287 lines
35 KiB
XML
287 lines
35 KiB
XML
<document id="A6011C2B9DF330F59975DA0562CFD32D" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFCCFFFDFF04F913FE9BFBF4" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tuberculobasis inversa Selys 1876, comb. nov." docType="treatment" docVersion="12" lastPageNumber="23" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="22" updateTime="1698585892014" updateUser="plazi">
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<mods:title id="07837CBD7274D235D15C93ABFE4BFB3A">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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<mods:namePart id="5467F8F4D9F2B9C8AAEE56E841093105">Machado, Angelo B. M.</mods:namePart>
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<mods:title id="B65EBE320862FA34BAFFF9503CFC4D6A">Zootaxa</mods:title>
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<mods:part id="6816BA7E5C2B2576072628DB491C6A0E">
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<mods:date id="046027E3635D09BB5151EB7FA5C7C594">2009</mods:date>
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<mods:detail id="D6DD62D5F6B4097579AADA4EA3BDF8FD" type="volume">
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<mods:number id="4760A10D36E34F4FA6D6C33A6E7537A1">2108</mods:number>
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<mods:identifier id="3F0311C35E56AD86CB9FFE22F3C5FA79" type="DOI">10.5281/zenodo.187806</mods:identifier>
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<subSubSection id="D3A195D6FFCCFFFEFF04F913FE48F897" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="9B04C65DFFCCFFFEFF04F913FD78F8A8" blockId="21.[151,735,1780,1841]" box="[151,735,1780,1806]" pageId="21" pageNumber="22">
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<heading id="C04C7131FFCCFFFEFF04F913FD78F8A8" bold="true" box="[151,735,1780,1806]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<emphasis id="A9CF1A4FFFCCFFFEFF04F913FD78F8A8" bold="true" box="[151,735,1780,1806]" pageId="21" pageNumber="22">
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<taxonomicName id="5CBBBDDEFFCCFFFEFF04F913FDF5F8A8" ID-CoL="7CXXK" authority="Selys, 1876" authorityName="Selys" authorityYear="1876" box="[151,594,1780,1806]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="inversa" status="comb. nov.">
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<emphasis id="A9CF1A4FFFCCFFFEFF04F913FE08F8A8" bold="true" box="[151,431,1780,1806]" italics="true" pageId="21" pageNumber="22">Tuberculobasis inversa</emphasis>
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(
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<bibRefCitation id="FF2ABBACFFCCFFFEFE2DF913FDEDF8A8" author="Selys" box="[446,586,1780,1806]" pageId="21" pageNumber="36" refString="Selys-Longchamps, E. de (1876) Synopsis des Agrionines, 5 me legion: Agrion (suite). Le genre Agrion Bulletin de l´Academie royale de Belgique, (2) 41, 247 - 322, 496 - 539, 1233 - 1309 [1 - 282 reprint]." type="journal article" year="1876">Selys, 1876</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="B2FCA734FFCCFFFEFDCAF913FD78F8A8" box="[601,735,1780,1806]" pageId="21" pageNumber="22">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9B04C65DFFCCFFFEFF04F8F0FE48F897" blockId="21.[151,735,1780,1841]" box="[151,495,1815,1841]" pageId="21" pageNumber="22">
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<figureCitation id="0380DAD8FFCCFFFEFF04F8F0FEE5F897" box="[151,322,1815,1841]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="21" pageNumber="22">Figures 31–32</figureCitation>
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,
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<figureCitation id="0380DAD8FFCCFFFEFEDDF8F0FEC9F897" box="[334,366,1815,1841]" captionStart="FIGURES 47 – 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 – 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="21" pageNumber="22">49</figureCitation>
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,
