345 lines
38 KiB
XML
345 lines
38 KiB
XML
<document id="F19461609513974E48250D3D10347D55" ID-DOI="10.11646/zootaxa.5507.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13757579" ID-ZooBank="C5A55C22-3D3B-4B49-9591-148B0FB7E8C3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1726221267114" checkinUser="plazi" docAuthor="Telnov, Dmitry" docDate="2024" docId="03C351407B27FFE9FF1C881285A5FBC4" docLanguage="en" docName="zootaxa.5507.2.4.pdf" docOrigin="Zootaxa 5507 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5507.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nitorus Telnov 2007" docType="key" docVersion="1" lastPageNumber="311" masterDocId="FFFA29387B3EFFF3FF8B8A788038FFB1" masterDocTitle="New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum" masterLastPageNumber="312" masterPageNumber="285" pageNumber="310" updateTime="1726661670562" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="30CCF4EBCF3EDB060A5644BFEB673AF8">New species and records of Anthicidae Latreille, 1819 (Coleoptera) from Taiwan with a key to Asian Nitorus Telnov, 2007 species with gibbose pronotum</mods:title>
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<mods:name id="1E2CDA3C07E830935DEF857F458D8B3B" type="personal">
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<mods:namePart id="16E56F4A157CE8AA1F326C7EB9D1F0C4">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="E0F6E0B70AC1AE6379A12455C07CB3B7">Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD London, United Kingdom & Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401 Daugavpils, Latvia & Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004 Rīga, Latvia</mods:affiliation>
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<mods:title id="D96349FC918AA1E779A4A36EBEBA06F0">Zootaxa</mods:title>
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<mods:date id="9589CBA40F33AE6566461D042796CB7E">2024</mods:date>
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<mods:number id="146489D79E6D68F9B80AAF1921084EE5">2024-09-12</mods:number>
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<mods:number id="31CC65F74F6BA2614A82CDB74C991F35">5507</mods:number>
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<mods:url id="B5211F535E71C344D3554D03719901B0">http://dx.doi.org/10.11646/zootaxa.5507.2.4</mods:url>
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<subSubSection id="C370B3DD7B27FFEAFF1C88128458FD34" box="[151,1120,618,645]" pageId="25" pageNumber="310" type="nomenclature">
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<paragraph id="8BD5E0567B27FFEAFF1C88128458FD34" blockId="25.[151,1120,618,645]" box="[151,1120,618,645]" pageId="25" pageNumber="310">
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<heading id="D09D573A7B27FFEAFF1C88128458FD34" bold="true" box="[151,1120,618,645]" fontSize="11" level="1" pageId="25" pageNumber="310" reason="1">
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<emphasis id="B91E3C447B27FFEAFF1C88128458FD34" bold="true" box="[151,1120,618,645]" pageId="25" pageNumber="310">
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Addition. Key to Asian species of
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<taxonomicName id="4C6A9BD57B27FFEAFDB288138300FD35" authority="Telnov, 2007" authorityName="Telnov" authorityYear="2007" box="[569,824,618,645]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="genus">
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<emphasis id="B91E3C447B27FFEAFDB2881382ACFD34" bold="true" box="[569,660,619,645]" italics="true" pageId="25" pageNumber="310">Nitorus</emphasis>
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Telnov, 2007
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</taxonomicName>
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with gibbose pronotum
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C370B3DD7B27FFEAFF1C88CB835EFC5C" pageId="25" pageNumber="310" type="description">
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<paragraph id="8BD5E0567B27FFEAFF1C88CB8328FCA4" blockId="25.[151,1437,691,1005]" pageId="25" pageNumber="310">
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This is the first attempt to list and key the species of
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<taxonomicName id="4C6A9BD57B27FFEAFD7488CB836BFD7D" authorityName="Telnov" authorityYear="2007" box="[767,851,691,716]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="genus">
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<emphasis id="B91E3C447B27FFEAFD7488CB836BFD7D" box="[767,851,691,716]" italics="true" pageId="25" pageNumber="310">Nitorus</emphasis>
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</taxonomicName>
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from Asian mainland and Sundaland with dorsally strongly gibbose and, in some species, saddle-like depressed pronotum. Proper identification is possible for both sexes, but the present key is focused primarily on males.
