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<document id="AEB13641F3C86E4790DD0F7E0E201ECA" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFD3FFE0FF04FB3BFD5EFD92" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Denticulobasis dunklei Machado, 2009, sp. nov." docType="treatment" docVersion="12" lastPageNumber="12" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="11" updateTime="1698585892014" updateUser="plazi">
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<mods:title id="9C7AA3C8ADD21DF0F29D9FA162BF2467">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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<mods:namePart id="3DF9B585E3EC11768ED8D3C161A45F95">Machado, Angelo B. M.</mods:namePart>
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<mods:date id="E5FE371942CA0BB8C1F39B9271AFA135">2009</mods:date>
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<mods:number id="C718457FDE83DB7BC387947638A2800E">2108</mods:number>
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<mods:identifier id="050CA9A79B28A015D431190E32EE8F9A" type="DOI">10.5281/zenodo.187806</mods:identifier>
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<treatment id="1312774BFFD3FFE0FF04FB3BFD5EFD92" ID-DOI="http://doi.org/10.5281/zenodo.4391355" ID-GBIF-Taxon="119617248" ID-Zenodo-Dep="4391355" LSID="urn:lsid:plazi:treatment:1312774BFFD3FFE0FF04FB3BFD5EFD92" httpUri="http://treatment.plazi.org/id/1312774BFFD3FFE0FF04FB3BFD5EFD92" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="D3A195D6FFD3FFE1FF04FB3BFDDFFABF" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="9B04C65DFFD3FFE1FF04FB3BFDB6FB50" blockId="10.[151,632,1244,1305]" box="[151,529,1244,1270]" pageId="10" pageNumber="11">
<heading id="C04C7131FFD3FFE1FF04FB3BFDB6FB50" bold="true" box="[151,529,1244,1270]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="A9CF1A4FFFD3FFE1FF04FB3BFDB6FB50" bold="true" box="[151,529,1244,1270]" pageId="10" pageNumber="11">
<taxonomicName id="5CBBBDDEFFD3FFE1FF04FB3BFE0BFB50" ID-CoL="34TX3" box="[151,428,1244,1270]" class="Insecta" family="Coenagrionidae" genus="Denticulobasis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dunklei" status="sp. nov.">
<emphasis id="A9CF1A4FFFD3FFE1FF04FB3BFE0BFB50" bold="true" box="[151,428,1244,1270]" italics="true" pageId="10" pageNumber="11">Denticulobasis dunklei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFD3FFE1FE27FB3BFDB6FB50" box="[436,529,1244,1270]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B04C65DFFD3FFE1FF04FB18FDDFFABF" blockId="10.[151,632,1244,1305]" box="[151,632,1279,1305]" pageId="10" pageNumber="11">
<figureCitation id="0380DAD8FFD3FFE1FF04FB18FEA1FABF" box="[151,262,1279,1305]" captionStart="FIGURE 2" captionStartId="4.[151,255,1888,1912]" captionTargetBox="[479,1102,1096,1850]" captionTargetId="figure@4.[472,1115,1084,1865]" captionTargetPageId="4" captionText="FIGURE 2. Ovipositors: denticulated part of ventral border of lateral valve. SEM: (a) Denticulobasis dunklei in lateral view; (b) T. williamsoni ventro-lateral view showing borders of two valves; (c) L. vacillans in lateral view." httpUri="https://zenodo.org/record/187808/files/figure.png" pageId="10" pageNumber="11">Figures 2</figureCitation>
a, 4, 5, 9, 12, 14, 16, 20, 21, 87.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFD3FFE1FF04FAABFCE2FA2B" pageId="10" pageNumber="11" type="etymology">
<paragraph id="9B04C65DFFD3FFE1FF04FAABFCE2FA2B" blockId="10.[151,1437,1356,2001]" pageId="10" pageNumber="11">
<emphasis id="A9CF1A4FFFD3FFE1FF04FAABFE93FAC0" bold="true" box="[151,308,1356,1382]" pageId="10" pageNumber="11">Etymology:</emphasis>
Named after my colleague Sidney W. Dunkle, who collected the specimens in
<collectingCountry id="E3AC86CDFFD3FFE1FAA4FAABFAD2FAC0" box="[1335,1397,1356,1382]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
, in recognition of his important contributions to Odonatology.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFD3FFE1FF55FA7EFDF0F9A7" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="9B04C65DFFD3FFE1FF55FA7EFDF0F9A7" blockId="10.[151,1437,1356,2001]" pageId="10" pageNumber="11">
<emphasis id="A9CF1A4FFFD3FFE1FF55FA7EFDB0FA15" bold="true" box="[198,535,1433,1459]" pageId="10" pageNumber="11">
<typeStatus id="440078FFFFD3FFE1FF55FA7EFEA4FA15" box="[198,259,1433,1459]" pageId="10" pageNumber="11">Type</typeStatus>
specimens:
<typeStatus id="440078FFFFD3FFE1FE36FA7EFDB0FA15" box="[421,535,1433,1459]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
(3) and
<emphasis id="A9CF1A4FFFD3FFE1FD1CFA7EFD55FA15" bold="true" box="[655,754,1433,1459]" pageId="10" pageNumber="11">allotype</emphasis>
(Ƥ).