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<figureCitation id="0380DAD8FFCCFFFEFEE9F8F0FE3DF897" box="[378,410,1815,1841]" captionStart="FIGURES 57 – 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 – 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 – 59); T. costalimai, paratypes (60 – 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="21" pageNumber="22">63</figureCitation>
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, 66–68
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A195D6FFCCFFFDFF04F882FD16FEEA" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="reference_group">
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<paragraph id="9B04C65DFFCCFFFEFF04F882FCA6F8DD" blockId="21.[151,1316,1892,2012]" box="[151,769,1892,1916]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="BBABE173FFCCFFFEFF04F882FCA6F8DD" box="[151,769,1892,1916]" pageId="21" pageNumber="22">
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<taxonomicName id="5CBBBDDEFFCCFFFEFF04F882FD8AF8DD" authority="Selys (1876: 969)" authorityName="Selys" authorityPageNumber="969" authorityYear="1876" box="[151,557,1892,1916]" class="Insecta" family="Coenagrionidae" genus="Telagrion" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="inversum">
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<emphasis id="A9CF1A4FFFCCFFFEFF04F882FEC1F8DA" box="[151,358,1893,1916]" italics="true" pageId="21" pageNumber="22">Telagrion inversum</emphasis>
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<treatmentCitation id="1A1AE04CFFCCFFFEFEFFF883FD8AF8DD" author="Selys" box="[364,557,1892,1915]" page="969" pageId="21" pageNumber="22" year="1876">
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<bibRefCitation id="FF2ABBACFFCCFFFEFEFFF883FD8AF8DD" author="Selys" box="[364,557,1892,1915]" pageId="21" pageNumber="36" refString="Selys-Longchamps, E. de (1876) Synopsis des Agrionines, 5 me legion: Agrion (suite). Le genre Agrion Bulletin de l´Academie royale de Belgique, (2) 41, 247 - 322, 496 - 539, 1233 - 1309 [1 - 282 reprint]." type="journal article" year="1876">Selys (1876: 969)</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="1A1AE04CFFCCFFFEFDABF883FD5AF8DD" author="Kirby" box="[568,765,1892,1915]" page="155" pageId="21" pageNumber="22" year="1890">
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<bibRefCitation id="FF2ABBACFFCCFFFEFDABF883FD5AF8DD" author="Kirby" box="[568,765,1892,1915]" pageId="21" pageNumber="36" refString="Kirby, W. F. (1890) A synonymic catalogue of Neuroptera Odonata, or dragonflies. With an appendix of fossil species. Gurney & Jackson, London." type="book" year="1890">Kirby (1890: 155)</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="9B04C65DFFCCFFFEFF04F862FDC4F83D" blockId="21.[151,1316,1892,2012]" box="[151,611,1924,1948]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="BBABE173FFCCFFFEFF04F862FDC4F83D" box="[151,611,1924,1948]" pageId="21" pageNumber="22">
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<taxonomicName id="5CBBBDDEFFCCFFFEFF04F862FDC7F83D" ID-CoL="3TJZS" authority="Santos (1961: 171)" authorityName="Santos" authorityPageNumber="171" authorityYear="1961" box="[151,608,1924,1948]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tuberculata">
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<emphasis id="A9CF1A4FFFCCFFFEFF04F862FE2CF83A" box="[151,395,1925,1948]" italics="true" pageId="21" pageNumber="22">Leptobasis tuberculata</emphasis>
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<treatmentCitation id="1A1AE04CFFCCFFFEFE01F863FDC7F83D" author="Santos" box="[402,608,1924,1947]" page="171" pageId="21" pageNumber="22" year="1961">
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<bibRefCitation id="FF2ABBACFFCCFFFEFE01F863FDC7F83D" author="Santos" box="[402,608,1924,1947]" pageId="21" pageNumber="36" refString="Santos, N. D. (1961) " Leptobasis tuberculata " SP. N. e notas sobre " Leptobasis vacillans " Selys, 1877 (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (2), 171 - 174." type="journal article" year="1961">Santos (1961: 171)</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="9B04C65DFFCCFFFEFF04F842FCE6F81D" blockId="21.[151,1316,1892,2012]" box="[151,833,1956,1980]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="BBABE173FFCCFFFEFF04F842FCE6F81D" box="[151,833,1956,1980]" pageId="21" pageNumber="22">