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</paragraph>
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<paragraph id="8BD5E0567B27FFEAFF4C8967835EFC5C" blockId="25.[151,1437,691,1005]" pageId="25" pageNumber="310">
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Several Asian
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<taxonomicName id="4C6A9BD57B27FFEAFEE089678187FC89" authorityName="Telnov" authorityYear="2007" box="[363,447,799,824]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="genus">
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<emphasis id="B91E3C447B27FFEAFEE089678187FC89" box="[363,447,799,824]" italics="true" pageId="25" pageNumber="310">Nitorus</emphasis>
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</taxonomicName>
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(for instance,
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<taxonomicName id="4C6A9BD57B27FFEAFDEF89678392FC88" authority="(Pic, 1931)" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[612,938,799,825]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="philipinnensis">
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<emphasis id="B91E3C447B27FFEAFDEF8967831BFC88" box="[612,803,799,825]" italics="true" pageId="25" pageNumber="310">N. philipinnensis</emphasis>
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(Pic, 1931)
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</taxonomicName>
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,
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<taxonomicName id="4C6A9BD57B27FFEAFC3C896784FFFC88" authority="(Pic, 1894)" baseAuthorityName="Pic" baseAuthorityYear="1894" box="[951,1223,799,825]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="sumatrae">
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<emphasis id="B91E3C447B27FFEAFC3C89678479FC89" box="[951,1089,799,824]" italics="true" pageId="25" pageNumber="310">N. sumatrae</emphasis>
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(Pic, 1894)
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</taxonomicName>
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etc.) are known to have slightly bigibbose pronotum (gibbosity is not prominent in lateral view). These species are not considered in the present key.
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<taxonomicName id="4C6A9BD57B27FFEAFEE4891F82DDFC30" authority="(Pic, 1923)" baseAuthorityName="Pic" baseAuthorityYear="1923" box="[367,741,871,897]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="testaceitarsis">
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<emphasis id="B91E3C447B27FFEAFEE4891F8267FC31" box="[367,607,871,896]" italics="true" pageId="25" pageNumber="310">Nitorus testaceitarsis</emphasis>
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(Pic, 1923)
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</taxonomicName>
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from Greater Sunda Islands is unknown to me. According to the original description (Pic 1923), base of pronotum is bituberculate in this species. However, it is unclear if the anterior pronotal lobe is gibbose or not and does the lateral pronotal constriction continues onto pronotal disc or not. I therefore decided not to include this species to the present key.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C370B3DD7B27FFE9FF1C8E6085A5FBC4" lastPageId="26" lastPageNumber="311" pageId="25" pageNumber="310" type="key">
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<key id="FFFBB7677B27FFE9FF1C8E6085A5FBC4" lastPageId="26" lastPageNumber="311" pageId="25" pageNumber="310">
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<keyStep id="309EF8F37B27FFEAFF1C8E6085A4FBFB" pageId="25" pageNumber="310">
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<paragraph id="8BD5E0567B27FFEAFF1C8E6085A4FB9C" blockId="25.[151,1437,1048,2038]" box="[151,1436,1048,1070]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8E6085A4FB9C" box="[151,1436,1048,1070]" pageId="25" pageNumber="310">1 Pronotum in lateral view with distinct gibbosity on anterior, with or without another gibbosity on posterior lobe.......... 3</keyLead>
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</paragraph>
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<paragraph id="8BD5E0567B27FFEAFF1C8E4D85A4FBFB" blockId="25.[151,1437,1048,2038]" box="[151,1436,1077,1099]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8E4D85A4FBFB" box="[151,1436,1077,1099]" pageId="25" pageNumber="310">- Pronotum in lateral view with distinct gibbosity on posterior lobe only, anterior lobe slightly convex to flattened.......... 2</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309EF8F37B27FFEAFF1C8E2A85A4FB4A" pageId="25" pageNumber="310">
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||
<paragraph id="8BD5E0567B27FFEAFF1C8E2A85A4FB13" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8E2A85A4FB13" pageId="25" pageNumber="310">
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2 Lateral constriction continues onto pronotal dorsum forming moderately deep impression; head strongly constricted posterior to compound eyes, posterior head nearly tapered; dorsum uniformly brown with paler legs and antennae.................................................................................................
