treatments-xml/data/21/71/72/217172DB856A5B3687DF4E8B15784A37.xml

<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.18.10007" ID-GBIF-Taxon="182224609" ID-Pensoft-Pub="1314-4049-18-1" ID-Pensoft-UUID="742AFFD8FF9DFFE0FFE1351B1C21FF99" ID-Zenodo-Dep="575611" ModsDocID="1314-4049-18-1" checkinTime="1481875202450" checkinUser="pensoft" docAuthor="Petersen, Ronald H. &amp; Hughes, Karen W." docDate="2016" docId="217172DB856A5B3687DF4E8B15784A37" docLanguage="en" docName="MycoKeys 18: 1-122" docOrigin="MycoKeys 18" docPubDate="2016-12-15" docSource="http://dx.doi.org/10.3897/mycokeys.18.10007" docTitle="Gymnopus sublaccatus R. H. Petersen 2016, sp. nov." docType="treatment" docVersion="2" id="742AFFD8FF9DFFE0FFE1351B1C21FF99" lastPageId="100" lastPageNumber="101" masterDocId="742AFFD8FF9DFFE0FFE1351B1C21FF99" masterDocTitle="Micromphale sect. Perforantia (Agaricales, Basidiomycetes); Expansion and phylogenetic placement" masterLastPageNumber="122" masterPageNumber="1" pageId="92" pageNumber="93" updateTime="1643498509278" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Micromphale sect. Perforantia (Agaricales, Basidiomycetes); Expansion and phylogenetic placement</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petersen, Ronald H.</mods:namePart>
<mods:affiliation>Ecology &amp; Evolutionary Biology, University of Tennessee, Knoxville, TN 37996 - 1100 USA</mods:affiliation>
<mods:nameIdentifier type="email">repete@utk.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hughes, Karen W.</mods:namePart>
<mods:affiliation>Ecology &amp; Evolutionary Biology, University of Tennessee, Knoxville, TN 37996 - 1100 USA</mods:affiliation>
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<mods:title>MycoKeys</mods:title>
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<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-12-15</mods:number>
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<mods:detail type="volume">
<mods:number>18</mods:number>
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<mods:start>1</mods:start>
<mods:end>122</mods:end>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.18.10007</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.18.10007</mods:identifier>
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<treatment ID-GBIF-Taxon="182224609" LSID="urn:lsid:plazi:treatment:217172DB856A5B3687DF4E8B15784A37" httpUri="http://treatment.plazi.org/id/217172DB856A5B3687DF4E8B15784A37" lastPageId="100" lastPageNumber="101" pageId="92" pageNumber="93">
<subSubSection pageId="92" pageNumber="93" type="nomenclature">
<paragraph pageId="92" pageNumber="93">
11. 
<taxonomicName LSID="217172DB-856A-5B36-87DF-4E8B15784A37" authority="R. H. Petersen" authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="92" pageNumber="93" phylum="Basidiomycota" rank="species" species="sublaccatus" status="sp. nov.">Gymnopus sublaccatus R.H. Petersen</taxonomicName>
<taxonomicNameLabel pageId="92" pageNumber="93">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="93" pageNumber="94" type="holotype">
<paragraph pageId="93" pageNumber="94">
<pageBreakToken pageId="93" pageNumber="94" start="start">Holotype</pageBreakToken>
.
</paragraph>
<paragraph pageId="93" pageNumber="94">
<emphasis bold="true" pageId="93" pageNumber="94">Canada</emphasis>
, 
<emphasis bold="true" pageId="93" pageNumber="94">British Columbia</emphasis>
, Victoria, Saanich Peninsula, Observatory Hill, 
<geoCoordinate degrees="48.5262" direction="north" orientation="latitude" precision="5" value="48.5262">N48.5262°</geoCoordinate>
, 
<geoCoordinate degrees="123.422" direction="west" orientation="longitude" precision="55" value="-123.422">W123.422°</geoCoordinate>
, 4.XI.2011, coll &amp; det O. Ceska (as 
<taxonomicName class="Basidiomycetes" family="Marasmiaceae" genus="Marasmius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Marasmius androsaceus" order="Agaricales" pageId="93" pageNumber="94" phylum="Basidiomycota" rank="species" species="androsaceus">
<emphasis italics="true" pageId="93" pageNumber="94">Marasmius androsaceus</emphasis>
</taxonomicName>
), UBC-F-25212.
