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<document id="E26632F85302636C1AB52682230E727E" ID-CLB-Dataset="26453" ID-DOI="10.11646/zootaxa.4619.2.10" ID-GBIF-Dataset="39031e6f-c6e2-4bdc-9a21-1000bdc1d443" ID-ISSN="1175-5326" ID-Zenodo-Dep="3249539" ID-ZooBank="F461B05C-CDD2-466B-92F7-10629300F507" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1560916724472" checkinUser="plazi" docAuthor="Bravo, Freddy &amp; Araújo, Maíra Xavier" docDate="2019" docId="03CD39647D42FFFFBDFCF96EFA92FC03" docLanguage="en" docName="zootaxa.4619.2.10.pdf" docOrigin="Zootaxa 4619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bryopharsos paulistensis Bravo &amp; Araújo 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="368" masterDocId="FFF4411C7D43FFFBBD6BFFA7FF8DFFC8" masterDocTitle="Three new species of Bryopharsos Quate, 1996 (Diptera, Psychodidae, Psychodinae) from the Neotropical Region" masterLastPageNumber="370" masterPageNumber="364" pageNumber="365" updateTime="1698736224280" updateUser="ExternalLinkService">
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<mods:title id="489BA67C468DC4D5AAFB6FBADB84E46F">Three new species of Bryopharsos Quate, 1996 (Diptera, Psychodidae, Psychodinae) from the Neotropical Region</mods:title>
</mods:titleInfo>
<mods:name id="C7D9B2AF64C0C69570D5785F9708063F" type="personal">
<mods:role id="E601D52FF001CF49A11CA0DABD5FDA2E">
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<mods:namePart id="73436BA2254671D2D7AFFD9062984A80">Bravo, Freddy</mods:namePart>
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<mods:namePart id="C75D01CA193FF7E932D04C56545048F1">Araújo, Maíra Xavier</mods:namePart>
</mods:name>
<mods:typeOfResource id="71830F555E790FDD70F5830B2345BACD">text</mods:typeOfResource>
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<mods:title id="6461D87B734AEDF7E1E99FE21EC2C45D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2D579B8B947BE13B289EDEC7FCF39686">
<mods:date id="5D5FED84EE6BBB635FC4B36061D18A1E">2019</mods:date>
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<mods:number id="C9BADDAACA317434DBD59383AE88C00F">2019-06-19</mods:number>
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<mods:number id="FEA15C2ADD25F276B9CB568D04066045">4619</mods:number>
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<mods:identifier id="2B2F61B1225AC9B67BE8B7383BA349BF" type="DOI">10.11646/zootaxa.4619.2.10</mods:identifier>
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<treatment id="03CD39647D42FFFFBDFCF96EFA92FC03" ID-GBIF-Taxon="158297419" LSID="urn:lsid:plazi:treatment:03CD39647D42FFFFBDFCF96EFA92FC03" httpUri="http://treatment.plazi.org/id/03CD39647D42FFFFBDFCF96EFA92FC03" lastPageId="4" lastPageNumber="368" pageId="1" pageNumber="365">
<subSubSection id="C37EDBF97D42FFFABDFCF96EFDA7F92B" box="[151,554,1737,1764]" pageId="1" pageNumber="365" type="nomenclature">
<paragraph id="8BDB88727D42FFFABDFCF96EFDA7F92B" blockId="1.[151,554,1737,1799]" box="[151,554,1737,1764]" pageId="1" pageNumber="365">
<heading id="D0933F1E7D42FFFABDFCF96EFDA7F92B" bold="true" box="[151,554,1737,1764]" fontSize="11" level="1" pageId="1" pageNumber="365" reason="1">
<emphasis id="B91054607D42FFFABDFCF96EFDA7F92B" bold="true" box="[151,554,1737,1764]" pageId="1" pageNumber="365">
<taxonomicName id="4C64F3F17D42FFFABDFCF96EFE4BF92C" authority="Bravo &amp; Araújo, 2019" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[151,454,1737,1764]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="paulistensis" status="sp. nov.">
<emphasis id="B91054607D42FFFABDFCF96EFE4BF92C" bold="true" box="[151,454,1737,1764]" italics="true" pageId="1" pageNumber="365">Bryopharsos paulistensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223E91B7D42FFFABCA6F96EFDA7F92B" box="[461,554,1737,1763]" pageId="1" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37EDBF97D42FFFABDFCF94AFEA3F8CF" box="[151,302,1773,1799]" pageId="1" pageNumber="365" type="description">
<paragraph id="8BDB88727D42FFFABDFCF94AFEA3F8CF" blockId="1.[151,554,1737,1799]" box="[151,302,1773,1799]" pageId="1" pageNumber="365">
(
<figureCitation id="135F94F77D42FFFABDF4F94AFEABF8CF" box="[159,294,1773,1799]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="1" pageNumber="365">Figs. 1324</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37EDBF97D42FFFABDFCF892FBCFF85F" pageId="1" pageNumber="365" type="materials_examined">
<paragraph id="8BDB88727D42FFFABDFCF892FBCFF85F" blockId="1.[151,1437,1844,2015]" pageId="1" pageNumber="365">
<emphasis id="B91054607D42FFFABDFCF892FEC5F887" bold="true" box="[151,328,1844,1871]" pageId="1" pageNumber="365">Type material.</emphasis>
<materialsCitation id="3B0C822F7D42FFFABC39F892FBCFF85F" ID-GBIF-Occurrence="2268317509" collectingDate="2004-07-25" collectingDateMax="2005-01-08" collectingDateMin="2004-07-25" collectionCode="MZFS" country="Brazil" location="Sete Barras" pageId="1" pageNumber="365" specimenCount="12" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="54DF36D07D42FFFABC39F892FE30F887" box="[338,445,1845,1871]" pageId="1" pageNumber="365" type="holotype">Holotype</typeStatus>
, male:
<collectingCountry id="F373C8E27D42FFFABF7FF892FDF4F887" box="[532,633,1845,1871]" name="Brazil" pageId="1" pageNumber="365">BRAZIL</collectingCountry>
,
<collectingRegion id="49A046907D42FFFABFE0F892FC8FF887" box="[651,770,1845,1871]" country="Brazil" name="Sao Paulo" pageId="1" pageNumber="365">São Paulo</collectingRegion>
,
<location id="8EBBDEA97D42FFFABE7BF892FC1AF887" LSID="urn:lsid:plazi:treatment:03CD39647D42FFFFBDFCF96EFA92FC03:8EBBDEA97D42FFFABE7BF892FC1AF887" box="[784,919,1845,1871]" country="Brazil" name="Sete Barras" pageId="1" pageNumber="365" stateProvince="Sao Paulo">Sete Barras</location>
,
<collectingDate id="EF9E575A7D42FFFABECEF892FBDFF887" box="[933,1106,1845,1871]" pageId="1" pageNumber="365" value="2004-11-13" valueMax="2004-11-27" valueMin="2004-11-13">
<date id="FFDAAEB27D42FFFABECEF892FBDFF887" box="[933,1106,1845,1871]" pageId="1" pageNumber="365" value="2004-11-13" valueMax="2004-11-27" valueMin="2004-11-13">1327.XI.2004</date>
</collectingDate>
,
<collectingMethod id="5225F0657D42FFFAB90BF892FB7CF887" box="[1120,1265,1845,1871]" pageId="1" pageNumber="365">Malaise trap</collectingMethod>
, without name of collector (
<collectionCode id="ED7510B77D42FFFABC40F8FEFEF7F8BB" box="[299,378,1881,1907]" pageId="1" pageNumber="365">MZFS</collectionCode>
).
<specimenCount id="9D6243FB7D42FFFABCE5F8FEFD9DF8BB" box="[398,528,1881,1907]" pageId="1" pageNumber="365" type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
male with same data of
<typeStatus id="54DF36D07D42FFFABE45F8FEFC1DF8BB" box="[814,912,1881,1907]" pageId="1" pageNumber="365" type="holotype">holotype</typeStatus>
except by date of collect:
<specimenCount id="9D6243FB7D42FFFAB9AFF8FEFAB6F8BB" box="[1220,1339,1881,1907]" pageId="1" pageNumber="365" type="generic" typeStatus="paratype">1 paratype</specimenCount>
<collectingDate id="EF9E575A7D42FFFAB82FF8FFFE81F85F" pageId="1" pageNumber="365" value="2004-07-25" valueMax="2004-11-07" valueMin="2004-07-25">
<date id="FFDAAEB27D42FFFAB82FF8FFFE81F85F" pageId="1" pageNumber="365" value="2004-07-25" valueMax="2004-11-07" valueMin="2004-07-25">25.VII 7.XI.2004</date>
</collectingDate>
;
<specimenCount id="9D6243FB7D42FFFABC73F8DAFE15F85F" box="[280,408,1917,1943]" pageId="1" pageNumber="365" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
<collectingDate id="EF9E575A7D42FFFABCF4F8DAFDCEF85F" box="[415,579,1917,1943]" pageId="1" pageNumber="365" value="2004-10-16" valueMax="2004-10-30" valueMin="2004-10-16">
<date id="FFDAAEB27D42FFFABCF4F8DAFDCEF85F" box="[415,579,1917,1943]" pageId="1" pageNumber="365" value="2004-10-16" valueMax="2004-10-30" valueMin="2004-10-16">1630.X.2004</date>
</collectingDate>
;
<specimenCount id="9D6243FB7D42FFFABF25F8DAFD43F85F" box="[590,718,1917,1943]" pageId="1" pageNumber="365" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
<collectingDate id="EF9E575A7D42FFFABFBEF8DBFCD2F85F" box="[725,863,1916,1943]" pageId="1" pageNumber="365" value="2004-12-24">
<date id="FFDAAEB27D42FFFABFBEF8DBFCD2F85F" box="[725,863,1916,1943]" pageId="1" pageNumber="365" value="2004-12-24">24.XII.2004</date>
</collectingDate>
<collectingDate id="EF9E575A7D42FFFABE04F8DBFC5AF85F" box="[879,983,1916,1943]" pageId="1" pageNumber="365" value="2005-01-08">
<date id="FFDAAEB27D42FFFABE04F8DBFC5AF85F" box="[879,983,1916,1943]" pageId="1" pageNumber="365" value="2005-01-08">08.I.2005</date>
</collectingDate>
(
<collectionCode id="ED7510B77D42FFFABE8DF8DAFBB8F85F" box="[998,1077,1917,1943]" pageId="1" pageNumber="365">MZFS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37EDBF97D42FFFABDACF806FE51F817" pageId="1" pageNumber="365" type="etymology">
<paragraph id="8BDB88727D42FFFABDACF806FE51F817" blockId="1.[151,1437,1844,2015]" pageId="1" pageNumber="365">
<emphasis id="B91054607D42FFFABDACF806FEC3F873" bold="true" box="[199,334,1953,1979]" pageId="1" pageNumber="365">Etymology.</emphasis>
The specific name
<taxonomicName id="4C64F3F17D42FFFABF45F806FD3FF873" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[558,690,1953,1979]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="paulistensis">
<emphasis id="B91054607D42FFFABF45F806FD3FF873" box="[558,690,1953,1979]" italics="true" pageId="1" pageNumber="365">paulistensis</emphasis>
</taxonomicName>
is based on the State where the specimen of the new species was collected, namely
<collectingRegion id="49A046907D42FFFABC0FF862FE55F817" box="[356,472,1989,2015]" country="Brazil" name="Sao Paulo" pageId="1" pageNumber="365">São Paulo</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="DF1BD8FA7D41FFF9BDFCF8A8FA19F813" httpUri="https://zenodo.org/record/3249541/files/figure.png" pageId="2" pageNumber="366" startId="2.[151,264,1807,1832]" targetBox="[151,1436,181,1780]" targetPageId="2">
<paragraph id="8BDB88727D41FFF9BDFCF8A8FA19F813" blockId="2.[151,1437,1807,2012]" pageId="2" pageNumber="366">
<emphasis id="B91054607D41FFF9BDFCF8A8FEC4F8E0" bold="true" box="[151,329,1807,1832]" pageId="2" pageNumber="366">FIGURES 112.</emphasis>
<taxonomicName id="4C64F3F17D41FFF9BC39F8B7FDCAF8EF" authority="Bravo &amp; Araújo, 2019" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[338,583,1808,1832]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="uncinatum" status="sp. nov.">
<emphasis id="B91054607D41FFF9BC39F8B7FDCAF8EF" box="[338,583,1808,1832]" italics="true" pageId="2" pageNumber="366">Bryopharsos uncinatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223E91B7D41FFF9BF24F8B7FD2FF8E0" box="[591,674,1808,1832]" pageId="2" pageNumber="366" rank="species">
<emphasis id="B91054607D41FFF9BF24F8B7FD2FF8E0" bold="true" box="[591,674,1808,1832]" pageId="2" pageNumber="366">sp. nov.</emphasis>
</taxonomicNameLabel>
(1) Head, anterior; (2) head, posterior; (3) labellum; (4) cardo-stipes; (5) scape, pedicel and four basal flagellomeres; (6) apical flagellomeres and apiculus; (7) flagellomere and leaf-shape ascoid; (8) wing; (9) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (10) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); (11) epandrium, cerci and hypoproct (dorsal); (12) epandrium, cerci, hypoproct and epiproct (ventral). Abbreviations: ae = aedeagus; agx = anterior expansion of gonocoxal apodemes; cde = cardo-stipes; ep = epiproct; hyp = hypandrium; lbm = labellum; lc = lacinia; pgx = posterior expansion of gonocoxal apodemes; pm = paramere.
</paragraph>
</caption>
<caption id="DF1BD8FA7D40FFF8BDFCF95AFE06F825" httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="3" pageNumber="367" startId="3.[151,264,1789,1814]" targetBox="[151,1436,181,1769]" targetPageId="3">
<paragraph id="8BDB88727D40FFF8BDFCF95AFE06F825" blockId="3.[151,1437,1789,2030]" pageId="3" pageNumber="367">
<emphasis id="B91054607D40FFF8BDFCF95AFED5F8DE" bold="true" box="[151,344,1789,1814]" pageId="3" pageNumber="367">FIGURES 1324.</emphasis>
<taxonomicName id="4C64F3F17D40FFF8BC0AF95AFDE8F8DD" authority="Bravo &amp; Araújo, 2019" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[353,613,1789,1813]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="367" phylum="Arthropoda" rank="species" species="paulistensis" status="sp. nov.">
<emphasis id="B91054607D40FFF8BC0AF95AFDE8F8DD" box="[353,613,1789,1813]" italics="true" pageId="3" pageNumber="367">Bryopharsos paulistensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223E91B7D40FFF8BF05F95AFD4FF8DD" box="[622,706,1789,1813]" pageId="3" pageNumber="367" rank="species">
<emphasis id="B91054607D40FFF8BF05F95AFD4FF8DD" bold="true" box="[622,706,1789,1813]" pageId="3" pageNumber="367">sp. nov.</emphasis>
</taxonomicNameLabel>
(13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1.
</paragraph>
</caption>
<subSubSection id="C37EDBF97D47FFFFBDACFF3FFC21FF33" pageId="4" pageNumber="368" type="diagnosis">
<paragraph id="8BDB88727D47FFFFBDACFF3FFC21FF33" blockId="4.[151,1437,152,971]" pageId="4" pageNumber="368">
<emphasis id="B91054607D47FFFFBDACFF3FFECFFF7B" bold="true" box="[199,322,152,179]" pageId="4" pageNumber="368">Diagnosis.</emphasis>
Male. Eye bridge with five facet rows; wing twice as long than as wide; ejaculatory apodeme subrectangular in dorsal view; cercus narrower in the one-half apical portion, with one tenaculum; aedeagus long, expanded, hemispheric basally and narrow apically with pointed apex.
</paragraph>
</subSubSection>
<subSubSection id="C37EDBF97D47FFFFBDACFEA3FE2FFCDF" pageId="4" pageNumber="368" type="description">
<paragraph id="8BDB88727D47FFFFBDACFEA3FBA7FD3B" blockId="4.[151,1437,152,971]" pageId="4" pageNumber="368">
<emphasis id="B91054607D47FFFFBDACFEA3FED4FED7" bold="true" box="[199,345,260,287]" pageId="4" pageNumber="368">Description.</emphasis>
Male. Head vertex length almost twice the width of eye bridge; eye bridge with five facet rows; pedicel shorter than scape (
<figureCitation id="135F94F77D47FFFFBCAFFE8EFDFFFE8C" box="[452,626,297,324]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Figs. 13 and 16</figureCitation>
). Occipital setae present and distributed into one basal block; post-ocular alveoli absent (
<figureCitation id="135F94F77D47FFFFBC29FEEAFE1BFEAF" box="[322,406,333,359]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Fig. 14</figureCitation>
). Labellum very short, ending before the apex of labrum; maxillary lacinia short, knife like well sclerotized (
<figureCitation id="135F94F77D47FFFFBC31FED6FE22FE43" box="[346,431,368,395]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Fig. 15</figureCitation>
). Fourteen flagellomeres, eccentric except first flagellomere that is spherical; internodes of flagellomeres 14 short; internodes of flagellomeres 413 0.7 times the length of nodes; flagellomere 14 without internode (
<figureCitation id="135F94F77D47FFFFBC7BFE1EFE4CFE1B" box="[272,449,441,467]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Figs. 16 and 17</figureCitation>
). One pair of ascoids per flagellomere, leaf shaped (
<figureCitation id="135F94F77D47FFFFB962FE1EFBD1FE1B" box="[1033,1116,441,467]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Fig. 18</figureCitation>
). Thorax without allurement organs. Wing twice as long as wide; seta alveoli distributed uniformly on wing membrane (
<figureCitation id="135F94F77D47FFFFB9CEFE7AFB74FE3F" box="[1189,1273,477,503]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Fig. 19</figureCitation>
). Hypandrium narrow; gonostylus with blunt apex; aedeagus expanded, hemispheric basally and narrow apically with pointed apex; ejaculatory apodeme rectangular in dorsal view; paramere with the same length of aedeagus and capitate apex (
<figureCitation id="135F94F77D47FFFFBDF4FDEEFEDEFDAB" box="[159,339,585,611]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Figs. 20 and 21</figureCitation>
). Gonocoxal apodeme anteriorly projected as two thin blades ending in an acute apex, not fused, and posteriorly projected as a pair of lateral, ovoid lobes with long bristles (
<figureCitation id="135F94F77D47FFFFBEB6FDCAFBBCFD4F" box="[989,1073,621,647]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Fig. 20</figureCitation>
). Epandrium with posterior margin slightly concave; cercus with apical one half narrower than base with one apical tenaculum (
<typeStatus id="54DF36D07D47FFFFB9AEFD36FAA4FD63" box="[1221,1321,657,683]" pageId="4" pageNumber="368" type="holotype">holotype</typeStatus>
with right cercus with two tenacula); subepandrial sclerite plate-like with distal arms sclerotized where cerci are articulated; epiproct smaller than hypoproct, both with apical micropilosity (
<figureCitation id="135F94F77D47FFFFBE01FD7EFB90FD3B" box="[874,1053,729,755]" captionStart="FIGURES 1324" captionStartId="3.[151,264,1789,1814]" captionTargetBox="[151,1436,181,1769]" captionTargetId="figure@3.[151,1436,181,1769]" captionTargetPageId="3" captionText="FIGURES 1324. Bryopharsos paulistensis sp. nov. (13) Head, anterior; (14) head, posterior; (15) cardo-stipes, labellum and first segment of palpus; (16) scape, pedicel and three basal flagellomeres; (17) apical flagellomeres and apiculus; (18) flagellomere and leaf-shape ascoid; (19) wing; (20) gonocoxites, gonostyli, hypandrium and ejaculatory apodeme (dorsal); (21) gonocoxites, gonostyli, gonocoxal apodemes and ejaculatory apodeme (ventral); posterior projection of gonocoxal apodeme not figured; (22) epandrium, hypoproct and epiproct (ventral); (23) cercus with one tenaculum; (24) right cercus of holotype with two tenacula.Abbreviations: ae = aedeagus; lbm = labellum; pgx = posterior expansion of gonocoxal apodemes; pm = paramere; ps1 = palpus segment 1." httpUri="https://zenodo.org/record/3249543/files/figure.png" pageId="4" pageNumber="368">Figs. 22, 23, 24</figureCitation>
).