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<taxonomicName id="4C6A9BD57B27FFEAFB3A8EF685A4FB13" authority="(Pic, 1913)" baseAuthorityName="Pic" baseAuthorityYear="1913" box="[1201,1436,1165,1187]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="peguensis">
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<emphasis id="B91E3C447B27FFEAFB3A8EF68514FB12" box="[1201,1324,1166,1187]" italics="true" pageId="25" pageNumber="310">N. peguensis</emphasis>
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(
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<bibRefCitation id="EFFB9DA77B27FFEAFAB28EF585ADFB13" author="Pic, M." box="[1337,1429,1165,1187]" pageId="25" pageNumber="310" pagination="129 - 135" refId="ref13523" refString="Pic, M. (1913) H. Sauter's Formosa-Ausbeute. Anthicidae, Pedilidae und Hylophilidae. Archiv fur Naturgeschichte, Series A, 79, 129 - 135." type="journal article" year="1913">Pic, 1913</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BD5E0567B27FFEAFF1C8ED385A4FB4A" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8ED385A4FB4A" pageId="25" pageNumber="310">
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- Lateral constriction not or hardly continues onto pronotal dorsum; head posterior to compound eyes rounded in broad arc; forebody paler yellowish to rufous, each elytron with pale spot in postbasal impression area......................................................................... some Japanese specimens of
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<taxonomicName id="4C6A9BD57B27FFEAFBA78E9E8506FB4A" authority="(Marseul, 1876)" baseAuthorityName="Marseul" baseAuthorityYear="1876" box="[1068,1342,1253,1275]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="trigibber">
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<emphasis id="B91E3C447B27FFEAFBA78E9E84A5FB4B" box="[1068,1181,1253,1275]" italics="true" pageId="25" pageNumber="310">N. trigibber</emphasis>
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(Marseul, 1876)
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</taxonomicName>
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(
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<figureCitation id="1351FCD37B27FFEAFAC08E9D85ADFB4A" box="[1355,1429,1253,1275]" captionStart="FIGURE 6" captionStartId="9.[151,250,1407,1431]" captionTargetBox="[271,1317,192,1369]" captionTargetId="figure-18@9.[258,1330,181,1382]" captionTargetPageId="9" captionText="FIGURE 6. Nitorus trigibber (Marseul, 1876). A—♀ from ‘14km W of Chihshang’, habitus, dorsal view; B—ditto, lateral view; C—♀ from ‘Japan’ collected by Lewis, forebody, lateral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13757595" httpUri="https://zenodo.org/record/13757595/files/figure.png" pageId="25" pageNumber="310">Fig. 6C</figureCitation>
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)
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309EF8F37B27FFEAFF1C8F7A85A5FAC1" pageId="25" pageNumber="310">
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<paragraph id="8BD5E0567B27FFEAFF1C8F7A85A4FA84" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8F7A85A4FA84" pageId="25" pageNumber="310">3 Pronotum in lateral view with distinct gibbosity on anterior and more or less strong gibbosity also on posterior lobe, lateral constriction continues onto pronotal dorsum in a form of saddle-like impression................................... 4</keyLead>
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</paragraph>
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<paragraph id="8BD5E0567B27FFEAFF1C8F4585A5FAC1" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8F4585A5FAC1" pageId="25" pageNumber="310">
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- Pronotum in lateral view with gibbosity on anterior lobe only; posterior lobe not gibbose, lateral constriction does not continue onto pronotal dorsum..........................................................