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<taxonomicName id="5CBBBDDEFFCCFFFEFF04F842FD8BF81D" authority="Santos (1965: 9)" authorityName="Santos" authorityPageNumber="9" authorityYear="1965" box="[151,556,1956,1980]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="inversum">
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<emphasis id="A9CF1A4FFFCCFFFEFF04F842FED4F81A" box="[151,371,1957,1980]" italics="true" pageId="21" pageNumber="22">Leptobasis inversum</emphasis>
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<treatmentCitation id="1A1AE04CFFCCFFFEFEEAF843FD8BF81D" author="Santos" box="[377,556,1956,1979]" page="9" pageId="21" pageNumber="22" year="1965">
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<bibRefCitation id="FF2ABBACFFCCFFFEFEEAF843FD8BF81D" author="Santos" box="[377,556,1956,1979]" pageId="21" pageNumber="36" refString="Santos, N. D. (1965) Contribuicao ao conhecimento dos " odonata " da Regiao de Pocos de Caldas, MG. Minagrion gen. n., para Telagrion mecistogastrum Selys-Longchamp, 1865, com a descricao de uma nova especie (Odonata: Coenagriidae). Atas da Sociedade de Biologia do Rio de Janeiro, 9 (1), 8 - 12." type="journal article" year="1965">Santos (1965: 9)</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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||
;
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<bibRefCitation id="FF2ABBACFFCCFFFEFDABF843FC9AF81D" author="Bridges" box="[568,829,1956,1979]" pageId="21" pageNumber="35" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Bridges, Urbana, 905 pp." type="book" year="1994">Bridges (1994: VII:119)</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="9B04C65DFFCCFFFEFF04F822FA83F87D" blockId="21.[151,1316,1892,2012]" box="[151,1316,1988,2012]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="BBABE173FFCCFFFEFF04F822FA83F87D" box="[151,1316,1988,2012]" pageId="21" pageNumber="22">
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<taxonomicName id="5CBBBDDEFFCCFFFEFF04F822FD2EF87D" authority="Davies & Tobin (1984: 77)" authorityName="Davies & Tobin" authorityPageNumber="77" authorityYear="1984" box="[151,649,1988,2012]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="inversa">
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<emphasis id="A9CF1A4FFFCCFFFEFF04F822FEF8F87A" box="[151,351,1989,2012]" italics="true" pageId="21" pageNumber="22">Leptobasis inversa</emphasis>
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<treatmentCitation id="1A1AE04CFFCCFFFEFEF5F823FD2EF87D" author="Davies" box="[358,649,1988,2011]" page="77" pageId="21" pageNumber="22" year="1984">
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<bibRefCitation id="FF2ABBACFFCCFFFEFEF5F823FD2EF87D" author="Davies" box="[358,649,1988,2011]" pageId="21" pageNumber="35" refString="Davies, D. A. L. & Tobin, P. (1984) The dragonflies of the world: A systematic list of the extant species of Odonata. Vol. 1. Zygoptera, Anisozigoptera Societas Internationalis Odonatologica Rapid Communications (supplement), 3, 1 - 127." type="journal article" year="1984">Davies & Tobin (1984: 77)</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="1A1AE04CFFCCFFFEFD07F823FCCCF87D" author="Garrison" box="[660,875,1988,2011]" page="13" pageId="21" pageNumber="22" year="1991">
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<bibRefCitation id="FF2ABBACFFCCFFFEFD07F823FCCCF87D" author="Garrison" box="[660,875,1988,2011]" pageId="21" pageNumber="35" refString="Garrison, R. W. (1991) A synonymic list of the new world Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison (1991: 13)</bibRefCitation>
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</treatmentCitation>
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||
;
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||
<treatmentCitation id="1A1AE04CFFCCFFFEFCE5F823FB88F87D" author="Tsuda" box="[886,1071,1988,2011]" page="38" pageId="21" pageNumber="22" year="2000">