<collectingCountry id="E3AC86CDFFD3FFE1FCAFFA7DFC2DFA12" box="[828,906,1434,1460]" name="Peru" pageId="10" pageNumber="11">PERU</collectingCountry>
, Loreto Department. Explorama Lodge,
<quantity id="5C436BB8FFD3FFE1FAEDFA7DFF70FA7C" metricMagnitude="4" metricUnit="m" metricValue="8.04672" pageId="10" pageNumber="11" unit="mi" value="50.0">50 miles</quantity>
NE of Iquitos on Amazon River at the junction with Yanomano River (
<geoCoordinate id="FE8FA09AFFD3FFE1FBBCFA27FB15FA7C" box="[1071,1202,1472,1498]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-3.3663888">3º21´59´´S</geoCoordinate>
,
<geoCoordinate id="FE8FA09AFFD3FFE1FB53FA27FAF8FA7C" box="[1216,1375,1472,1498]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-72.79889">72º47´56´´W</geoCoordinate>
),
<date id="EF05E09DFFD3FFE1FAE7FA27FEB7F9A7" pageId="10" pageNumber="11" value="1989-08-24">24- VIII-1989</date>
, S.W. Dunkley leg.(
<collectionCode id="FDAA5E98FFD3FFE1FE69FA00FDECF9A7" box="[506,587,1511,1537]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="10" pageNumber="11">FSCA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFD3FFE0FF55F9EAFD5EFD92" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="9B04C65DFFD3FFE1FF55F9EAFDB0F981" blockId="10.[151,1437,1356,2001]" box="[198,535,1549,1576]" pageId="10" pageNumber="11">
<emphasis id="A9CF1A4FFFD3FFE1FF55F9EAFEF9F981" bold="true" box="[198,350,1549,1575]" pageId="10" pageNumber="11">Description.</emphasis>
Male
<typeStatus id="440078FFFFD3FFE1FE3AF9E9FDA8F98E" box="[425,527,1550,1576]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9B04C65DFFD3FFE1FF55F9D3FF4BF93A" blockId="10.[151,1437,1356,2001]" pageId="10" pageNumber="11">
<emphasis id="A9CF1A4FFFD3FFE1FF55F9D3FEAEF9E8" bold="true" box="[198,265,1588,1614]" pageId="10" pageNumber="11">Head</emphasis>
. Labium whitish yellow. Labrum, base of mandibles, and genae greyish brown. Anteclypeus, postclypeus, frons, antennae, and top of head brown with large blue postocular spots. Rear of head whitish yellow.