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<taxonomicName id="4C6A9BD57B27FFEAFBFF8F2385A5FAC1" authority="(Telnov, 2005)" baseAuthorityName="Telnov" baseAuthorityYear="2005" box="[1140,1437,1370,1392]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="dromedarius">
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<emphasis id="B91E3C447B27FFEAFBFF8F238532FADE" box="[1140,1290,1370,1392]" italics="true" pageId="25" pageNumber="310">N. dromedarius</emphasis>
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(
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<bibRefCitation id="EFFB9DA77B27FFEAFA9C8F2285ADFAC1" author="Telnov, D." box="[1303,1429,1370,1392]" pageId="25" pageNumber="310" pagination="161 - 180" refId="ref13627" refString="Telnov, D. (2005) Anthicidae (Coleoptera) aus der Sammlung des Naturhistorischen Museums in Basel. Teil II: Bemerkenswerte und neue Notoxini, Endomiini, Anthicini (Anthicinae) aus der palaarktischen und orientalischen Region. Entomologica basiliensia et collectionis Frey, 27, 161 - 180." type="journal article" year="2005">Telnov, 2005</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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||
<keyStep id="309EF8F37B27FFEAFF1C8F0085A4FA54" pageId="25" pageNumber="310">
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<paragraph id="8BD5E0567B27FFEAFF1C8F0085A4FA1A" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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<keyLead id="309B43637B27FFEAFF1C8F0085A4FA1A" pageId="25" pageNumber="310">4 Pronotum in lateral view with anterior and posterior pronotal gibbosities nearly equally high, gibbosities apically tapered, not semicircular (broadly rounded).......................................................................... 5</keyLead>
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</paragraph>
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<paragraph id="8BD5E0567B27FFEAFF1C8FCB85A4FA54" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
|
||
<keyLead id="309B43637B27FFEAFF1C8FCB85A4FA54" pageId="25" pageNumber="310">- Pronotum in lateral view with anterior gibbosity significantly stronger than posterior or both gibbosities semicircular (not tapered apically)..................................................................................... 10</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309EF8F37B27FFEAFF1C8F9585A4F902" pageId="25" pageNumber="310">
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<paragraph id="8BD5E0567B27FFEAFF1C8F9585A4F98C" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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||
<keyLead id="309B43637B27FFEAFF1C8F9585A4F98C" pageId="25" pageNumber="310">5 Dorsal head minutely punctate; lateral margins of anterior pronotal lobe broadly rounded in dorsal view; width ratio of anterior to posterior pronotal lobe about 1.3 or less; elytra slightly convex or nearly flattened in dorsal aspect, postbasal transverse impression distinct; elytron with more or less wide pale transverse spot or fascia in postbasal impression area............ 7</keyLead>
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</paragraph>
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<paragraph id="8BD5E0567B27FFEAFF1C8C3D85A4F902" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
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||
<keyLead id="309B43637B27FFEAFF1C8C3D85A4F902" pageId="25" pageNumber="310">- Dorsal head with distinct punctures; lateral margins of anterior pronotal lobe expanded medially, subangulate in dorsal view; width ratio of anterior to posterior pronotal lobe about 1.4 or more; elytra distinctly convex in dorsal aspect, postbasal transverse impression not indicated; elytron without pale transverse spot or fascia in postbasal impression area, maximum dark reddish brown at base but not contrastingly paler than rest of elytra.................................................... 6</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309EF8F37B27FFEAFF1C8CC285A4F8D3" pageId="25" pageNumber="310">
|
||
<paragraph id="8BD5E0567B27FFEAFF1C8CC285A5F8BB" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
|
||
<keyLead id="309B43637B27FFEAFF1C8CC285A5F8BB" pageId="25" pageNumber="310">
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6 Dorsal head with large deep punctures; width ratio of anterior to posterior pronotal lobe about 1.5; lateral sides or pronotum in constriction area with distinct longitudinal subparallel wrinkles; aedeagus as in fig. 18, apex of median lobe narrowly pointed..............................................................................
|
||
<taxonomicName id="4C6A9BD57B27FFEAFB6B8C8E8574F8BB" authorityName="Telnov" authorityYear="2024" box="[1248,1356,1781,1803]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="sathulaz" status="sp. nov.">
|
||
<emphasis id="B91E3C447B27FFEAFB6B8C8E8574F8BB" box="[1248,1356,1781,1803]" italics="true" pageId="25" pageNumber="310">N. sathulaz</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D813F7B27FFEAFAD98C8D85A5F8BB" box="[1362,1437,1781,1802]" pageId="25" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B27FFEAFF1C8D6B85A4F8D3" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
|
||
<keyLead id="309B43637B27FFEAFF1C8D6B85A4F8D3" pageId="25" pageNumber="310">
|
||
- Dorsal head with distinct but rather moderate and shallow punctures; width ratio of anterior to posterior pronotal lobe about 1.4; lateral sides or pronotum in constriction area not or indistinctly longitudinally wrinkled; aedeagus not as in fig. 18, apex of median lobe less slender, rounded.................................................
|
||
<taxonomicName id="4C6A9BD57B27FFEAFBF58D3685A4F8D3" authority="(Pic, 1901)" baseAuthorityName="Pic" baseAuthorityYear="1901" box="[1150,1436,1869,1891]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="monstrosicollis">
|
||
<emphasis id="B91E3C447B27FFEAFBF58D368514F8D3" box="[1150,1324,1869,1891]" italics="true" pageId="25" pageNumber="310">N. monstrosicollis</emphasis>
|
||
(Pic, 1901)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B27FFEAFF1C8D1385A4F866" pageId="25" pageNumber="310">
|
||
<paragraph id="8BD5E0567B27FFEAFF1C8D1385A4F82C" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
|
||
<keyLead id="309B43637B27FFEAFF1C8D1385A4F82C" pageId="25" pageNumber="310">7 Pronotum in lateral view with anterior gibbosity somewhat stronger (stronger elevated or/and larger) than posterior or both gibbosities nearly equally strong; head somewhat constricted at base; aedeagus not as in figs 2F–I & 7................. 9</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B27FFEAFF1C8DDD85A4F866" blockId="25.[151,1437,1048,2038]" pageId="25" pageNumber="310">
|
||
<keyLead id="309B43637B27FFEAFF1C8DDD85A4F866" pageId="25" pageNumber="310">- Pronotum in lateral view with posterior gibbosity somewhat stronger elevated than anterior; aedeagus as in figs 2F–I or 7... ................................................................................................... 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B27FFE9FF1C8D9885A4FF1C" lastPageId="26" lastPageNumber="311" pageId="25" pageNumber="310">
|
||
<paragraph id="8BD5E0567B27FFEAFF1C8D9885A4F844" blockId="25.[151,1437,1048,2038]" box="[151,1436,2016,2038]" pageId="25" pageNumber="310">
|
||
<keyLead id="309B43637B27FFEAFF1C8D9885A4F844" box="[151,1436,2016,2038]" pageId="25" pageNumber="310">
|
||
8 Head somewhat constricted at base; aedeagus as in fig. 2F–I....................................
|
||
<taxonomicName id="4C6A9BD57B27FFEAFB5C8D988572F844" authorityName="Telnov" authorityYear="2024" box="[1239,1354,2016,2037]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="310" phylum="Arthropoda" rank="species" species="mediator" status="sp. nov.">
|
||
<emphasis id="B91E3C447B27FFEAFB5C8D988572F844" box="[1239,1354,2016,2037]" italics="true" pageId="25" pageNumber="310">N. mediator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91E3C447B27FFEAFADB8D9885A4F844" bold="true" box="[1360,1436,2016,2037]" pageId="25" pageNumber="310">
|
||
<taxonomicNameLabel id="A22D813F7B27FFEAFADB8D9885A4F844" box="[1360,1436,2016,2037]" pageId="25" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8AE085A4FF1C" blockId="26.[151,1437,151,1141]" box="[151,1436,151,173]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8AE085A4FF1C" box="[151,1436,151,173]" pageId="26" pageNumber="311">
|
||
- Head rounded in broad arc posterior to compound eyes; aedeagus as in fig. 7................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FB018AE085A4FF1C" authority="(Marseul, 1876)" baseAuthorityName="Marseul" baseAuthorityYear="1876" box="[1162,1436,151,173]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="trigibber">
|
||
<emphasis id="B91E3C447B24FFE9FB018AE084C3FF1D" box="[1162,1275,151,173]" italics="true" pageId="26" pageNumber="311">N. trigibber</emphasis>
|
||
(Marseul, 1876)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C8ACD85A4FEEC" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8ACD85A4FEB4" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8ACD85A4FEB4" pageId="26" pageNumber="311">
|
||
9 Pronotum in lateral view with anterior gibbosity broad, obtuse angulate; posterior gibbosity slightly less strongly elevated compared to anterior; dorsum generally uniformly brown, narrow pale transverse fascia on elytra in postbasal impression area.............................................................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FB238A8885A4FEB4" authority="(Pic, 1913)" baseAuthorityName="Pic" baseAuthorityYear="1913" box="[1192,1436,239,261]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="bigibbosus">
