,1615]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="FCDCA431FF8AF9765A7CF983F636F9F9" box="[351,617,1591,1615]" italics="true" pageId="26" pageNumber="27">Eucorystes intermedius</emphasis>
</taxonomicName>
shows a much weaker dorsal ornament, whereas
<taxonomicName id="09A803A0FF8AF9765F99F983F1C3F9F9" authorityName="Ilyin &amp; Pistshikova" authorityYear="2005" box="[1210,1436,1591,1615]" class="Malacostraca" family="Raninidae" genus="Eucorystes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mangyshlakensis">
<emphasis id="FCDCA431FF8AF9765F99F983F1C3F9F9" box="[1210,1436,1591,1615]" italics="true" pageId="26" pageNumber="27">E. mangyshlakensis</emphasis>
</taxonomicName>
exhibits deeper branchiocardiac grooves and merged, rather than isolated protogastric strap-like lobes, which have a more forwardly directed course.
</paragraph>
</subSubSection>
</treatment>
</document>