</paragraph>
</subSubSection>
<subSubSection pageId="94" pageNumber="95" type="etymology">
<paragraph pageId="94" pageNumber="95">
<pageBreakToken pageId="94" pageNumber="95" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="94" pageNumber="95">sub- = Latin: less than; laccatus = Latin: appearing polished or varnished, referring to varnished appearance of dried pileus.</paragraph>
</subSubSection>
<subSubSection pageId="94" pageNumber="95" type="diagnosis">
<paragraph pageId="94" pageNumber="95">Diagnosis.</paragraph>
<paragraph pageId="94" pageNumber="95">
1) Fruiting on needles of 
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Coniferopsida" family="Cupressaceae" genus="Thuja" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Thuja" order="Pinales" pageId="94" pageNumber="95" phylum="Gymnospermae" rank="genus">
<emphasis italics="true" pageId="94" pageNumber="95">Thuja</emphasis>
</taxonomicName>
(and 
<taxonomicName authorityName="E.A.Carriere" authorityYear="1867" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pseudotsuga" order="Pinales" pageId="94" pageNumber="95" phylum="Coniferophyta" rank="genus">
<emphasis italics="true" pageId="94" pageNumber="95">Pseudotsuga</emphasis>
</taxonomicName>
); 2) cheilocystidia rare, small, clavate, smooth, without setulae; 3) stipe near ochraceous buff above, downward rusty brown; 4) pileal hairs usually smooth, rarely roughened; 5) dried pileus sublaccate; 6) pileipellis of smooth, unencrusted, repent hyphae in slime matrix; 7) rhizomorphs inconspicuous, with minute black basal pad.
</paragraph>
</subSubSection>
<subSubSection lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95" type="description">
<paragraph pageId="94" pageNumber="95">The following description is based solely on dried material.</paragraph>
<paragraph lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95">
<emphasis bold="true" pageId="94" pageNumber="95">Basidiomata</emphasis>
(Fig. 
<figureCitation captionStart="Figure 78" captionStartId="F78" captionText="Figure 78. Gymnopus sublaccatus. A Basidiomata showing adventitious rhizomorphs and stipe insertion to pileus B Basidiospores C Substrate scales with back spots and rhizomorphs. Standard bars: A = 20 mm; B = 5 µm; C = not to scale. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure78" httpUri="https://binary.pensoft.net/fig/115142" pageId="94" pageNumber="95">78A</figureCitation>
) diminutive. 
<emphasis bold="true" pageId="94" pageNumber="95">Pileus</emphasis>
2-8 mm broad, convex to plano-convex, minutely laccate, vaguely tuberculate, not striate or sulcate; disc 
<pageBreakToken pageId="95" pageNumber="96" start="start">about</pageBreakToken>
&quot;sayal brown&quot; 6C5 to &quot;ochraceous buff&quot; 5A5; limb and margin about &quot;light ochraceous buff&quot; 5A4 to &quot;tilleul buff&quot; 7B2. 
<emphasis bold="true" pageId="95" pageNumber="96">Lamellae</emphasis>
(Fig. 
<figureCitation captionStart="Figure 78" captionStartId="F78" captionText="Figure 78. Gymnopus sublaccatus. A Basidiomata showing adventitious rhizomorphs and stipe insertion to pileus B Basidiospores C Substrate scales with back spots and rhizomorphs. Standard bars: A = 20 mm; B = 5 µm; C = not to scale. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure78" httpUri="https://binary.pensoft.net/fig/115142" pageId="95" pageNumber="96">78A</figureCitation>
) pseudocollariate (dried), adnate, distant, thickish, total lamellae = 20-23, through lamellae = 9-11, now (dried) &quot;tilleul buff&quot; 7B2 to &quot;olive buff&quot; 3B3; lamellulae in a single rank. 
<emphasis bold="true" pageId="95" pageNumber="96">Stipe</emphasis>
18-25 
<normalizedToken originalValue="×">x</normalizedToken>
0.6-0.8 mm, terete, equal, hollow to lightly stuffed, subinsititious, minutely shaggy 
<pageBreakToken pageId="96" pageNumber="97" start="start">to</pageBreakToken>
silky above, downward becoming minutely barbed (35
<normalizedToken originalValue="×">x</normalizedToken>
), above about &quot;ochraceous buff&quot; 5A5, downward through &quot;army brown&quot; 8D5, increasingly dark to rusty brown (
<normalizedToken originalValue="“Prout’s">&quot;Prout's</normalizedToken>
brown&quot; 5F6 to &quot;Vandyke brown&quot; 7E6), sometimes slightly expanded at base; 
<pageBreakToken pageId="97" pageNumber="98" start="start">stipe</pageBreakToken>
medulla white. 
<emphasis bold="true" pageId="97" pageNumber="98">Rhizomorphs</emphasis>
(Fig. 
<figureCitation captionStart="Figure 78" captionStartId="F78" captionText="Figure 78. Gymnopus sublaccatus. A Basidiomata showing adventitious rhizomorphs and stipe insertion to pileus B Basidiospores C Substrate scales with back spots and rhizomorphs. Standard bars: A = 20 mm; B = 5 µm; C = not to scale. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure78" httpUri="https://binary.pensoft.net/fig/115142" pageId="97" pageNumber="98">78C</figureCitation>
) (if produced) inconspicuous, often represented only by minute, black basal pads. 