</paragraph>
<paragraph id="8BDB88727D47FFFFBDACFD5AFE2FFCDF" blockId="4.[151,1437,152,971]" box="[199,418,765,791]" pageId="4" pageNumber="368">
<emphasis id="B91054607D47FFFFBDACFD5AFEA8FCDF" bold="true" box="[199,293,765,791]" pageId="4" pageNumber="368">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37EDBF97D47FFFFBDACFC86FA92FC03" pageId="4" pageNumber="368" type="discussion">
<paragraph id="8BDB88727D47FFFFBDACFC86FA92FC03" blockId="4.[151,1437,152,971]" pageId="4" pageNumber="368">
<emphasis id="B91054607D47FFFFBDACFC86FEB6FCF3" bold="true" box="[199,315,801,827]" pageId="4" pageNumber="368">Remarks.</emphasis>
The new species
<taxonomicName id="4C64F3F17D47FFFFBF6DFC86FC99FCF3" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[518,788,801,827]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="uncinatum">
<emphasis id="B91054607D47FFFFBF6DFC86FC99FCF3" box="[518,788,801,827]" italics="true" pageId="4" pageNumber="368">Bryopharsos uncinatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C64F3F17D47FFFFBE27FC85FC7CFCF3" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[844,1009,801,827]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="paulistensis">
<emphasis id="B91054607D47FFFFBE27FC85FC7CFCF3" box="[844,1009,801,827]" italics="true" pageId="4" pageNumber="368">B. paulistensis</emphasis>
</taxonomicName>
have only one tenaculum in the cerci (except for the
<typeStatus id="54DF36D07D47FFFFBC2DFCE2FE26FC97" box="[326,427,837,863]" pageId="4" pageNumber="368" type="holotype">holotype</typeStatus>
, which has two tenacula in one cercus). The presence of one tenaculum is unique, as the other species of
<taxonomicName id="4C64F3F17D47FFFFBC3EFCCEFE69FC4B" authorityName=": Quate" authorityYear="1999" box="[341,484,873,899]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B91054607D47FFFFBC3EFCCEFE69FC4B" box="[341,484,873,899]" italics="true" pageId="4" pageNumber="368">Bryopharsos</emphasis>
</taxonomicName>
have three to five tenacula.
<taxonomicName id="4C64F3F17D47FFFFBE44FCCEFBB3FC4B" authorityName="Bravo &amp; Araújo" authorityYear="2019" box="[815,1086,873,899]" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="uncinatum">
<emphasis id="B91054607D47FFFFBE44FCCEFBB3FC4B" box="[815,1086,873,899]" italics="true" pageId="4" pageNumber="368">Bryopharsos uncinatum</emphasis>
</taxonomicName>
has five facet rows in the eye bridge, and the specimens does not have lobes in the anterior projection of the gonocoxal apodemes, while
<taxonomicName id="4C64F3F17D47FFFFB821FC29FF7CFC03" authorityName="Bravo &amp; Araújo" authorityYear="2019" class="Insecta" family="Psychodidae" genus="Bryopharsos" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="368" phylum="Arthropoda" rank="species" species="paulistensis">
<emphasis id="B91054607D47FFFFB821FC29FF7CFC03" italics="true" pageId="4" pageNumber="368">B. paulistensis</emphasis>
</taxonomicName>
has four facet rows and lobes are present in the anterior projection of the gonocoxal apodemes.
</paragraph>
</subSubSection>
</treatment>
</document>