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<bibRefCitation id="FF2ABBACFFCCFFFEFCE5F823FB88F87D" author="Tsuda" box="[886,1071,1988,2011]" pageId="21" pageNumber="36" refString="Tsuda, S. (2000) A distributional list of world Odonata. Tsuda-Osaka, 430 pp." type="book" year="2000">Tsuda (2000: 38)</bibRefCitation>
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</treatmentCitation>
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||
;
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||
<treatmentCitation id="1A1AE04CFFCCFFFEFBA9F823FAB8F87D" author="Lencioni" box="[1082,1311,1988,2011]" page="156" pageId="21" pageNumber="22" year="2006">
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<bibRefCitation id="FF2ABBACFFCCFFFEFBA9F823FAB8F87D" author="Lencioni" box="[1082,1311,1988,2011]" pageId="21" pageNumber="36" refString="Lencioni, F. A. A. (2006). Damselflies of Brazil. An illustrated identification guide. II. Coenagrionidae, All Print Edit., Sao Paulo, 419 pp." type="book" year="2006">Lencioni (2006: 156)</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="9B04C65DFFCFFFFDFF04FF59FD16FEEA" blockId="22.[151,1437,190,1106]" pageId="22" pageNumber="23">
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||
<emphasis id="A9CF1A4FFFCFFFFDFF04FF59FD9AFF7E" bold="true" box="[151,573,190,216]" pageId="22" pageNumber="23">
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||
Specimens examined:
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||
<typeStatus id="440078FFFFCFFFFDFE51FF59FD9AFF7E" box="[450,573,190,216]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
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||
3.
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<collectingCountry id="E3AC86CDFFCFFFFDFDFDFF59FD47FF7E" box="[622,736,190,216]" name="Brazil" pageId="22" pageNumber="23">BRASIL</collectingCountry>
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||
, Pará State, Santarém (
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,
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<geoCoordinate id="FE8FA09AFFCFFFFDFBE1FF59FB42FF7E" box="[1138,1253,190,216]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-54.7">54º42´W</geoCoordinate>
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||
) (IRSN). 1 3, Amazonas State, Tefé (
|
||
<geoCoordinate id="FE8FA09AFFCFFFFDFE50FF02FDB9FF59" box="[451,542,229,255]" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="-3.3666668">3º22´S</geoCoordinate>
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,
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||
<geoCoordinate id="FE8FA09AFFCFFFFDFDBFFF02FD04FF59" box="[556,675,229,255]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-64.7">64º42´W</geoCoordinate>
|
||
),
|
||
<date id="EF05E09DFFCFFFFDFD29FF02FC95FF59" box="[698,818,229,255]" pageId="22" pageNumber="23" value="1960-12">XII.1960</date>
|
||
, Carvalho leg. (A.B.M.M.). 1 Ƥ,
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||
<collectingCountry id="E3AC86CDFFCFFFFDFB75FF02FA9DFF59" box="[1254,1338,229,255]" name="Peru" pageId="22" pageNumber="23">PERU</collectingCountry>
|
||
, Loreto Department. Explorama Lodge,
|
||
<quantity id="5C436BB8FFCFFFFDFDB2FEEBFD2DFE80" box="[545,650,268,294]" metricMagnitude="4" metricUnit="m" metricValue="8.04672" pageId="22" pageNumber="23" unit="mi" value="50.0">50 miles</quantity>
|
||
NE of Iquitos on Amazon River at the junction with Yanomano River,
|
||
<date id="EF05E09DFFCFFFFDFF76FED5FE26FEEA" box="[229,385,306,332]" pageId="22" pageNumber="23" value="1989-08-30">30.VIII.1989</date>
|
||
, S.W. Dunkle leg (SWD).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A195D6FFCFFFFDFF55FEBFFB1BFED5" box="[198,1212,344,371]" pageId="22" pageNumber="23" type="distribution">
|
||
<paragraph id="9B04C65DFFCFFFFDFF55FEBFFB1BFED5" blockId="22.[151,1437,190,1106]" box="[198,1212,344,371]" pageId="22" pageNumber="23">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFF55FEBFFEF8FED4" bold="true" box="[198,351,344,370]" pageId="22" pageNumber="23">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="E3AC86CDFFCFFFFDFEFDFEBEFE1EFED5" box="[366,441,345,371]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
|
||
, Amazonas, Pará, and Rondônia States.