|
||
<emphasis id="B91E3C447B24FFE9FB238A888514FEB5" box="[1192,1324,239,261]" italics="true" pageId="26" pageNumber="311">N. bigibbosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFB9DA77B24FFE9FAB28A9785ADFEB4" author="Pic, M." box="[1337,1429,239,261]" pageId="26" pageNumber="311" pagination="129 - 135" refId="ref13523" refString="Pic, M. (1913) H. Sauter's Formosa-Ausbeute. Anthicidae, Pedilidae und Hylophilidae. Archiv fur Naturgeschichte, Series A, 79, 129 - 135." type="journal article" year="1913">Pic, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8B7585A4FEEC" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8B7585A4FEEC" pageId="26" pageNumber="311">
|
||
- Pronotum in lateral view with anterior gibbosity rather slender, subacutely angulate; posterior gibbosity same strongly elevated as anterior; pronotum paler than head and elytra, elytra with rather broad pale transverse fascia in postbasal impression area...................................................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FB278B3085A4FEEC" authority="(Pic, 1901)" baseAuthorityName="Pic" baseAuthorityYear="1901" box="[1196,1436,327,349]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91E3C447B24FFE9FB278B308514FEED" box="[1196,1324,327,349]" italics="true" pageId="26" pageNumber="311">N. bidentatus</emphasis>
|
||
(Pic, 1901)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C8B1D85A5FE41" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8B1D85A4FECB" blockId="26.[151,1437,151,1141]" box="[151,1436,357,379]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8B1D85A4FECB" box="[151,1436,357,379]" pageId="26" pageNumber="311">10 Elytra uniformly dark or with pale transverse band or spot in postbasal impression area............................ 12</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8BFB85A5FE41" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8BFB85A5FE41" pageId="26" pageNumber="311">
|
||
- Elytra largely pale yellowish rufous with darkened median area or dark with two pale areas on each elytron—postbasal and postmedian................................................................................................. 11 [some specimens of
|
||
<taxonomicName id="4C6A9BD57B24FFE9FDA98BC58397FE63" authority="(Krekich-Strassoldo, 1914)" baseAuthorityName="Krekich-Strassoldo" baseAuthorityYear="1914" box="[546,943,445,467]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="B91E3C447B24FFE9FDA98BC582AFFE63" box="[546,663,445,466]" italics="true" pageId="26" pageNumber="311">N. arcuatus</emphasis>
|
||
(Krekich-Strassoldo, 1914)
|
||
</taxonomicName>
|
||
are similarly coloured, but aedeagus is different, protibia without modifications, head base broadly rounded, Pronotum in lateral view with two semicircular gibbosities]
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C8B8085A4FDD4" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8B8085A4FD9B" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8B8085A4FD9B" pageId="26" pageNumber="311">
|
||
<specimenCount id="9D6C2BDF7B24FFE9FF1C8B80812BFDBC" box="[151,275,503,525]" count="11" pageId="26" pageNumber="311" type="male">11 Male</specimenCount>
|
||
protibia without modifications; antennomeres 6‒8 not modified (widened / flattened); aedeagus trident-shaped apically, as in fig. 4F–H............................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FC0D886D8488FD9B" authority="(Pic, 1913)" baseAuthorityName="Pic" baseAuthorityYear="1913" box="[902,1200,533,555]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="mediomaculatus">
|
||
<emphasis id="B91E3C447B24FFE9FC0D886D8407FD9B" box="[902,1087,533,554]" italics="true" pageId="26" pageNumber="311">N. mediomaculatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFB9DA77B24FFE9FBC7886D8491FD9B" author="Pic, M." box="[1100,1193,533,555]" pageId="26" pageNumber="311" pagination="129 - 135" refId="ref13523" refString="Pic, M. (1913) H. Sauter's Formosa-Ausbeute. Anthicidae, Pedilidae und Hylophilidae. Archiv fur Naturgeschichte, Series A, 79, 129 - 135." type="journal article" year="1913">Pic, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B91E3C447B24FFE9FB3D886D851AFD9A" bold="true" box="[1206,1314,533,555]" pageId="26" pageNumber="311">stat. restit.</emphasis>
|
||
,
|
||
<emphasis id="B91E3C447B24FFE9FAA5886D85A4FD9B" bold="true" box="[1326,1436,533,554]" pageId="26" pageNumber="311">comb. nov.</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C884B85A4FDD4" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C884B85A4FDD4" pageId="26" pageNumber="311">
|
||
- Male protibia distinctly sinuous; antennomeres 6‒8 widened and somewhat dorso-ventrally flattened; aedeagus not as in fig. 4................................................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FBBB882885A4FDD4" authority="(Krekich-Strassoldo, 1913)" baseAuthorityName="Krekich-Strassoldo" baseAuthorityYear="1913" box="[1072,1436,591,613]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="lepidus">
|
||
<emphasis id="B91E3C447B24FFE9FBBB882884A9FDD5" box="[1072,1169,591,613]" italics="true" pageId="26" pageNumber="311">N. lepidus</emphasis>
|
||
(Krekich-Strassoldo, 1913)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C881585A4FD2E" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C881585A4FD33" blockId="26.[151,1437,151,1141]" box="[151,1436,621,643]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C881585A4FD33" box="[151,1436,621,643]" pageId="26" pageNumber="311">12 Elytra uniformly coloured, without pale spots or bands....................................................... 13</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C88F385A4FD2E" blockId="26.[151,1437,151,1141]" box="[151,1436,650,672]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C88F385A4FD2E" box="[151,1436,650,672]" pageId="26" pageNumber="311">- Elytra with more or less prominent pale spot or transverse band in postbasal transverse impression area................ 14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C88D085A4FC83" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C88D085A4FD46" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C88D085A4FD46" pageId="26" pageNumber="311">
|
||
13 Head distinctly constricted posterior to compound eyes; pronotum slender, about as wide as head between compound eyes, in dorsal view hourglass-shaped, anterior lobe significantly wider than posterior; metatibia not modified (widened or thickened).......................................................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FB11889B85A4FD46" authority="(Pic, 1914)" baseAuthorityName="Pic" baseAuthorityYear="1914" box="[1178,1436,738,760]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="subconiceps">
|
||
<emphasis id="B91E3C447B24FFE9FB11889B8513FD46" box="[1178,1323,738,760]" italics="true" pageId="26" pageNumber="311">N. subconiceps</emphasis>
|
||
(Pic, 1914)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C897885A4FC83" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C897885A4FC83" pageId="26" pageNumber="311">
|
||
- Head rounded posterior to compound eyes; pronotum in dorsal view either subcylindrical with bulged anterior lobe, not hourglass-shaped, anterior lobe less strongly wider than posterior; metatibia slightly widened........