<emphasis bold="true" pageId="97" pageNumber="98">Taste</emphasis>
and 
<emphasis bold="true" pageId="97" pageNumber="98">odor</emphasis>
not recorded.
</paragraph>
<caption ID-Zenodo-Dep="984935" doi="10.3897/mycokeys.18.10007.figure78" httpUri="https://binary.pensoft.net/fig/115142" pageId="97" pageNumber="98" start="Figure 78" startId="F78">
<paragraph pageId="97" pageNumber="98">
<emphasis bold="true" pageId="97" pageNumber="98">Figure 78.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="97" pageNumber="98" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="97" pageNumber="98">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="97" pageNumber="98">A</emphasis>
Basidiomata showing adventitious rhizomorphs and stipe insertion to pileus 
<emphasis bold="true" pageId="97" pageNumber="98">B</emphasis>
Basidiospores 
<emphasis bold="true" pageId="97" pageNumber="98">C</emphasis>
Substrate scales with back spots and rhizomorphs. Standard bars: 
<emphasis bold="true" pageId="97" pageNumber="98">A</emphasis>
= 20 mm; 
<emphasis bold="true" pageId="97" pageNumber="98">B</emphasis>
= 5 
<normalizedToken originalValue="µm">µm</normalizedToken>
; 
<emphasis bold="true" pageId="97" pageNumber="98">C</emphasis>
= not to scale. UBC 25212.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="99" lastPageNumber="100" pageId="97" pageNumber="98" type="habitat">
<paragraph pageId="97" pageNumber="98">Habitat and phenology.</paragraph>
<paragraph pageId="97" pageNumber="98">
Fruiting on needles of 
<taxonomicName class="Coniferopsida" family="Cupressaceae" genus="Thuja" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Thuja plicata" order="Pinales" pageId="97" pageNumber="98" phylum="Gymnospermae" rank="species" species="plicata">
<emphasis italics="true" pageId="97" pageNumber="98">Thuja plicata</emphasis>
</taxonomicName>
, less often on 
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="97" pageNumber="98" phylum="Coniferophyta" rank="species" species="menziesii">
<emphasis italics="true" pageId="97" pageNumber="98">Pseudotsuga menziesii</emphasis>
</taxonomicName>
; British Columbia; Autumn to early Winter.
</paragraph>
<paragraph lastPageId="99" lastPageNumber="100" pageId="97" pageNumber="98">
<emphasis bold="true" pageId="97" pageNumber="98">Pileipellis</emphasis>
involved in a slime matrix which includes detersile encrusting material, of the following elements: 1) pileal hairs (Fig. 
<figureCitation captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Gymnopus sublaccatus. Pileipellis elements. A-E Pileal hairs F Secondary septa of pileipellis hypha. Standard bars = 10 µm. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure79" httpUri="https://binary.pensoft.net/fig/115144" pageId="97" pageNumber="98">79</figureCitation>
) -150 
<normalizedToken originalValue="×">x</normalizedToken>
3-6 
<normalizedToken originalValue="µm">µm</normalizedToken>
, erect, firm-walled, smooth or rarely weakly roughened, slightly subcapitulate; 2) repent hyphae 4.5-7.5 
<normalizedToken originalValue="µm">µm</normalizedToken>
diam, firm-walled, conspicuously clamped but often secondarily septate, without discernable slime sheath. Pileus and lamellar tramae loosely interwoven; hyphae 3-5.5 
<normalizedToken originalValue="µm">µm</normalizedToken>
diam, smooth, firm-walled, without slime sheath, conspicuously clamped. 
<emphasis bold="true" pageId="97" pageNumber="98">Pleurocystidia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 80" captionStartId="F80" captionText="Figure 80. Gymnopus sublaccatus. Hymenial elements. A-D Pleurocystidia E Basidiole F-H Basidia. Standard bars = 10 µm. UBC 15356." figureDoi="10.3897/mycokeys.18.10007.figure80" httpUri="https://binary.pensoft.net/fig/115145" pageId="97" pageNumber="98">80A-D</figureCitation>
) 25-35(-41) 
<normalizedToken originalValue="×">x</normalizedToken>
7-8 
<normalizedToken originalValue="µm">µm</normalizedToken>
, fusiform to clavo-fusiform, without partitioned contents but with vague vacuolated area in midsection (PhC), conspicuously clamped. Basidioles (Fig. 
