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<document id="882E4BDCF4682C8DC6B80B3AB2EAAF09" ID-DOI="10.5281/zenodo.207900" ID-GBIF-Dataset="0686de91-8308-4a42-a495-4c6c976dc3df" ID-ISSN="1175-5326" ID-Zenodo-Dep="207900" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460300107413" checkinUser="plazi" docAuthor="Lazányi, Eszter &amp; Korsós, Zoltán" docDate="2011" docId="03EC87F1FFE1E765FF7FFA52FB9A3B85" docLanguage="en" docName="zt02864p056.pdf" docOrigin="Zootaxa 2864" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Megaphyllum projectum" docType="key" docVersion="9" lastPageNumber="53" masterDocId="FFD5FF89FFEBE76FFFE8FFD1FFB23C14" masterDocTitle="Revision of the Megaphyllum projectum Verhoeff species complex (Myriapoda: Diplopoda: Julida: Julidae)" masterLastPageNumber="56" masterPageNumber="43" pageNumber="53" updateTime="1698252441926" updateUser="plazi">
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<mods:title id="D0C5DC0CAFEB0C9C8BB884D7D958CBBE">Revision of the Megaphyllum projectum Verhoeff species complex (Myriapoda: Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="ED1060EF24108AC19002557ED1A802C8">Lazányi, Eszter</mods:namePart>
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<treatment id="03EC87F1FFE1E765FF7FFA52FB9A3B85" ID-DOI="http://doi.org/10.5281/zenodo.6183759" ID-GBIF-Taxon="119399748" ID-Zenodo-Dep="6183759" LSID="urn:lsid:plazi:treatment:03EC87F1FFE1E765FF7FFA52FB9A3B85" httpUri="http://treatment.plazi.org/id/03EC87F1FFE1E765FF7FFA52FB9A3B85" lastPageNumber="53" pageId="10" pageNumber="53">
<subSubSection id="C35F656CFFE1E765FF7FFA52FE9D3A12" pageId="10" pageNumber="53" type="nomenclature">
<paragraph id="8BFA36E7FFE1E765FF7FFA52FD43398A" blockId="10.[151,753,1411,1438]" box="[151,753,1411,1438]" pageId="10" pageNumber="53">
<heading id="D0B2818BFFE1E765FF7FFA52FD43398A" bold="true" box="[151,753,1411,1438]" fontSize="11" level="1" pageId="10" pageNumber="53" reason="1">
<emphasis id="B931EAF5FFE1E765FF7FFA52FD43398A" bold="true" box="[151,753,1411,1438]" pageId="10" pageNumber="53">
Key to the subspecies of
<taxonomicName id="4C454D64FFE1E765FE20FA55FD43398A" ID-CoL="8KR7G" box="[456,753,1412,1438]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" species="projectum">
<emphasis id="B931EAF5FFE1E765FE20FA55FD43398A" bold="true" box="[456,753,1412,1438]" italics="true" pageId="10" pageNumber="53">Megaphyllum projectum</emphasis>
</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="8BFA36E7FFE1E765FF7FFA1BFE9D3A12" blockId="10.[151,1435,1482,1542]" pageId="10" pageNumber="53">
Based on male and female genitalia characters; female of
<taxonomicName id="4C454D64FFE1E765FCACFA1DFC5239F7" box="[836,992,1483,1507]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="deubeli">
<emphasis id="B931EAF5FFE1E765FCACFA1DFC5239F7" box="[836,992,1483,1507]" italics="true" pageId="10" pageNumber="53">M. p. deubeli</emphasis>
</taxonomicName>
not included because it needs further investigation.