|
||
<taxonomicName id="4C6A9BD57B24FFE9FB4D896585A4FC83" authority="(Pic, 1901)" baseAuthorityName="Pic" baseAuthorityYear="1901" box="[1222,1436,797,819]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis id="B91E3C447B24FFE9FB4D89658513FC83" box="[1222,1323,797,818]" italics="true" pageId="26" pageNumber="311">N. collaris</emphasis>
|
||
(Pic, 1901)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C894285A4FC16" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C894285A4FCDC" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C894285A4FCDC" pageId="26" pageNumber="311">14 Pronotum or entire dorsal forebody distinctly paler than elytra; two or several basal antennomeres distinctly paler than rest of antenna............................................................................................ 15</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C890D85A4FC16" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C890D85A4FC16" pageId="26" pageNumber="311">
|
||
- Dorsal forebody about the same colour as elytra; basal (one) antennomere distinctly paler than rest of antenna..........................................................................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FB0189EB85A4FC16" authority="(Pic, 1922)" baseAuthorityName="Pic" baseAuthorityYear="1922" box="[1162,1436,914,936]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="luteofasciatus">
|
||
<emphasis id="B91E3C447B24FFE9FB0189EB8513FC16" box="[1162,1323,914,936]" italics="true" pageId="26" pageNumber="311">N. luteofasciatus</emphasis>
|
||
(Pic, 1922)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309EF8F37B24FFE9FF1C89D785A5FBC4" pageId="26" pageNumber="311">
|
||
<paragraph id="8BD5E0567B24FFE9FF1C89D785A4FC4E" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C89D785A4FC4E" pageId="26" pageNumber="311">
|
||
15 Posterior lobe of pronotum entire, not bilobate; profemur yellowish, protibia entirely or at least medially dark; compound eye comparatively smaller; elytra slightly convex with moderately prominent postbasal transverse impression; metatibia slightly arched................................................................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FBA9899385A4FC4E" authority="(Krekich-Strassoldo, 1914)" baseAuthorityName="Krekich-Strassoldo" baseAuthorityYear="1914" box="[1058,1436,1002,1024]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="arcuatus">
|
||
<emphasis id="B91E3C447B24FFE9FBA9899384A9FBB1" box="[1058,1169,1003,1024]" italics="true" pageId="26" pageNumber="311">N. arcuatus</emphasis>
|
||
(Krekich-Strassoldo, 1914)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD5E0567B24FFE9FF1C8E7085A5FBC4" blockId="26.[151,1437,151,1141]" pageId="26" pageNumber="311">
|
||
<keyLead id="309B43637B24FFE9FF1C8E7085A5FBC4" pageId="26" pageNumber="311">
|
||
- Posterior lobe of pronotum bilobate (with a short median longitudinal impression and a lateral gibbosity either side of it); protibia not contrastingly darker than profemur; compound eye comparatively larger; elytra flattened, without or with vaguely defined postbasal transverse impression; metatibia not modified.......................
|
||
<taxonomicName id="4C6A9BD57B24FFE9FBEB8E3B8125FBC4" authority="(Krekich-Strassoldo, 1929)" baseAuthorityName="Krekich-Strassoldo" baseAuthorityYear="1929" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="bifidus">
|
||
<emphasis id="B91E3C447B24FFE9FBEB8E3B84FAFBE6" box="[1120,1218,1090,1112]" italics="true" pageId="26" pageNumber="311">N. bifidus</emphasis>
|
||
(Krekich-Strassoldo, 1929)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6A9BD57B24FFE9FEC58E188256FBC4" authority="(Bonadona, 1981)" baseAuthorityName="Bonadona" baseAuthorityYear="1981" box="[334,622,1119,1141]" class="Insecta" family="Anthicidae" genus="Nitorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="311" phylum="Arthropoda" rank="species" species="gibber">
|
||
<emphasis id="B91E3C447B24FFE9FEC58E188196FBC5" box="[334,430,1119,1141]" italics="true" pageId="26" pageNumber="311">N. gibber</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFB9DA77B24FFE9FE308E27825EFBC4" author="Bonadona, P." box="[443,614,1119,1141]" pageId="26" pageNumber="311" pagination="193 - 212" refId="ref12991" refString="Bonadona, P. (1981) Anthicidae inedits (Coleoptera) des Iles Philippines et l'Archipel Bismarck (Insecta, Coleoptera). Steenstrupia, 7 (8), 193 - 212." type="journal article" year="1981">Bonadona, 1981</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, likely conspecific, the differential diagnosis in
|
||
<bibRefCitation id="EFFB9DA77B24FFE9FBD38E278530FBC4" author="Bonadona, P." box="[1112,1288,1119,1141]" pageId="26" pageNumber="311" pagination="193 - 212" refId="ref12991" refString="Bonadona, P. (1981) Anthicidae inedits (Coleoptera) des Iles Philippines et l'Archipel Bismarck (Insecta, Coleoptera). Steenstrupia, 7 (8), 193 - 212." type="journal article" year="1981">Bonadona (1981)</bibRefCitation>
|
||
is suboptimal.
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |