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<document id="61823E0D1F5DA5B897B7B70B5874C3F7" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFC9FFFAFF04FB3DFA3BFE80" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tuberculobasis costalimai Santos 1957, comb. nov." docType="treatment" docVersion="12" lastPageNumber="17" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="17" updateTime="1698585892014" updateUser="plazi">
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<mods:title id="26BD8067FDECF3239BCFC58F5C3866D2">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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<mods:namePart id="B47A1AAF330A84B205B30C9D3A82F09D">Machado, Angelo B. M.</mods:namePart>
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<mods:date id="FBFDB6142CF67686C1C5E64895D16D01">2009</mods:date>
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<treatment id="1312774BFFC9FFFAFF04FB3DFA3BFE80" ID-DOI="http://doi.org/10.5281/zenodo.4391369" ID-GBIF-Taxon="119617256" ID-Zenodo-Dep="4391369" LSID="urn:lsid:plazi:treatment:1312774BFFC9FFFAFF04FB3DFA3BFE80" httpUri="http://treatment.plazi.org/id/1312774BFFC9FFFAFF04FB3DFA3BFE80" lastPageId="17" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="D3A195D6FFC9FFFBFF04FB3DFE51FAB1" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="9B04C65DFFC9FFFBFF04FB3DFCB1FB52" blockId="16.[151,790,1242,1303]" box="[151,790,1242,1268]" pageId="16" pageNumber="17">
<heading id="C04C7131FFC9FFFBFF04FB3DFCB1FB52" bold="true" box="[151,790,1242,1268]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="A9CF1A4FFFC9FFFBFF04FB3DFCB1FB52" bold="true" box="[151,790,1242,1268]" pageId="16" pageNumber="17">
<taxonomicName id="5CBBBDDEFFC9FFFBFF04FB3DFD2EFB52" ID-CoL="7CXXJ" authority="Santos, 1957" authorityName="Santos" authorityYear="1957" box="[151,649,1242,1268]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="costalimai" status="comb. nov.">
<emphasis id="A9CF1A4FFFC9FFFBFF04FB3DFE75FB52" bold="true" box="[151,466,1242,1268]" italics="true" pageId="16" pageNumber="17">Tuberculobasis costalimai</emphasis>
(
<bibRefCitation id="FF2ABBACFFC9FFFBFE72FB3DFD26FB52" author="Santos" box="[481,641,1242,1268]" pageId="16" pageNumber="36" refString="Santos, N. D. (1957) Leptobasis costa-limai sp. nov. (Odonata-Coenagriidae). Boletim do Museu Nacional do Rio de Janeiro, nova serie Zoologia, 157, 1 - 6." type="journal article" year="1957">Santos, 1957</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFC9FFFBFD03FB3DFCB1FB52" box="[656,790,1242,1268]" pageId="16" pageNumber="17">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B04C65DFFC9FFFBFF04FB1AFE51FAB1" blockId="16.[151,790,1242,1303]" box="[151,502,1277,1303]" pageId="16" pageNumber="17">
<figureCitation id="0380DAD8FFC9FFFBFF04FB1AFEE5FAB1" box="[151,322,1277,1303]" captionStart="FIGURES 25 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="16" pageNumber="17">Figures 2728</figureCitation>
,
<figureCitation id="0380DAD8FFC9FFFBFEDDFB1AFEC9FAB1" box="[334,366,1277,1303]" captionStart="FIGURES 47 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="16" pageNumber="17">48</figureCitation>
,
<figureCitation id="0380DAD8FFC9FFFBFEE9FB1AFE61FAB1" box="[378,454,1277,1303]" captionStart="FIGURES 57 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 59); T. costalimai, paratypes (60 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="16" pageNumber="17">6062</figureCitation>
,
<figureCitation id="0380DAD8FFC9FFFBFE41FB1AFE54FAB1" box="[466,499,1277,1303]" captionStart="FIGURES 87 94" captionStartId="29.[151,269,1856,1880]" captionTargetBox="[282,1311,194,1855]" captionTargetId="figure@29.[273,1313,117,1921]" captionTargetPageId="29" captionText="FIGURES 87 94. Penis in lateral and ectal views: D. dunklei, holotype (87); D. garrisoni, holotype (88); T. costalimai, paratype (89); T. geijskesi, holotype (90); T. mammilaris, lectotype (91); T. tirio, paratype (92); T. williamsoni, paratype (93); Leptobasis vacillans, Panamá (94) [91 by R. W. Garrison]." httpUri="https://zenodo.org/record/187820/files/figure.png" pageId="16" pageNumber="17">89</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC9FFFBFF04FAADFCA1F92A" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="9B04C65DFFC9FFFBFF04FAADFCECFA27" blockId="16.[151,1435,1354,1409]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="BBABE173FFC9FFFBFF04FAADFCECFA27" pageId="16" pageNumber="17">
<taxonomicName id="5CBBBDDEFFC9FFFBFF04FAADFDE5FAC7" ID-CoL="9WYD7" authority="Santos (1957: 1)" authorityName="Santos" authorityPageNumber="1" authorityYear="1957" box="[151,578,1354,1377]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="costalimai">
<emphasis id="A9CF1A4FFFC9FFFBFF04FAADFE26FAC7" box="[151,385,1354,1377]" italics="true" pageId="16" pageNumber="17">Leptobasis costalimai</emphasis>
:
<treatmentCitation id="1A1AE04CFFC9FFFBFE1CFAADFDE5FAC7" author="Santos" box="[399,578,1354,1377]" page="1" pageId="16" pageNumber="17" year="1957">
<bibRefCitation id="FF2ABBACFFC9FFFBFE1CFAADFDE5FAC7" author="Santos" box="[399,578,1354,1377]" pageId="16" pageNumber="36" refString="Santos, N. D. (1957) Leptobasis costa-limai sp. nov. (Odonata-Coenagriidae). Boletim do Museu Nacional do Rio de Janeiro, nova serie Zoologia, 157, 1 - 6." type="journal article" year="1957">Santos (1957: 1)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="1A1AE04CFFC9FFFBFDC3FAADFCC0FAC7" author="Davies" box="[592,871,1354,1377]" page="7" pageId="16" pageNumber="17" year="1984">
<bibRefCitation id="FF2ABBACFFC9FFFBFDC3FAADFCC0FAC7" author="Davies" box="[592,871,1354,1377]" pageId="16" pageNumber="35" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The dragonflies of the world: A systematic list of the extant species of Odonata. Vol. 1. Zygoptera, Anisozigoptera Societas Internationalis Odonatologica Rapid Communications (supplement), 3, 1 - 127." type="journal article" year="1984">Davies &amp; Tobin (1984: 7)</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="FF2ABBACFFC9FFFBFCE7FAADFBCAFAC7" author="Bridges" box="[884,1133,1354,1377]" pageId="16" pageNumber="35" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Odonata of the world (Third Edition). Bridges, Urbana, 905 pp." type="book" year="1994">
Bridges (1994:
<date id="EF05E09DFFC9FFFBFB88FAADFBC2FAC7" box="[1051,1125,1354,1377]" pageId="16" pageNumber="17" value="1959-07">VII.59</date>
)
</bibRefCitation>
;
<treatmentCitation id="1A1AE04CFFC9FFFBFBEAFAADFAF7FAC7" author="Garrison" box="[1145,1360,1354,1377]" page="13" pageId="16" pageNumber="17" year="1991">
<bibRefCitation id="FF2ABBACFFC9FFFBFBEAFAADFAF7FAC7" author="Garrison" box="[1145,1360,1354,1377]" pageId="16" pageNumber="35" refString="Garrison, R. W. (1991) A synonymic list of the new world Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison (1991: 13)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1A1AE04CFFC9FFFBFACFFAADFE9EFA27" author="Tsuda" page="38" pageId="16" pageNumber="17" year="2000">
<bibRefCitation id="FF2ABBACFFC9FFFBFACFFAADFE9EFA27" author="Tsuda" pageId="16" pageNumber="36" refString="Tsuda, S. (2000) A distributional list of world Odonata. Tsuda-Osaka, 430 pp." type="book" year="2000">Tsuda (2000: 38)</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="FF2ABBACFFC9FFFBFED6FA8DFDF2FA27" author="Costa" box="[325,597,1386,1409]" pageId="16" pageNumber="35" refString="Costa, J. M. &amp; Oldrini, B. B. (2005). Diversidade e distribuicao dos odonata (Insecta) no estado do Espirito Santo, Brasil. Publicacoes Avulsas do Museu Nacional do Rio de Janeiro, 107, 1 - 15." type="journal article" year="2005">Costa &amp; Oldrini (2005:8)</bibRefCitation>
;
<treatmentCitation id="1A1AE04CFFC9FFFBFDF2FA8DFCE1FA27" author="Lencioni" box="[609,838,1386,1409]" page="155" pageId="16" pageNumber="17" year="2006">
<bibRefCitation id="FF2ABBACFFC9FFFBFDF2FA8DFCE1FA27" author="Lencioni" box="[609,838,1386,1409]" pageId="16" pageNumber="36" refString="Lencioni, F. A. A. (2006). Damselflies of Brazil. An illustrated identification guide. II. Coenagrionidae, All Print Edit., Sao Paulo, 419 pp." type="book" year="2006">Lencioni (2006: 155)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9B04C65DFFC9FFFBFF04FA57FCA1F92A" blockId="16.[151,1437,1456,2024]" pageId="16" pageNumber="17">
<emphasis id="A9CF1A4FFFC9FFFBFF04FA57FE3AFA6C" bold="true" box="[151,413,1456,1482]" pageId="16" pageNumber="17">Specimens examined</emphasis>
:
<emphasis id="A9CF1A4FFFC9FFFBFE3EFA57FD87FA6C" bold="true" box="[429,544,1456,1482]" pageId="16" pageNumber="17">
<typeStatus id="440078FFFFC9FFFBFE3EFA57FD87FA6C" box="[429,544,1456,1482]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
(3) and
<emphasis id="A9CF1A4FFFC9FFFBFD1EFA57FD5FFA6C" bold="true" box="[653,760,1456,1482]" pageId="16" pageNumber="17">Allotype</emphasis>
(Ƥ):
<collectingCountry id="E3AC86CDFFC9FFFBFCAFFA56FC0DFA6D" box="[828,938,1457,1483]" name="Brazil" pageId="16" pageNumber="17">BRAZIL</collectingCountry>
, Espirito Santo State, Conceição da Barra (
<geoCoordinate id="FE8FA09AFFC9FFFBFF33FA3FFEAFFA54" box="[160,264,1496,1522]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="-18.583334">18º35´S</geoCoordinate>
,
<geoCoordinate id="FE8FA09AFFC9FFFBFE86FA3FFE25FA54" box="[277,386,1496,1522]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="-39.75">39º45´W</geoCoordinate>
),
<date id="EF05E09DFFC9FFFBFE04FA3FFE56FA54" box="[407,497,1496,1522]" pageId="16" pageNumber="17" value="1944-10">X-1944</date>
(Ribeirão do Engano, a small and shallow river, 45 meters wide), N.D. Santos leg. (MNRJ).
<emphasis id="A9CF1A4FFFC9FFFBFED7FA19FE76F9BE" bold="true" box="[324,465,1534,1560]" pageId="16" pageNumber="17">
<typeStatus id="440078FFFFC9FFFBFED7FA19FE6BF9BE" box="[324,460,1534,1560]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
:
</emphasis>
20 3, 8 Ƥ same data as
<typeStatus id="440078FFFFC9FFFBFC95FA19FCD1F9BE" box="[774,886,1534,1560]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
. Other specimens: Rio de Janeiro State,
<collectingCountry id="E3AC86CDFFC9FFFBFAFDFA19FF52F999" name="Saint Vincent and the Grenadines" pageId="16" pageNumber="17">São Vicente</collectingCountry>
, flooded area of the São João River,
<date id="EF05E09DFFC9FFFBFD22F9C2FCE7F999" box="[689,832,1573,1599]" pageId="16" pageNumber="17" value="1978-09-16">16-IX-1978</date>
, Santos leg.: 29 3, 6 Ƥ;
<date id="EF05E09DFFC9FFFBFBF8F9C2FAA7F999" box="[1131,1280,1573,1599]" pageId="16" pageNumber="17" value="1980-08-07">7-VIII-1980</date>
, 20 3, 25 Ƥ, Santos &amp; Mesquita leg. (MNRJ); Minas Gerais State, Rio Doce State Park 11,
<date id="EF05E09DFFC9FFFBFBA8F9ABFBF1F9C0" box="[1083,1110,1612,1638]" pageId="16" pageNumber="17" value="1980-12-13">13</date>
-XII-1980 (
<date id="EF05E09DFFC9FFFBFB72F9ABFA94F9C0" box="[1249,1331,1611,1638]" pageId="16" pageNumber="17" value="1935-02-03">35 3 2</date>
Ƥ), data Pereira &amp; Vulcano leg. (ABMM). Total 104 3,
<specimenCount id="8DBD0DD4FFC9FFFBFD56F995FD45F92A" box="[709,738,1650,1676]" pageId="16" pageNumber="17" type="generic">42</specimenCount>
Ƥ.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC9FFFBFF55F97FFB30F915" box="[198,1175,1688,1715]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="9B04C65DFFC9FFFBFF55F97FFB30F915" blockId="16.[151,1437,1456,2024]" box="[198,1175,1688,1715]" pageId="16" pageNumber="17">
<emphasis id="A9CF1A4FFFC9FFFBFF55F97FFEC0F914" bold="true" box="[198,359,1688,1714]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="E3AC86CDFFC9FFFBFEFDF97EFE7CF915" box="[366,475,1689,1715]" name="Brazil" pageId="16" pageNumber="17">BRAZIL</collectingCountry>
, States of Rio de Janeiro, Minas Gerais, and Espírito Santo.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC9FFFAFF55F958FA3BFE80" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="9B04C65DFFC9FFFAFF55F958FA3BFE80" blockId="16.[151,1437,1456,2024]" lastBlockId="17.[151,1437,152,294]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="A9CF1A4FFFC9FFFBFF55F958FEE7F97F" bold="true" box="[198,320,1727,1753]" pageId="16" pageNumber="17">Remarks.</emphasis>
<taxonomicName id="5CBBBDDEFFC9FFFBFED8F927FE41F97F" box="[331,486,1728,1753]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="costalimai">
<emphasis id="A9CF1A4FFFC9FFFBFED8F927FE41F97F" box="[331,486,1728,1753]" italics="true" pageId="16" pageNumber="17">T. costalimai</emphasis>
</taxonomicName>
can be easily identified by its large size (abdomen 3234; HW 1920), male mesepisternal tubercle adjacent to mid-dorsal carina (
<figureCitation id="0380DAD8FFC9FFFBFC9EF901FCC2F8A6" box="[781,869,1766,1792]" captionStart="FIGURES 25 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="16" pageNumber="17">Fig. 27</figureCitation>
), and ventral process of male cercus curved and directed ventro-medially (
<figureCitation id="0380DAD8FFC9FFFBFE5CF8EAFD8DF881" box="[463,554,1805,1831]" captionStart="FIGURES 57 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 59); T. costalimai, paratypes (60 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="16" pageNumber="17">Fig. 62</figureCitation>
). The description of male and female by
<bibRefCitation id="FF2ABBACFFC9FFFBFB8AF8EAFB66F881" author="Santos" box="[1049,1217,1805,1831]" pageId="16" pageNumber="36" refString="Santos, N. D. (1957) Leptobasis costa-limai sp. nov. (Odonata-Coenagriidae). Boletim do Museu Nacional do Rio de Janeiro, nova serie Zoologia, 157, 1 - 6." type="journal article" year="1957">Santos (1957)</bibRefCitation>
is adequate, and I provide here new illustrations of cerci (
<figureCitation id="0380DAD8FFC9FFFBFDFAF8D3FD56F8E8" box="[617,753,1844,1870]" captionStart="FIGURES 57 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 59); T. costalimai, paratypes (60 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="16" pageNumber="17">Figs 6062</figureCitation>
) and thorax (
<figureCitation id="0380DAD8FFC9FFFBFC1CF8D3FBB0F8E8" box="[911,1047,1844,1870]" captionStart="FIGURES 25 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="16" pageNumber="17">Figs 2728</figureCitation>
,
<figureCitation id="0380DAD8FFC9FFFBFBB7F8D3FBF5F8E8" box="[1060,1106,1844,1870]" captionStart="FIGURE 101" captionStartId="32.[151,255,1622,1646]" captionText="FIGURE 101. Distribution map of Denticulobasis and Tuberculobasis." pageId="16" pageNumber="17">101</figureCitation>
a).
<bibRefCitation id="FF2ABBACFFC9FFFBFBE5F8D3FABBF8E8" author="Santos" box="[1142,1308,1844,1870]" pageId="16" pageNumber="36" refString="Santos, N. D. (1957) Leptobasis costa-limai sp. nov. (Odonata-Coenagriidae). Boletim do Museu Nacional do Rio de Janeiro, nova serie Zoologia, 157, 1 - 6." type="journal article" year="1957">Santos (1957)</bibRefCitation>
stated that female thorax does not have the dorsal prominences described for male. Although considerably smaller than in males, tubercles could be detected in all
<specimenCount id="8DBD0DD4FFC9FFFBFD00F866FCB4F83D" box="[659,787,1921,1947]" pageId="16" pageNumber="17" type="female">20 females</specimenCount>
I examined including the allotype and eight specimens of the
<typeStatus id="440078FFFFC9FFFBFF77F84FFEBFF864" box="[228,280,1960,1986]" pageId="16" pageNumber="17">type</typeStatus>
series. In ten specimens (50 %) tubercles were well-developed, in six (30 %) very small, and in four (20%) poorly-defined. Since in all specimens of the female
<typeStatus id="440078FFFFC9FFFBFC2FF829FC56F84E" box="[956,1009,1998,2024]" pageId="16" pageNumber="17">type</typeStatus>
series examined by
<bibRefCitation id="FF2ABBACFFC9FFFBFB63F829FA3AF84E" author="Santos" box="[1264,1437,1998,2024]" pageId="16" pageNumber="36" refString="Santos, N. D. (1957) Leptobasis costa-limai sp. nov. (Odonata-Coenagriidae). Boletim do Museu Nacional do Rio de Janeiro, nova serie Zoologia, 157, 1 - 6." type="journal article" year="1957">Santos (1957)</bibRefCitation>
mesepisternal tubercles could be distinguished it becomes evident that he overlooked these tubercles, certainly much less conspicuous than the &quot;prominences&quot; he had described in the males. In seven females (35%) tubercles were connected with mid-dorsal carina and in 15 (75%) posterior margin of mesostigmal plates curved back and penetrated between tubercle of mesostigmal plate and mesepisternal tubercle (
<figureCitation id="0380DAD8FFC8FFFAFAA4FEEBFA29FE80" box="[1335,1422,268,294]" captionStart="FIGURES 47 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="17" pageNumber="18">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>