</paragraph>
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<subSubSection id="C35F656CFFE1E765FF7FF9E7FB9A3B85" pageId="10" pageNumber="53" type="key">
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<paragraph id="8BFA36E7FFE1E765FF7FF9E7FA2E3A5F" blockId="10.[151,1436,1590,1937]" box="[151,1436,1590,1611]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF9E7FA2E3A5F" box="[151,1436,1590,1611]" pageId="10" pageNumber="53">
<specimenCount id="9D43FD6EFFE1E765FF7FF9E7FEA53A5F" box="[151,279,1590,1611]" pageId="10" pageNumber="53" type="male">1 Male</specimenCount>
............................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BFA36E7FFE1E765FF7FF985FA2E3A7D" blockId="10.[151,1436,1590,1937]" box="[151,1436,1620,1641]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF985FA2E3A7D" box="[151,1436,1620,1641]" pageId="10" pageNumber="53">- Female.............................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E42FFE1E765FF7FF9A0FA2E3AEF" pageId="10" pageNumber="53">
<paragraph id="8BFA36E7FFE1E765FF7FF9A0FA2E3AB7" blockId="10.[151,1436,1590,1937]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF9A0FA2E3AB7" pageId="10" pageNumber="53">
2 Anterior process of opisthomere rudimentary, shorter than the well-developed posterior process (
<figureCitation id="137E2A62FFE1E765FB60F9A0FB623A92" box="[1160,1232,1649,1670]" captionStart="FIGURES 9 13" captionStartId="7.[151,264,1720,1743]" captionTargetBox="[184,1436,193,1709]" captionTargetId="figure@7.[151,1469,193,1710]" captionTargetPageId="7" captionText="FIGURES 9 13. (9) Left promere of the M. p. deubeli specimen from Bǎile Borşa, Maramureş Mts., Romania, mesal view. (10) Left opisthomere of the same individual, lateral view (scale bars: 0.2 mm). The same opisthomere from (11) caudo-lateral view; (12) lateral view; (13) antero-lateral view (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/207903/files/figure.png" pageId="10" pageNumber="53">Fig. 10</figureCitation>
). Promere triangular, without apical serrated lobe................................................
<taxonomicName id="4C454D64FFE1E765FBC2F95FFA2E3AB7" ID-CoL="8KR5L" authority="Verhoeff, 1899" authorityName="Verhoeff" authorityYear="1899" box="[1066,1436,1678,1699]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="deubeli">
<emphasis id="B931EAF5FFE1E765FBC2F95FFB453AB7" box="[1066,1271,1678,1699]" italics="true" pageId="10" pageNumber="53">M. projectum deubeli</emphasis>
(
<bibRefCitation id="EFD44B16FFE1E765FAEBF95FFA273AB7" author="Verhoeff" box="[1283,1429,1678,1699]" pageId="10" pageNumber="53" refString="Verhoeff, C. (1899) Beitrage zur Kenntniss palaarktischer Myriopoden. IX. Aufsatz: Zur Systematik, Phylogenie und vergleichenden Morphologie der Juliden und uber einige andere Diplopoden. Archiv fur Naturgeschichte, 65 (1 / 3), 183 - 221." type="journal article" year="1899">Verhoeff, 1899</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36E7FFE1E765FF7FF97DFA2E3AEF" blockId="10.[151,1436,1590,1937]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF97DFA2E3AEF" pageId="10" pageNumber="53">
- Anterior and posterior processes of opisthomere of approximately the same length and development (
<figureCitation id="137E2A62FFE1E765FB51F97DFB463AD5" box="[1209,1268,1708,1729]" captionStart="FIGURES 1 7" captionStartId="4.[151,264,1545,1568]" captionTargetBox="[165,1436,269,1501]" captionTargetId="figure@4.[151,1436,267,1501]" captionTargetPageId="4" captionText="FIGURES 1 7. (1) In situ gonopods of a M. p. projectum male, from ANP, Hungary, caudal view. (2) Right promere of a M. p. projectum individual, from ANP, Hungary, mesal view. (3) Right opisthomere of the same individual, lateral view (scale bars: 0.5 mm). (4) Same individual, apical end of left opisthomere, mesal view (scale bar: 0.1 mm). Right opishomere of the same individual (5) caudo-lateral view; (6) lateral view; (7) antero-lateral view (scale bars: 0.2 mm)." httpUri="https://zenodo.org/record/207901/files/figure.png" pageId="10" pageNumber="53">Fig. 3</figureCitation>
). Promere with a serrated apical lobe (by
<taxonomicName id="4C454D64FFE1E765FE3AF918FDF13ACA" ID-CoL="8KR5F" box="[466,579,1737,1758]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="kochi">
<emphasis id="B931EAF5FFE1E765FE3AF918FDF13ACA" box="[466,579,1737,1758]" italics="true" pageId="10" pageNumber="53">M. p. kochi</emphasis>
</taxonomicName>
this serrated lobe sometimes not visible under a standard light microscope, just under stereo microscope, at high magnification).................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E42FFE1E765FF7FF8D5FA2E3B22" pageId="10" pageNumber="53">
<paragraph id="8BFA36E7FFE1E765FF7FF8D5FA2E3B0D" blockId="10.[151,1436,1590,1937]" box="[151,1436,1796,1817]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF8D5FA2E3B0D" box="[151,1436,1796,1817]" pageId="10" pageNumber="53">
3 Promere tapering, the whole gonopod
<emphasis id="B931EAF5FFE1E765FDB8F8D5FD3E3B0D" box="[592,652,1796,1817]" italics="true" pageId="10" pageNumber="53">in situ</emphasis>
appears to be thin (
<figureCitation id="137E2A62FFE1E765FCA0F8D5FC093B0D" box="[840,955,1796,1817]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1201,1224]" captionTargetBox="[155,1411,254,1157]" captionTargetId="figure@8.[151,1436,254,1158]" captionTargetPageId="8" captionText="FIGURES 14 16. (14) In situ gonopods of a M. p. kochi male, from Oława, Poland, caudal view. (15) Right promere of a M. p. kochi male from Potsdam, Germany: caudo-mesal, mesal and antero-mesal views. (16) Right promere of a M. p. projectum male from ANP, Aggtelek, Patkós Valley, Hungary: caudo-mesal, mesal and antero-mesal views. (scale bars 0.2 mm)" httpUri="https://zenodo.org/record/207904/files/figure.png" pageId="10" pageNumber="53">Figs. 1415</figureCitation>
).........
<taxonomicName id="4C454D64FFE1E765FBD4F8D5FA2E3B0D" ID-CoL="8KR5F" authority="Verhoeff, 1907" authorityName="Verhoeff" authorityYear="1907" box="[1084,1436,1796,1817]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="kochi">
<emphasis id="B931EAF5FFE1E765FBD4F8D5FB453B0D" box="[1084,1271,1796,1817]" italics="true" pageId="10" pageNumber="53">M. projectum kochi</emphasis>
(
<bibRefCitation id="EFD44B16FFE1E765FAEBF8D5FA273B0D" author="Verhoeff" box="[1283,1429,1796,1817]" pageId="10" pageNumber="53" refString="Verhoeff, C. (1907) Uber Diplopoden 6. (26.) Aufsatz: Tausendfuβler aus Brandenburg und andere Formen aus Ostdeutschland und Osterreich-Ungarn. Mitteilungen aus dem Zoologischen Museum in Berlin, 3, 291 - 337." type="journal article" year="1907">Verhoeff, 1907</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36E7FFE1E765FF7FF8F0FA2E3B22" blockId="10.[151,1436,1590,1937]" box="[151,1436,1825,1846]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF8F0FA2E3B22" box="[151,1436,1825,1846]" pageId="10" pageNumber="53">
- Promere not tapering, the whole gonopod
<emphasis id="B931EAF5FFE1E765FD9DF8F0FD023B22" box="[629,688,1825,1846]" italics="true" pageId="10" pageNumber="53">in situ</emphasis>
appears to be stout (
<figureCitation id="137E2A62FFE1E765FC9EF8F0FC0B3B22" box="[886,953,1825,1846]" captionStart="FIGURES 1 7" captionStartId="4.[151,264,1545,1568]" captionTargetBox="[165,1436,269,1501]" captionTargetId="figure@4.[151,1436,267,1501]" captionTargetPageId="4" captionText="FIGURES 1 7. (1) In situ gonopods of a M. p. projectum male, from ANP, Hungary, caudal view. (2) Right promere of a M. p. projectum individual, from ANP, Hungary, mesal view. (3) Right opisthomere of the same individual, lateral view (scale bars: 0.5 mm). (4) Same individual, apical end of left opisthomere, mesal view (scale bar: 0.1 mm). Right opishomere of the same individual (5) caudo-lateral view; (6) lateral view; (7) antero-lateral view (scale bars: 0.2 mm)." httpUri="https://zenodo.org/record/207901/files/figure.png" pageId="10" pageNumber="53">Figs. 1</figureCitation>
,
<figureCitation id="137E2A62FFE1E765FC2DF8F0FC6C3B22" box="[965,990,1825,1846]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1201,1224]" captionTargetBox="[155,1411,254,1157]" captionTargetId="figure@8.[151,1436,254,1158]" captionTargetPageId="8" captionText="FIGURES 14 16. (14) In situ gonopods of a M. p. kochi male, from Oława, Poland, caudal view. (15) Right promere of a M. p. kochi male from Potsdam, Germany: caudo-mesal, mesal and antero-mesal views. (16) Right promere of a M. p. projectum male from ANP, Aggtelek, Patkós Valley, Hungary: caudo-mesal, mesal and antero-mesal views. (scale bars 0.2 mm)" httpUri="https://zenodo.org/record/207904/files/figure.png" pageId="10" pageNumber="53">16</figureCitation>
)....
<taxonomicName id="4C454D64FFE1E765FBC8F8F0FA2E3B22" ID-CoL="8KR7P" authority="Verhoeff, 1894" authorityName="Verhoeff" authorityYear="1894" box="[1056,1436,1825,1846]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="projectum">
<emphasis id="B931EAF5FFE1E765FBC8F8F0FAB43B22" box="[1056,1286,1825,1846]" italics="true" pageId="10" pageNumber="53">M. projectum projectum</emphasis>
Verhoeff, 1894
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B12E42FFE1E765FF7FF8EFFB9A3B85" pageId="10" pageNumber="53">
<paragraph id="8BFA36E7FFE1E765FF7FF8EFFA2E3B65" blockId="10.[151,1436,1590,1937]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF8EFFA2E3B65" pageId="10" pageNumber="53">
4 Ampullas well-developed, dorsal ampulla wide, bulbous, ventral ampulla elongated (
<figureCitation id="137E2A62FFE1E765FBF8F8EFFBE33B47" box="[1040,1105,1854,1875]" captionStart="FIGURE 8. M. p" captionStartId="5.[151,250,1741,1764]" captionTargetBox="[172,1411,577,1684]" captionTargetId="figure@5.[160,1432,553,1704]" captionTargetPageId="5" captionText="FIGURE 8. M. p. projectum genitalia variation: (a) (c) female vulvae, (d) (i) males, right and left opisthomeres from mesal view. (a) ANP, Szin, Szelcepuszta, left vulva caudal, right vulva mesal view; (b) ANP, Jósvafő, Tohonya Valley, left vulva caudal, right vulva mesal view; (c) Romania, Maramureşului Basin, Rona de Sus, left vulva caudal, right vulva anterior view. (d) (h) five males from the same vial, ANP, Aggtelek, Ménes valley. (i) ANP, Aggtelek, Patkós Valley (scale bars: 0.2 mm)." httpUri="https://zenodo.org/record/207902/files/figure.png" pageId="10" pageNumber="53">Figs. 8</figureCitation>
ac)............................................................................................
<taxonomicName id="4C454D64FFE1E765FBC8F88DFA2E3B65" ID-CoL="8KR7P" authority="Verhoeff, 1894" authorityName="Verhoeff" authorityYear="1894" box="[1056,1436,1884,1905]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="projectum">
<emphasis id="B931EAF5FFE1E765FBC8F88DFAB43B65" box="[1056,1286,1884,1905]" italics="true" pageId="10" pageNumber="53">M. projectum projectum</emphasis>
Verhoeff, 1894
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFA36E7FFE1E765FF7FF8ADFB9A3B85" blockId="10.[151,1436,1590,1937]" box="[151,1064,1916,1937]" pageId="10" pageNumber="53">
<keyLead id="30B495D2FFE1E765FF7FF8ADFB9A3B85" box="[151,1064,1916,1937]" pageId="10" pageNumber="53">
- Ampullas thin, elongated, dorsal ampulla not bulbous
<taxonomicName id="4C454D64FFE1E765FD21F8ADFB9A3B85" ID-CoL="8KR5F" authority="Verhoeff, 1907" authorityName="Verhoeff" authorityYear="1907" box="[713,1064,1916,1937]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="10" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="projectum" subSpecies="kochi">
<emphasis id="B931EAF5FFE1E765FD21F8ADFC313B85" box="[713,899,1916,1937]" italics="true" pageId="10" pageNumber="53">M. projectum kochi</emphasis>
(
<bibRefCitation id="EFD44B16FFE1E765FC67F8ADFB933B85" author="Verhoeff" box="[911,1057,1916,1937]" pageId="10" pageNumber="53" refString="Verhoeff, C. (1907) Uber Diplopoden 6. (26.) Aufsatz: Tausendfuβler aus Brandenburg und andere Formen aus Ostdeutschland und Osterreich-Ungarn. Mitteilungen aus dem Zoologischen Museum in Berlin, 3, 291 - 337." type="journal article" year="1907">Verhoeff, 1907</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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