<figureCitation captionStart="Figure 80" captionStartId="F80" captionText="Figure 80. Gymnopus sublaccatus. Hymenial elements. A-D Pleurocystidia E Basidiole F-H Basidia. Standard bars = 10 µm. UBC 15356." figureDoi="10.3897/mycokeys.18.10007.figure80" httpUri="https://binary.pensoft.net/fig/115145" pageId="97" pageNumber="98">80E</figureCitation>
) clavate; 
<emphasis bold="true" pageId="97" pageNumber="98">basidia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 80" captionStartId="F80" captionText="Figure 80. Gymnopus sublaccatus. Hymenial elements. A-D Pleurocystidia E Basidiole F-H Basidia. Standard bars = 10 µm. UBC 15356." figureDoi="10.3897/mycokeys.18.10007.figure80" httpUri="https://binary.pensoft.net/fig/115145" pageId="97" pageNumber="98">80F-H</figureCitation>
) 24-27 
<normalizedToken originalValue="×">x</normalizedToken>
8-9 
<normalizedToken originalValue="µm">µm</normalizedToken>
, clavate, 4-sterigmate, clamped; contents multigranular at maturity; effete pleurocystidia and basidia emptying but not col 
<pageBreakToken pageId="98" pageNumber="99" start="start">lapsing</pageBreakToken>
(
<normalizedToken originalValue="“husking”">&quot;husking&quot;</normalizedToken>
Fig. 
<figureCitation captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Gymnopus sublaccatus. A Subbasidial hyphae appearing beaded B Effete hymenial structures without collapse (&quot; husking &quot;) C Stipe medullary hyphae. Standard bars = 10 µm. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure81" httpUri="https://binary.pensoft.net/fig/115146" pageId="98" pageNumber="99">81B</figureCitation>
); subbasidial hyphae (Fig. 
<figureCitation captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Gymnopus sublaccatus. A Subbasidial hyphae appearing beaded B Effete hymenial structures without collapse (&quot; husking &quot;) C Stipe medullary hyphae. Standard bars = 10 µm. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure81" httpUri="https://binary.pensoft.net/fig/115146" pageId="98" pageNumber="99">81A</figureCitation>
) rupturing at clamp connections, appearing beaded. 
<emphasis bold="true" pageId="98" pageNumber="99">Basidiospores</emphasis>
(Fig. 
<figureCitation captionStart="Figure 78" captionStartId="F78" captionText="Figure 78. Gymnopus sublaccatus. A Basidiomata showing adventitious rhizomorphs and stipe insertion to pileus B Basidiospores C Substrate scales with back spots and rhizomorphs. Standard bars: A = 20 mm; B = 5 µm; C = not to scale. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure78" httpUri="https://binary.pensoft.net/fig/115142" pageId="98" pageNumber="99">78B</figureCitation>
) (6.5-)7-7.5(-8) 
<normalizedToken originalValue="×">x</normalizedToken>
(3.5-)4-5 
<normalizedToken originalValue="µm">µm</normalizedToken>
(Q = 1.40-1.88; Qm = 1.67; Lm = 7.20 
<normalizedToken originalValue="µm">µm</normalizedToken>
), rotund-ellipsoid, smooth, thin-walled, inamyloid. 
<emphasis bold="true" pageId="98" pageNumber="99">Cheilocystidia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Gymnopus sublaccatus. Cheilocystidia. Standard bars = 10 µm. UBS 15356." figureDoi="10.3897/mycokeys.18.10007.figure82" httpUri="https://binary.pensoft.net/fig/115147" pageId="98" pageNumber="99">82</figureCitation>
) very locally common, usually rare to absent, 14-22 
<normalizedToken originalValue="×">x</normalizedToken>
8-12 
<normalizedToken originalValue="µm">µm</normalizedToken>
, clavate to utriform, thin-walled, obscurely clamped; contents homogeneous. 
<emphasis bold="true" pageId="98" pageNumber="99">Stipe medullary hyphae</emphasis>
(Fig. 
<figureCitation captionStart="Figure 81" captionStartId="F81" captionText="Figure 81. Gymnopus sublaccatus. A Subbasidial hyphae appearing beaded B Effete hymenial structures without collapse (&quot; husking &quot;) C Stipe medullary hyphae. Standard bars = 10 µm. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure81" httpUri="https://binary.pensoft.net/fig/115146" pageId="98" pageNumber="99">81C</figureCitation>
) 4-12 
<normalizedToken originalValue="µm">µm</normalizedToken>
diam, strictly parallel, apparently free (no discernable gelatinized matrix, hyaline, thick-walled (wall -0.7 
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, hyaline, non-gelatinized), obscurely clamped, often with small side-branches ranging from lobate to rudimentarily branched. 
<emphasis bold="true" pageId="98" pageNumber="99">Stipe cortical hyphae</emphasis>
4-8 
<normalizedToken originalValue="µm">µm</normalizedToken>
diam, strictly parallel, apparently free, thick-walled (wall -1.2 
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, especially at stipe surface), pigmented (yellow-brown, KOH + PhC), appearing moderately dextrinoid (IKI + PhC), or not so (IKI + BF), often producing broad-based side branches. 
<emphasis bold="true" pageId="98" pageNumber="99">Caulocystidia</emphasis>
(Figs 
<figureCitation captionStart="Figure 83" captionStartId="F83" captionText="Figure 83. Gymnopus sublaccatus. Caulocystidia from stipe apex. Standard bars = 10 µm. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure83" httpUri="https://binary.pensoft.net/fig/115148" pageId="98" pageNumber="99">83</figureCitation>
, 
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Gymnopus sublaccatus. Caulocystidia from stipe base. Standard bars = 10 µm. UBC 25212." figureDoi="10.3897/mycokeys.18.10007.figure84" httpUri="https://binary.pensoft.net/fig/115149" pageId="98" pageNumber="99">84</figureCitation>
) -45 
<normalizedToken originalValue="µm">µm</normalizedToken>
at stipe apex, -140 
<normalizedToken originalValue="µm">µm</normalizedToken>
<pageBreakToken pageId="99" pageNumber="100" start="start">at</pageBreakToken>
stipe base, 
<normalizedToken originalValue="×">x</normalizedToken>
5.5-10 
<normalizedToken originalValue="µm">µm</normalizedToken>
, ranging from lobate to elongate-digitate, without basal clamp, often internally secondarily septate, sometimes branched in one rank, thick-walled [wall -4 
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, often occluding cell lumen, pigmented (yellow-brown PhC)].
</paragraph>
<caption ID-Zenodo-Dep="984937" doi="10.3897/mycokeys.18.10007.figure79" httpUri="https://binary.pensoft.net/fig/115144" pageId="99" pageNumber="100" start="Figure 79" startId="F79">
<paragraph pageId="99" pageNumber="100">
<emphasis bold="true" pageId="99" pageNumber="100">Figure 79.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. Pileipellis elements. 
<emphasis bold="true" pageId="99" pageNumber="100">A-E</emphasis>
Pileal hairs 
<emphasis bold="true" pageId="99" pageNumber="100">F</emphasis>
Secondary septa of pileipellis hypha. Standard bars = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
. UBC 25212.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="984941" doi="10.3897/mycokeys.18.10007.figure80" httpUri="https://binary.pensoft.net/fig/115145" pageId="99" pageNumber="100" start="Figure 80" startId="F80">
<paragraph pageId="99" pageNumber="100">
<emphasis bold="true" pageId="99" pageNumber="100">Figure 80.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. Hymenial elements. 
<emphasis bold="true" pageId="99" pageNumber="100">A-D</emphasis>
Pleurocystidia 
<emphasis bold="true" pageId="99" pageNumber="100">E</emphasis>
Basidiole 
<emphasis bold="true" pageId="99" pageNumber="100">F-H</emphasis>
Basidia. Standard bars = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
. UBC 15356.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="984943" doi="10.3897/mycokeys.18.10007.figure81" httpUri="https://binary.pensoft.net/fig/115146" pageId="99" pageNumber="100" start="Figure 81" startId="F81">
<paragraph pageId="99" pageNumber="100">
<emphasis bold="true" pageId="99" pageNumber="100">Figure 81.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="99" pageNumber="100">A</emphasis>
Subbasidial hyphae appearing beaded 
<emphasis bold="true" pageId="99" pageNumber="100">B</emphasis>
Effete hymenial structures without collapse (
<normalizedToken originalValue="“husking”">&quot;husking&quot;</normalizedToken>
) 
<emphasis bold="true" pageId="99" pageNumber="100">C</emphasis>
Stipe medullary hyphae. Standard bars = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
. UBC 25212.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="984945" doi="10.3897/mycokeys.18.10007.figure82" httpUri="https://binary.pensoft.net/fig/115147" pageId="99" pageNumber="100" start="Figure 82" startId="F82">
<paragraph pageId="99" pageNumber="100">
<emphasis bold="true" pageId="99" pageNumber="100">Figure 82.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. Cheilocystidia. Standard bars = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
. UBS 15356.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="984947" doi="10.3897/mycokeys.18.10007.figure83" httpUri="https://binary.pensoft.net/fig/115148" pageId="99" pageNumber="100" start="Figure 83" startId="F83">
<paragraph pageId="99" pageNumber="100">
<emphasis bold="true" pageId="99" pageNumber="100">Figure 83.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. Caulocystidia from stipe apex. Standard bars = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
. UBC 25212.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="984949" doi="10.3897/mycokeys.18.10007.figure84" httpUri="https://binary.pensoft.net/fig/115149" pageId="99" pageNumber="100" start="Figure 84" startId="F84">
<paragraph pageId="99" pageNumber="100">
<emphasis bold="true" pageId="99" pageNumber="100">Figure 84.</emphasis>
<taxonomicName authorityName="R. H. Petersen" authorityYear="2016" class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Gymnopus sublaccatus" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">Gymnopus sublaccatus</emphasis>
</taxonomicName>
. Caulocystidia from stipe base. Standard bars = 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
. UBC 25212.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100" type="comments">
<paragraph pageId="99" pageNumber="100">Commentary.</paragraph>
<paragraph pageId="99" pageNumber="100">
With slime matrix covering the pileus surface and involving hymenial structures, with characteristic clavate cheilocystidia and with vestured stipe, UBC 25212 seems certain to belong in sect. 
<taxonomicName class="Agaricomycetes" family="Omphalotaceae" genus="Gymnopus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Agaricales" pageId="99" pageNumber="100" phylum="Basidiomycota" rank="section" section="Perforantia">
<emphasis italics="true" pageId="99" pageNumber="100">Perforantia</emphasis>
</taxonomicName>
. There it joins 
<taxonomicName genus="G." lsidName="G. perforans" pageId="99" pageNumber="100" rank="species" species="perforans">
<emphasis italics="true" pageId="99" pageNumber="100">G. perforans</emphasis>
</taxonomicName>
, 
<taxonomicName genus="G." lsidName="G. foliiphilus" pageId="99" pageNumber="100" rank="species" species="foliiphilus">
<emphasis italics="true" pageId="99" pageNumber="100">G. foliiphilus</emphasis>
</taxonomicName>
and 
<taxonomicName genus="G." lsidName="G. sequoiae" pageId="99" pageNumber="100" rank="species" species="sequoiae">
<emphasis italics="true" pageId="99" pageNumber="100">G. sequoiae</emphasis>
</taxonomicName>
with the same general characters. From 
<taxonomicName genus="Ma." lsidName="Ma. androsaceus" pageId="99" pageNumber="100" rank="species" species="androsaceus">
<emphasis italics="true" pageId="99" pageNumber="100">Ma. androsaceus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="G." lsidName="G. sublaccatus" pageId="99" pageNumber="100" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">G. sublaccatus</emphasis>
</taxonomicName>
differs in pigmented (not black), vestured stipe. If its substrate preference is limited to 
<taxonomicName authorityName="E.A.Carriere" authorityYear="1867" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pseudotsuga" order="Pinales" pageId="99" pageNumber="100" phylum="Coniferophyta" rank="genus">
<emphasis italics="true" pageId="99" pageNumber="100">Pseudotsuga</emphasis>
</taxonomicName>
, this constitutes another difference. Finally, pileipellis of 
<taxonomicName genus="G." lsidName="G. androsaceus" pageId="99" pageNumber="100" rank="species" species="androsaceus">
<emphasis italics="true" pageId="99" pageNumber="100">G. androsaceus</emphasis>
</taxonomicName>
is characterized by diverticulate hyphal segments and broom cell-like hyphal termini, unlike that of 
<taxonomicName genus="G." lsidName="G. sublaccatus" pageId="99" pageNumber="100" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">G. sublaccatus</emphasis>
</taxonomicName>
which lacks these structures.
</paragraph>
<paragraph lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100">
Basidiomata of 
<taxonomicName genus="G." lsidName="G. sequoiae" pageId="99" pageNumber="100" rank="species" species="sequoiae">
<emphasis italics="true" pageId="99" pageNumber="100">G. sequoiae</emphasis>
</taxonomicName>
resemble those of 
<taxonomicName genus="M." lsidName="M. sublaccatus" pageId="99" pageNumber="100" rank="species" species="sublaccatus">
<emphasis italics="true" pageId="99" pageNumber="100">M. sublaccatus</emphasis>
</taxonomicName>
in stature and size, but seems limited to fruiting on needles of 
<taxonomicName class="Coniferopsida" family="Taxodiaceae" genus="Sequoia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sequoia sempervirens" order="Pinales" pageId="99" pageNumber="100" phylum="Gymnospermae" rank="species" species="sempervirens">
<emphasis italics="true" pageId="99" pageNumber="100">Sequoia sempervirens</emphasis>
</taxonomicName>
in northern California. Cheilocystidia are shaped like an incandescent bulb, sometimes slightly askew, but otherwise quite typical of cheilocystidia in this taxonomic complex (compare, for example, 
<pageBreakToken pageId="100" pageNumber="101" start="start">those</pageBreakToken>
of 
<taxonomicName genus="G." lsidName="G. bulliformis" pageId="100" pageNumber="101" rank="species" species="bulliformis">
<emphasis italics="true" pageId="100" pageNumber="101">G. bulliformis</emphasis>
</taxonomicName>
and of 
<taxonomicName genus="G." lsidName="G. sequoiae" pageId="100" pageNumber="101" rank="species" species="sequoiae">
<emphasis italics="true" pageId="100" pageNumber="101">G. sequoiae</emphasis>
</taxonomicName>
). Difficult to distinguish from immature basidia, such cheilocystidia are less obscure after several sightings.
</paragraph>
</subSubSection>
<subSubSection pageId="100" pageNumber="101" type="materials_examined">
<paragraph pageId="100" pageNumber="101">Specimens examined.</paragraph>
<paragraph pageId="100" pageNumber="101">
<materialsCitation accessionNumber="F18168" collectingDate="2006-09-08" collectorName="PK Kroeger" country="Canada" county="Queen Charlotte Islands" latitude="52.409725" location="Section Cove" longLatPrecision="17" longitude="-131.33194" municipality="Burnaby Island" specimenCount="1" stateProvince="British Columbia">
<emphasis bold="true" pageId="100" pageNumber="101">
<collectingCountry name="Canada">Canada</collectingCountry>
</emphasis>
, 
<emphasis bold="true" pageId="100" pageNumber="101">
<collectingRegion country="Canada" name="British Columbia">British Columbia</collectingRegion>
</emphasis>
, 
<collectingCounty>Queen Charlotte Islands</collectingCounty>
, 
<collectingMunicipality>Burnaby Island</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:217172DB856A5B3687DF4E8B15784A37:49821BCB0F9A5EF7DD5CBA51B9908705" country="Canada" county="Queen Charlotte Islands" latitude="52.409725" longLatPrecision="17" longitude="-131.33194" municipality="Burnaby Island" name="Section Cove" stateProvince="British Columbia">Section Cove</location>
, 
<geoCoordinate degrees="52" direction="north" minutes="24" orientation="latitude" precision="15" seconds="35" value="52.409725">N52°24'35&quot;</geoCoordinate>
, 
<geoCoordinate degrees="131" direction="west" minutes="19" orientation="longitude" precision="15" seconds="55" value="-131.33194">W131°19'55&quot;</geoCoordinate>
, 
<collectingDate value="2006-09-08">8.IX.2006</collectingDate>
, coll 
<collectorName>PK Kroeger</collectorName>
(as 
<taxonomicName class="Basidiomycetes" family="Marasmiaceae" genus="Marasmius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Marasmius androsaceus" order="Agaricales" pageId="100" pageNumber="101" phylum="Basidiomycota" rank="species" species="androsaceus">
<emphasis italics="true" pageId="100" pageNumber="101">Marasmius androsaceus</emphasis>
</taxonomicName>
), PK 489 (UBC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F18168">F18168</accessionNumber>
)
</materialsCitation>
; 
<materialsCitation accessionNumber="F16670" collectingDate="2008-09-11" collectorName="P. Kroeger" country="Canada" latitude="52.356388" location="Queen Charlotte Islands" longLatPrecision="17" longitude="-131.40666" specimenCount="1" stateProvince="British Columbia">
<location LSID="urn:lsid:plazi:treatment:217172DB856A5B3687DF4E8B15784A37:FB7E6D1D39E89552DF4336504D7B402E" country="Canada" latitude="52.356388" longLatPrecision="17" longitude="-131.40666" name="Queen Charlotte Islands" stateProvince="British Columbia">Queen Charlotte Islands</location>
, 
<geoCoordinate degrees="52" direction="north" minutes="21" orientation="latitude" precision="15" seconds="23" value="52.356388">N52°21'23&quot;</geoCoordinate>
, 
<geoCoordinate degrees="131" direction="west" minutes="24" orientation="longitude" precision="15" seconds="24" value="-131.40666">W131°24'24&quot;</geoCoordinate>
, 
<collectingDate value="2008-09-11">11.IX.2008</collectingDate>
, coll 
<collectorName>P. Kroeger</collectorName>
(as 
<taxonomicName class="Basidiomycetes" family="Marasmiaceae" genus="Marasmius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Marasmius androsaceus" order="Agaricales" pageId="100" pageNumber="101" phylum="Basidiomycota" rank="species" species="androsaceus">
<emphasis italics="true" pageId="100" pageNumber="101">Marasmius androsaceus</emphasis>
</taxonomicName>
), PK 5904 (UBC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F16670">F16670</accessionNumber>
)
</materialsCitation>
; 
<materialsCitation accessionNumber="F15356" collectingDate="2004-09-10" collectorName="P. Kroeger, B. &amp; C. Kendrick, Brown" country="Canada" latitude="52.15" location="Ellen Island" longLatPrecision="1085" longitude="-131.1" municipality="Queen Charlotte Islands" specimenCount="1" stateProvince="British Columbia">
<collectingMunicipality>Queen Charlotte Islands</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:217172DB856A5B3687DF4E8B15784A37:824606B66E467C037C151CAAA7BC42F0" country="Canada" latitude="52.15" longLatPrecision="1085" longitude="-131.1" municipality="Queen Charlotte Islands" name="Ellen Island" stateProvince="British Columbia">Ellen Island</location>
, 
<geoCoordinate degrees="52" direction="north" minutes="09" orientation="latitude" precision="925" value="52.15">N52°09'</geoCoordinate>
, 
<geoCoordinate degrees="131" direction="west" minutes="06" orientation="longitude" precision="925" value="-131.1">W131°06'</geoCoordinate>
, 
<collectingDate value="2004-09-10">10.IX.2004</collectingDate>
, coll &amp; 
<collectorName>P. Kroeger</collectorName>
, 
<collectorName>B. &amp; C. Kendrick</collectorName>
, J. 
<collectorName>Brown</collectorName>
, (as 
<taxonomicName class="Basidiomycetes" family="Marasmiaceae" genus="Marasmius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Marasmius androsaceus" order="Agaricales" pageId="100" pageNumber="101" phylum="Basidiomycota" rank="species" species="androsaceus">
<emphasis italics="true" pageId="100" pageNumber="101">Marasmius androsaceus</emphasis>
</taxonomicName>
), PK 2932 (UBC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F15356">F15356</accessionNumber>
)
</materialsCitation>
; 
<materialsCitation accessionNumber="F17675" collectingDate="2006-09-02" collectorName="P. Kroeger" country="Canada" latitude="52.166668" location="Ross Island" longLatPrecision="1085" longitude="-131.11667" municipality="Queen Charlotte Islands" specimenCount="1" stateProvince="British Columbia">
<collectingMunicipality>Queen Charlotte Islands</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:217172DB856A5B3687DF4E8B15784A37:76071FF1D66B9056CA15DF29FDB40E0C" country="Canada" latitude="52.166668" longLatPrecision="1085" longitude="-131.11667" municipality="Queen Charlotte Islands" name="Ross Island" stateProvince="British Columbia">Ross Island</location>
, 
<geoCoordinate degrees="52" direction="north" minutes="10" orientation="latitude" precision="925" value="52.166668">N52°10'</geoCoordinate>
, 
<geoCoordinate degrees="131" direction="west" minutes="07" orientation="longitude" precision="925" value="-131.11667">W131°07'</geoCoordinate>
, 
<collectingDate value="2006-09-02">2.IX.2006</collectingDate>
, coll 
<collectorName>P. Kroeger</collectorName>
(as 
<taxonomicName class="Basidiomycetes" family="Marasmiaceae" genus="Marasmius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Marasmius androsaceus" order="Agaricales" pageId="100" pageNumber="101" phylum="Basidiomycota" rank="species" species="androsaceus">
<emphasis italics="true" pageId="100" pageNumber="101">Marasmius androsaceus</emphasis>
</taxonomicName>
), PK 4347 (UBC 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F17675">F17675</accessionNumber>
)
</materialsCitation>
; 
<materialsCitation collectingDate="2011-11-04" collectorName="O. Ceska" country="Canada" county="Victoria" latitude="48.5262" location="Observatory Hill" longLatPrecision="36" longitude="-123.422" municipality="Saanich Peninsula" specimenCount="1" stateProvince="British Columbia" typeStatus="holotype">
<collectingCounty>Victoria</collectingCounty>
, 
<collectingMunicipality>Saanich Peninsula</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:217172DB856A5B3687DF4E8B15784A37:9F727E495B25C2415CD43792DA93EBBE" country="Canada" county="Victoria" latitude="48.5262" longLatPrecision="36" longitude="-123.422" municipality="Saanich Peninsula" name="Observatory Hill" stateProvince="British Columbia">Observatory Hill</location>
, 
<geoCoordinate degrees="48.5262" direction="north" orientation="latitude" precision="5" value="48.5262">N48.5262°</geoCoordinate>
, 
<geoCoordinate degrees="123.422" direction="west" orientation="longitude" precision="55" value="-123.422">W123.422°</geoCoordinate>
, 
<collectingDate value="2011-11-04">4.XI.2011</collectingDate>
, coll &amp; det 
<collectorName>O. Ceska</collectorName>
(as 
<taxonomicName class="Basidiomycetes" family="Marasmiaceae" genus="Marasmius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Marasmius androsaceus" order="Agaricales" pageId="100" pageNumber="101" phylum="Basidiomycota" rank="species" species="androsaceus">
<emphasis italics="true" pageId="100" pageNumber="101">Marasmius androsaceus</emphasis>
</taxonomicName>
), UBC-F-25212 (
<typeStatus>holotype</typeStatus>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>