|
||
<collectingCountry id="E3AC86CDFFCFFFFDFC03FEBEFC6FFED5" box="[912,968,345,371]" name="Peru" pageId="22" pageNumber="23">Peru</collectingCountry>
|
||
, Loreto Department.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A195D6FFCFFFFDFF55FE98FE9BFBF4" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="9B04C65DFFCFFFFDFF55FE98FE9BFBF4" blockId="22.[151,1437,190,1106]" pageId="22" pageNumber="23">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFF55FE98FE9FFE3F" bold="true" box="[198,312,383,409]" pageId="22" pageNumber="23">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFED4FE67FDFFFE3F" box="[327,600,384,409]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFED4FE67FDFFFE3F" box="[327,600,384,409]" italics="true" pageId="22" pageNumber="23">Tuberculobasis inversa</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFCA0FE67FC6FFE3C" author="Selys" box="[819,968,384,410]" pageId="22" pageNumber="36" refString="Selys-Longchamps, E. de (1876) Synopsis des Agrionines, 5 me legion: Agrion (suite). Le genre Agrion Bulletin de l´Academie royale de Belgique, (2) 41, 247 - 322, 496 - 539, 1233 - 1309 [1 - 282 reprint]." type="journal article" year="1876">Selys (1876)</bibRefCitation>
|
||
based on a single male from Santarém, Pará,
|
||
<collectingCountry id="E3AC86CDFFCFFFFDFF48FE41FE80FE66" box="[219,295,422,448]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
|
||
.
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFEA1FE41FE7FFE66" author="Santos" box="[306,472,422,448]" pageId="22" pageNumber="36" refString="Santos, N. D. (1961) " Leptobasis tuberculata " SP. N. e notas sobre " Leptobasis vacillans " Selys, 1877 (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (2), 171 - 174." type="journal article" year="1961">Santos (1961)</bibRefCitation>
|
||
described and illustrated both sexes of
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFC3CFE40FB19FE66" box="[943,1214,423,448]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tuberculata">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFC3CFE40FB19FE66" box="[943,1214,423,448]" italics="true" pageId="22" pageNumber="23">Leptobasis tuberculata</emphasis>
|
||
</taxonomicName>
|
||
from Porto Velho, Rondônia,
|
||
<collectingCountry id="E3AC86CDFFCFFFFDFE8AFE2AFEC6FE41" box="[281,353,461,487]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
|
||
but after examining the
|
||
<typeStatus id="440078FFFFCFFFFDFD14FE2AFD49FE41" box="[647,750,461,487]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFC84FE2AFC2BFE40" box="[791,908,461,486]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFC84FE2AFC2BFE40" box="[791,908,461,486]" italics="true" pageId="22" pageNumber="23">T. inversa</emphasis>
|
||
</taxonomicName>
|
||
synonymized
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFBAAFE2AFB47FE40" box="[1081,1248,461,486]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tuberculata">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFBAAFE2AFB47FE40" box="[1081,1248,461,486]" italics="true" pageId="22" pageNumber="23">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFAB6FE2AFEE0FDA8" authority="Santos, 1965" authorityName="Santos" authorityYear="1965" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFAB6FE2AFA3BFE40" box="[1317,1436,461,486]" italics="true" pageId="22" pageNumber="23">L. inversa</emphasis>
|
||
(
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFF0CFE13FE98FDA8" author="Santos" box="[159,319,500,526]" pageId="22" pageNumber="36" refString="Santos, N. D. (1965) Contribuicao ao conhecimento dos " odonata " da Regiao de Pocos de Caldas, MG. Minagrion gen. n., para Telagrion mecistogastrum Selys-Longchamp, 1865, com a descricao de uma nova especie (Odonata: Coenagriidae). Atas da Sociedade de Biologia do Rio de Janeiro, 9 (1), 8 - 12." type="journal article" year="1965">Santos, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="440078FFFFCFFFFDFE1AFE13FE57FDA8" box="[393,496,500,526]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
lacks head and right wings, but color is well preserved. The descriptions made by
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFF2DFDFDFEFCFD92" author="Selys" box="[190,347,538,564]" pageId="22" pageNumber="36" refString="Selys-Longchamps, E. de (1876) Synopsis des Agrionines, 5 me legion: Agrion (suite). Le genre Agrion Bulletin de l´Academie royale de Belgique, (2) 41, 247 - 322, 496 - 539, 1233 - 1309 [1 - 282 reprint]." type="journal article" year="1876">Selys (1876)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFE0BFDFDFDE3FD92" author="Santos" box="[408,580,538,564]" pageId="22" pageNumber="36" refString="Santos, N. D. (1961) " Leptobasis tuberculata " SP. N. e notas sobre " Leptobasis vacillans " Selys, 1877 (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (2), 171 - 174." type="journal article" year="1961">Santos (1961)</bibRefCitation>
|
||
are adequate. Until recently the only illustrations known for females specimen of
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFEA3FDA6FE04FDFC" box="[304,419,577,602]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFEA3FDA6FE04FDFC" box="[304,419,577,602]" italics="true" pageId="22" pageNumber="23">T. inversa</emphasis>
|
||
</taxonomicName>
|
||
were those provided by
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFD5DFDA6FCD2FDFD" author="Santos" box="[718,885,577,603]" pageId="22" pageNumber="36" refString="Santos, N. D. (1961) " Leptobasis tuberculata " SP. N. e notas sobre " Leptobasis vacillans " Selys, 1877 (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (2), 171 - 174." type="journal article" year="1961">Santos (1961)</bibRefCitation>
|
||
under
|
||
<emphasis id="A9CF1A4FFFCFFFFDFC5AFDA6FBDFFDFC" box="[969,1144,577,602]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFC5AFDA6FBD2FDFC" box="[969,1141,577,602]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tuberculata">L. tuberculata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
I had the opportunity to study and provide an illustration of the thoracic structures of a specimen from
|
||
<collectingCountry id="E3AC86CDFFCFFFFDFBAAFD8FFBC8FD24" box="[1081,1135,616,642]" name="Peru" pageId="22" pageNumber="23">Peru</collectingCountry>
|
||
that agreed well with the specimen described by
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFE3CFD69FDF0FD0E" author="Santos" box="[431,599,654,680]" pageId="22" pageNumber="36" refString="Santos, N. D. (1961) " Leptobasis tuberculata " SP. N. e notas sobre " Leptobasis vacillans " Selys, 1877 (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (2), 171 - 174." type="journal article" year="1961">Santos (1961)</bibRefCitation>
|
||
especially on the unique shape of the hind prothoracic lobe (
|
||
<figureCitation id="0380DAD8FFCFFFFDFAA7FD69FA28FD0E" box="[1332,1423,654,680]" captionStart="FIGURES 47 – 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 – 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="22" pageNumber="23">Fig. 49</figureCitation>
|
||
). R.W. Garrison identified as
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFE73FD52FDF4FD68" box="[480,595,693,718]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFE73FD52FDF4FD68" box="[480,595,693,718]" italics="true" pageId="22" pageNumber="23">T. inversa</emphasis>
|
||
</taxonomicName>
|
||
a male in his collection found in the same month and at the same place where the female illustrated herein had been collected and suggested, and I agree, that it is the female of
|
||
<emphasis id="A9CF1A4FFFCFFFFDFA15FD3BFF52FCBD" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFA15FD3BFF56FCBD" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">T. inversa</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
However, the Peruvian specimen has a pair of small but very evident mesepisternal tubercles that are apparently lacking in the Brazilian specimen described by Santos. I believe these small tubercle might have been overlooked by
|
||
<bibRefCitation id="FF2ABBACFFCFFFFDFE19FCB7FD97FCCC" author="Santos" box="[394,560,848,874]" pageId="22" pageNumber="36" refString="Santos, N. D. (1961) " Leptobasis tuberculata " SP. N. e notas sobre " Leptobasis vacillans " Selys, 1877 (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (2), 171 - 174." type="journal article" year="1961">Santos (1961)</bibRefCitation>
|
||
as he did with the tubercles of the females of
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFBC9FCB7FB54FCCF" box="[1114,1267,848,873]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFBC9FCB7FB54FCCF" box="[1114,1267,848,873]" italics="true" pageId="22" pageNumber="23">T. costalimai</emphasis>
|
||
</taxonomicName>
|
||