</paragraph>
<paragraph id="9B04C65DFFD3FFE1FF55F94FFA3BF822" blockId="10.[151,1437,1356,2001]" pageId="10" pageNumber="11">
<emphasis id="A9CF1A4FFFD3FFE1FF55F94FFE8DF964" bold="true" box="[198,298,1704,1730]" pageId="10" pageNumber="11">Thorax.</emphasis>
Prothorax: Propleuron and anterior lobe light blue. Pronotum light brown. Pterothorax: mesopleuron dominantly yellowish brown. Horns and area between them black, except at their latero-basal region, which is bluish. Metepisternum yellowish brown and metepimeron yellowish white. Three blue stripes; antehumeral, metepisternal, and a poorly defined metepimeral. Legs yellow. Wings hyaline, venation brown, pterostigma brownish yellow surrounded by yellow. Px in FW
<quantity id="5C436BB8FFD3FFE1FC44F8A4FB98F8FB" box="[983,1087,1859,1885]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.302" metricValueMin="3.048" pageId="10" pageNumber="11" unit="in" value="12.5" valueMax="13.0" valueMin="12.0">1213 in</quantity>
HW 11. R3 arising near Px
<quantity id="5C436BB8FFD3FFE1FA1EF8A4FF09F822" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="11" unit="in" value="6.0">6 in</quantity>
FW and
<quantity id="5C436BB8FFD3FFE1FE8AF88DFEE1F822" box="[281,326,1898,1924]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="11" unit="in" value="5.0">5 in</quantity>
HW. Petiolation distal to Ac by a distance 1.5 times length of Ac in FW and equal to it in HW.
</paragraph>
<paragraph id="9B04C65DFFD3FFE0FF55F877FBFCFF14" blockId="10.[151,1437,1356,2001]" lastBlockId="11.[151,1437,152,564]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="A9CF1A4FFFD3FFE1FF55F877FEEFF80C" bold="true" box="[198,328,1936,1962]" pageId="10" pageNumber="11">Abdomen.</emphasis>
S1 and S2 dorsally brown, laterally light blue. S36 dorsally brown, laterally yellow with black ring at distal 1/10. S7 dorsally reddish brown, laterally yellow with black ring at proximal ¼. S810 dorsally brownish yellow, laterally yellow. Cercus and paraproct brownish yellow.
</paragraph>
<paragraph id="9B04C65DFFD2FFE0FF55FF59FE08FDA8" blockId="11.[151,1437,152,564]" pageId="11" pageNumber="12">
<emphasis id="A9CF1A4FFFD2FFE0FF55FF59FE74FF7E" bold="true" box="[198,467,190,216]" pageId="11" pageNumber="12">Structural characters</emphasis>
.
<collectingCountry id="E3AC86CDFFD2FFE0FE77FF59FDB8FF7E" box="[484,543,190,216]" name="India" pageId="11" pageNumber="12">Hind</collectingCountry>
prothoracic lobe (
<figureCitation id="0380DAD8FFD2FFE0FD69FF59FCE5FF7E" box="[762,834,190,216]" captionStart="FIGURES 4 11" captionStartId="6.[151,273,1897,1921]" captionTargetBox="[257,1330,190,1837]" captionTargetId="figure@6.[211,1375,151,1877]" captionTargetPageId="6" captionText="FIGURES 4 11. Denticulobasis, male thorax in dorsal and lateral view: D. dunklei, holotype (4, 5); D. garrisoni, holotype (6, 7); female thorax in dorsal view: D. ariken, holotype (8); D. dunklei, allotype (9); and D. garrisoni, allotype (10); female S 8 10 in lateral view, D. garrisoni, allotype (11)." httpUri="https://zenodo.org/record/187810/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
) with lateral lobes minute, median lobe projecting caudally, subquadrangular and two-lipped; ventral lip continuous, and dorsal lip reduced to two flaps separated by an incision at mid-line. Mesepisternal horns in dorsal view (
<figureCitation id="0380DAD8FFD2FFE0FB8AFEEBFBC4FE80" box="[1049,1123,268,294]" captionStart="FIGURES 4 11" captionStartId="6.[151,273,1897,1921]" captionTargetBox="[257,1330,190,1837]" captionTargetId="figure@6.[211,1375,151,1877]" captionTargetPageId="6" captionText="FIGURES 4 11. Denticulobasis, male thorax in dorsal and lateral view: D. dunklei, holotype (4, 5); D. garrisoni, holotype (6, 7); female thorax in dorsal view: D. ariken, holotype (8); D. dunklei, allotype (9); and D. garrisoni, allotype (10); female S 8 10 in lateral view, D. garrisoni, allotype (11)." httpUri="https://zenodo.org/record/187810/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
) strongly diverging, each tapering to a blunt tip, mostly smooth with a transversely ridged portion at distal 1/5. Cercus in dorsal view slightly wider at apex than at base (excluding distal lobe) (
<figureCitation id="0380DAD8FFD2FFE0FCFCFEBEFC6EFED5" box="[879,969,345,371]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1977,2001]" captionTargetBox="[186,1411,190,1976]" captionTargetId="figure@7.[176,1420,183,2013]" captionTargetPageId="7" captionText="FIGURES 12 17. Denticulobasis, male anal appendages in dorsal, lateral, and ventro-posterior view in holotypes of D." httpUri="https://zenodo.org/record/187811/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
) with a whitish shallow subtriangular membranous depression at base (
<figureCitation id="0380DAD8FFD2FFE0FDBEFE67FD2FFE3C" box="[557,648,384,410]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1977,2001]" captionTargetBox="[186,1411,190,1976]" captionTargetId="figure@7.[176,1420,183,2013]" captionTargetPageId="7" captionText="FIGURES 12 17. Denticulobasis, male anal appendages in dorsal, lateral, and ventro-posterior view in holotypes of D." httpUri="https://zenodo.org/record/187811/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
). In ventro-posterior view (
<figureCitation id="0380DAD8FFD2FFE0FC48FE67FB93FE3C" box="[987,1076,384,410]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1977,2001]" captionTargetBox="[186,1411,190,1976]" captionTargetId="figure@7.[176,1420,183,2013]" captionTargetPageId="7" captionText="FIGURES 12 17. Denticulobasis, male anal appendages in dorsal, lateral, and ventro-posterior view in holotypes of D." httpUri="https://zenodo.org/record/187811/files/figure.png" pageId="11" pageNumber="12">Fig. 16</figureCitation>
) cercus is spoon-shaped, and continues apically as a distal lobe. A ventral transverse digitiform structure present at limit between main portion and distal lobe of cercus (
<figureCitation id="0380DAD8FFD2FFE0FDA1FE2AFD2CFE41" box="[562,651,461,487]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1977,2001]" captionTargetBox="[186,1411,190,1976]" captionTargetId="figure@7.[176,1420,183,2013]" captionTargetPageId="7" captionText="FIGURES 12 17. Denticulobasis, male anal appendages in dorsal, lateral, and ventro-posterior view in holotypes of D." httpUri="https://zenodo.org/record/187811/files/figure.png" pageId="11" pageNumber="12">Fig. 16</figureCitation>
) which is not visible in dorsal view (
<figureCitation id="0380DAD8FFD2FFE0FBC0FE2AFB08FE41" box="[1107,1199,461,487]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1977,2001]" captionTargetBox="[186,1411,190,1976]" captionTargetId="figure@7.[176,1420,183,2013]" captionTargetPageId="7" captionText="FIGURES 12 17. Denticulobasis, male anal appendages in dorsal, lateral, and ventro-posterior view in holotypes of D." httpUri="https://zenodo.org/record/187811/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
). Penis (
<figureCitation id="0380DAD8FFD2FFE0FA8AFE2AFAD6FE41" box="[1305,1393,461,487]" captionStart="FIGURES 87 94" captionStartId="29.[151,269,1856,1880]" captionTargetBox="[282,1311,194,1855]" captionTargetId="figure@29.[273,1313,117,1921]" captionTargetPageId="29" captionText="FIGURES 87 94. Penis in lateral and ectal views: D. dunklei, holotype (87); D. garrisoni, holotype (88); T. costalimai, paratype (89); T. geijskesi, holotype (90); T. mammilaris, lectotype (91); T. tirio, paratype (92); T. williamsoni, paratype (93); Leptobasis vacillans, Panamá (94) [91 by R. W. Garrison]." httpUri="https://zenodo.org/record/187820/files/figure.png" pageId="11" pageNumber="12">Fig. 87</figureCitation>
) as described for the genus.
</paragraph>
<paragraph id="9B04C65DFFD2FFE0FF55FDFDFD5EFD92" blockId="11.[151,1437,152,564]" box="[198,761,538,564]" pageId="11" pageNumber="12">
<emphasis id="A9CF1A4FFFD2FFE0FF55FDFDFE0DFD92" bold="true" box="[198,426,538,564]" pageId="11" pageNumber="12">Dimensions (mm).</emphasis>
Abdomen: 32.9; HW 18.82.
</paragraph>
</subSubSection>
</treatment>
</document>