(see Remarks under
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFF70FC90FEDAFC36" box="[227,381,887,912]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFF70FC90FEDAFC36" box="[227,381,887,912]" italics="true" pageId="22" pageNumber="23">T. costalimai</emphasis>
|
||
</taxonomicName>
|
||
). Besides, by its male character
|
||
<emphasis id="A9CF1A4FFFCFFFFDFC95FC90FCBCFC36" box="[774,795,887,912]" italics="true" pageId="22" pageNumber="23">T.</emphasis>
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFCB6FC91FCDCFC36" box="[805,891,886,912]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">inversa</taxonomicName>
|
||
belongs in the
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFBA7FC90FB17FC36" box="[1076,1200,887,912]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFBA7FC90FB17FC36" box="[1076,1200,887,912]" italics="true" pageId="22" pageNumber="23">costalimai</emphasis>
|
||
</taxonomicName>
|
||
group in which the known females (
|
||
<emphasis id="A9CF1A4FFFCFFFFDFECEFC7AFD10FC10" box="[349,695,925,950]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFECEFC7AFE5EFC10" box="[349,505,925,950]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
|
||
,
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFD95FC7AFD09FC10" box="[518,686,925,950]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="williamsoni">T. williamsoni</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
have tubercles, whereas the known females (
|
||
<emphasis id="A9CF1A4FFFCFFFFDFB47FC7AFF6DFC7B" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFB47FC7AFADDFC10" box="[1236,1402,925,950]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mammilaris">T.mammilaris</taxonomicName>
|
||
,
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFA15FC7AFF6DFC7B" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirio">T. tirio</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFE92FC23FE3FFC7B" box="[257,408,964,989]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="yanomami">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFE92FC23FE3FFC7B" box="[257,408,964,989]" italics="true" pageId="22" pageNumber="23">T. yanomami</emphasis>
|
||
</taxonomicName>
|
||
) of the
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFE60FC23FDD8FC7B" box="[499,639,964,989]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mammilaris">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFE60FC23FDD8FC7B" box="[499,639,964,989]" italics="true" pageId="22" pageNumber="23">mammilaris</emphasis>
|
||
</taxonomicName>
|
||
group the known females (
|
||
<emphasis id="A9CF1A4FFFCFFFFDFC2DFC23FB63FC7B" box="[958,1220,964,989]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFC2DFC23FBCDFC7B" box="[958,1130,964,989]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mammilaris">T. mammilaris</taxonomicName>
|
||
,
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFBE6FC23FB63FC7B" box="[1141,1220,964,989]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tirio">T. tirio</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFB6FFC23FA35FC7B" box="[1276,1426,964,989]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="yanomami">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFB6FFC23FA35FC7B" box="[1276,1426,964,989]" italics="true" pageId="22" pageNumber="23">T. yanomami</emphasis>
|
||
</taxonomicName>
|
||
) have no tubercles. In the
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFE2EFC0DFD90FBA5" box="[445,567,1002,1027]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFE2EFC0DFD90FBA5" box="[445,567,1002,1027]" italics="true" pageId="22" pageNumber="23">costalimai</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
<taxonomicName id="5CBBBDDEFFCFFFFDFD76FC0DFCFFFBA5" box="[741,856,1002,1027]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCFFFFDFD76FC0DFCFFFBA5" box="[741,856,1002,1027]" italics="true" pageId="22" pageNumber="23">T. inversa</emphasis>
|
||
</taxonomicName>
|
||
can be easily separated from the other species by its subparallel mesepisternal tubercles, with anterior borders connected to mesostigmal plate by a curved plate (
|
||
<figureCitation id="0380DAD8FFCFFFFDFF0CFBDFFE88FBF4" box="[159,303,1080,1106]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="22" pageNumber="23">Figs. 31, 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |