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<document id="DFBD9FD70C4DA140BAC9B544A081F44D" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFCD1F5DFF50FF27FC168C7B" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax gonaivensis Henderson, Fernandez &amp; Bacon" docType="treatment" docVersion="2" lastPageNumber="51" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="51" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:title id="1D5B2DEDC652D36A2E6C720DCA34836E">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<treatment id="03B387DAFFCD1F5DFF50FF27FC168C7B" ID-DOI="http://doi.org/10.5281/zenodo.8389303" ID-Zenodo-Dep="8389303" LSID="urn:lsid:plazi:treatment:03B387DAFFCD1F5DFF50FF27FC168C7B" httpUri="http://treatment.plazi.org/id/03B387DAFFCD1F5DFF50FF27FC168C7B" lastPageNumber="51" pageId="50" pageNumber="51">
<subSubSection id="C3006547FFCD1F5DFF50FF27FB1C891B" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8BA536CCFFCD1F5DFF50FF27FB1C891B" blockId="50.[136,1452,157,257]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF50FF27FB0C88A3" bold="true" box="[136,1269,157,185]" pageId="50" pageNumber="51">
1.16.
<taxonomicName id="4C1A4D4FFFCD1F5DFF1FFF27FC8288A3" authority="Henderson, Fernandez &amp; Bacon" authorityName="Henderson, Fernandez &amp; Bacon" box="[199,891,157,185]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="gonaivensis">
<emphasis id="B96EEADEFFCD1F5DFF1FFF27FE0B88A3" bold="true" box="[199,498,158,185]" italics="true" pageId="50" pageNumber="51">Coccothrinax gonaivensis</emphasis>
Henderson, Fernández &amp; Bacon
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFCD1F5DFC78FF27FB1688A3" author="Henderson, A. &amp; Fernandez, E. &amp; Montero, O. &amp; Clase, T. &amp; Gratacos, X. &amp; Bacon, C." box="[928,1263,158,185]" pageId="50" pageNumber="51" pagination="101 - 111" refId="ref62198" refString="Henderson, A., Fernandez, E., Montero, O., Clase, T., Gratacos, X. &amp; Bacon, C. (2023) Five new species of Coccothrinax (Cryosophileae, Coryphoideae, Arecaceae) from Hispaniola. Phytotaxa 589 (2): 101 - 111. https: // doi. org / 10.11646 / phytotaxa. 589.2.1" type="journal article" year="2023">
Henderson
<emphasis id="B96EEADEFFCD1F5DFBF2FF26FB9B88A2" bold="true" box="[1066,1122,158,185]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
(2023: 108)
</bibRefCitation>
.
</emphasis>
<materialsCitation id="3B723C91FFCD1F5DFB25FF27FB1C891B" collectingDate="1985-06-11" collectionCode="T, M, R, JBSD" collectorName="de Gonaives" country="Haiti" elevation="100" latitude="19.466667" location="Artibonite" longLatPrecision="1271" longitude="-72.71667" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Artibonite" typeStatus="holotype">
Type:—
<collectingCountry id="F30D765CFFCD1F5DFA81FF27FA5188A3" box="[1369,1448,158,184]" name="Haiti" pageId="50" pageNumber="51">HAITI</collectingCountry>
. Dept.
<collectingRegion id="49DEF82EFFCD1F5DFF12FF7BFEB988C7" box="[202,320,194,220]" country="Haiti" name="Artibonite" pageId="50" pageNumber="51">Artibonite</collectingRegion>
, Presquile du
<collectingRegion id="49DEF82EFFCD1F5DFE3AFF7BFD9088C7" box="[482,617,194,220]" country="Haiti" name="Nord-Ouest" pageId="50" pageNumber="51">Nord-Ouest</collectingRegion>
,
<quantity id="4CE29B29FFCD1F5DFDABFF7BFD3988C7" box="[627,704,194,221]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="50" pageNumber="51" unit="km" value="5.6">5.6 km</quantity>
al oeste
<collectorName id="26EF531AFFCD1F5DFCC7FF7BFC5088C7" box="[799,937,194,220]" pageId="50" pageNumber="51">de Gonaives</collectorName>
en la carretera costera a Anse Rouge,
<geoCoordinate id="EE2E500BFFCD1F5DFA93FF7BFA5288C7" box="[1355,1451,194,220]" degrees="19" direction="north" minutes="28" orientation="latitude" pageId="50" pageNumber="51" precision="925" value="19.466667">19°28N</geoCoordinate>
<geoCoordinate id="EE2E500BFFCD1F5DFF50FF5FFF16891B" box="[136,239,230,256]" degrees="72" direction="west" minutes="43" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-72.71667">72°43W</geoCoordinate>
,
<quantity id="4CE29B29FFCD1F5DFF22FF5FFEBA891A" box="[250,323,230,257]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="51" unit="m" value="100.0">
<elevation id="0037D1FFFFCD1F5DFF22FF5FFEBA891A" box="[250,323,230,257]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="51" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="FFA4100CFFCD1F5DFE97FF5FFE1E891A" box="[335,487,230,257]" pageId="50" pageNumber="51" value="1985-06-11">
<collectingDate id="EFE0E9E4FFCD1F5DFE97FF5FFE1E891A" box="[335,487,230,257]" pageId="50" pageNumber="51" value="1985-06-11">11 June 1985</collectingDate>
</date>
,
<emphasis id="B96EEADEFFCD1F5DFE2AFF5EFC56891B" box="[498,943,230,257]" italics="true" pageId="50" pageNumber="51">
<collectionCode id="ED0BAE09FFCD1F5DFE2AFF5EFDFA891B" box="[498,515,231,256]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="51">T</collectionCode>
. Zanoni,
<collectionCode id="ED0BAE09FFCD1F5DFDB2FF5EFD7C891B" box="[618,645,231,256]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="50" pageNumber="51" type="Herbarium">M</collectionCode>
. Mejía &amp;
<collectionCode id="ED0BAE09FFCD1F5DFD2DFF5EFCF0891B" box="[757,777,231,256]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="50" pageNumber="51">R</collectionCode>
. García 35154
</emphasis>
(
<typeStatus id="54A1886EFFCD1F5DFC66FF5FFBDB891B" box="[958,1058,230,256]" pageId="50" pageNumber="51">holotype</typeStatus>
<collectionCode id="ED0BAE09FFCD1F5DFBF0FF5FFB8A891B" box="[1064,1139,230,256]" country="Dominican Republic" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15541" lsid="urn:lsid:biocol.org:col:15541" name="Jardín Botánico Nacional Dr. Rafael M. Moscoso" pageId="50" pageNumber="51" type="Herbarium">JBSD</collectionCode>
!).
<figureCitation id="13212A49FFCD1F5DFB52FF5FFB1C891B" box="[1162,1253,230,256]" captionStart="PLATE 13" captionStartId="51.[136,214,1917,1938]" captionTargetBox="[151,1433,211,1891]" captionTargetId="figure-15@51.[150,1434,193,1893]" captionTargetPageId="51" captionText="PLATE 13. Coccothrinax gonaivensis, Haiti, with wedge-shaped leaf blades and segments not pendulous at the apices, giving the leaf a flat appearance. (Image by Eladio Fernández)." figureDoi="http://doi.org/10.5281/zenodo.8389383" httpUri="https://zenodo.org/record/8389383/files/figure.png" pageId="50" pageNumber="51">Plate 13</figureCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFCD1F5DFF50FE97FD278B7E" pageId="50" pageNumber="51" type="description">
<paragraph id="8BA536CCFFCD1F5DFF50FE97FD278B7E" blockId="50.[136,1452,302,1121]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF50FE97FF298953" bold="true" box="[136,208,302,328]" pageId="50" pageNumber="51">Stems</emphasis>
5.0(2.08.0) m long and
<quantity id="4CE29B29FFCD1F5DFE05FE97FDDF8952" box="[477,550,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="50" pageNumber="51" unit="cm" value="3.8">3.8 cm</quantity>
diameter,branching not recorded.
<emphasis id="B96EEADEFFCD1F5DFC46FE97FC098953" bold="true" box="[926,1008,302,328]" pageId="50" pageNumber="51">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 0.4(0.20.5) mm diameter, stout, loosely woven and forming a loose, hexagonal mesh, initially forming ligules at the apices; petioles 7.1(6.67.6) mm diameter just below the apex; palmans
<quantity id="4CE29B29FFCD1F5DFA86FECFFA52898A" box="[1374,1451,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="50" pageNumber="51" unit="cm" value="2.2">2.2 cm</quantity>
long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades wedge-shaped; segments 22(2123) per leaf, the middle ones 51.8(46.557.0) cm long and 1.9(1.82.0) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments tapering from base to apex, often folded, stiff and leathery, with or without scarcely developed shoulders, the apices sharply pointed and briefly splitting; middle leaf segment apices attenuate; leaf segments with not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, semi-persistent, interlocking, fimbriate hairs without an obvious center, without or with poorly developed transverse veinlets.
<emphasis id="B96EEADEFFCD1F5DFB98FDCBFB108A97" bold="true" box="[1088,1257,626,652]" pageId="50" pageNumber="51">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 4; proximalmost rachillae straight,
<quantity id="4CE29B29FFCD1F5DFB21FD03FABD8ACE" box="[1273,1348,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="50" pageNumber="51" unit="cm" value="3.8">3.8 cm</quantity>
long and
<quantity id="4CE29B29FFCD1F5DFF50FD67FF198AE2" box="[136,224,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="50" pageNumber="51" unit="mm" value="0.9">0.9 mm</quantity>
diameter in fruit; rachillae uneven at or near anthesis with lines of warty outgrowths, these often becoming more pronounced as fruits develop; stamens not recorded; fruit pedicels
<quantity id="4CE29B29FFCD1F5DFC77FCBBFBFD8B06" box="[943,1028,770,797]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="51" unit="mm" value="0.1">0.1 mm</quantity>
long;
<emphasis id="B96EEADEFFCD1F5DFB93FCBBFB758B07" bold="true" box="[1099,1164,770,796]" pageId="50" pageNumber="51">fruits</emphasis>
<quantity id="4CE29B29FFCD1F5DFB4BFCBBFB118B06" box="[1171,1256,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="50" pageNumber="51" unit="mm" value="3.9">3.9 mm</quantity>
long and 4.0 mm diameter, color not recorded; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces lobed, the lobes running from base of seeds approximately to equator.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFCD1F5DFF65FCD7FC168C7B" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8BA536CCFFCD1F5DFF65FCD7FABB8B93" blockId="50.[136,1452,302,1121]" box="[189,1346,878,905]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF65FCD7FDFF8B93" bold="true" box="[189,518,878,904]" pageId="50" pageNumber="51">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFCD1F5DFDDEFCD7FDC78B93" box="[518,574,878,904]" name="Haiti" pageId="50" pageNumber="51">Haiti</collectingCountry>
(
<collectingRegion id="49DEF82EFFCD1F5DFD94FCD7FD3A8B93" box="[588,707,878,904]" country="Haiti" name="Artibonite" pageId="50" pageNumber="51">Artibonite</collectingRegion>
) (
<figureCitation id="13212A49FFCD1F5DFD02FCD7FCD28B93" box="[730,811,878,904]" captionStart="FIGURE 12" captionStartId="47.[136,229,1777,1799]" captionTargetBox="[155,1429,469,1747]" captionTargetId="figure-147@47.[150,1434,464,1752]" captionTargetPageId="47" captionText="FIGURE 12. Distribution of Coccothrinax fagildei, C. fragrans, C. garciana, and C. gonaivensis." figureDoi="http://doi.org/10.5281/zenodo.8389379" httpUri="https://zenodo.org/record/8389379/files/figure.png" pageId="50" pageNumber="51">Fig. 12</figureCitation>
) on limestone scree at 86(72100) m elevation.
</paragraph>
<paragraph id="8BA536CCFFCD1F5DFF65FC2BFC168C7B" blockId="50.[136,1452,302,1121]" pageId="50" pageNumber="51">
<emphasis id="B96EEADEFFCD1F5DFF65FC2BFE528BB7" bold="true" box="[189,427,914,940]" pageId="50" pageNumber="51">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFCD1F5DFD12FC2BFC148BB7" box="[714,1005,914,940]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="gonaivensis">
<emphasis id="B96EEADEFFCD1F5DFD12FC2BFC148BB7" box="[714,1005,914,940]" italics="true" pageId="50" pageNumber="51">Coccothrinax gonaivensis</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. It is a member of a group of nine Hispaniolan species (see notes under
<taxonomicName id="4C1A4D4FFFCD1F5DFECCFC62FE508BEF" authorityName="Mejia &amp; Garcia" authorityYear="1997" box="[276,425,986,1013]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="boschiana">
<emphasis id="B96EEADEFFCD1F5DFECCFC62FE508BEF" box="[276,425,986,1013]" italics="true" pageId="50" pageNumber="51">C. boschiana</emphasis>
</taxonomicName>
). In this group it is similar to
<taxonomicName id="4C1A4D4FFFCD1F5DFCDEFC62FC748BEF" authorityName="Mejia &amp; Garcia" authorityYear="2013" box="[774,909,987,1013]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="jimenezii">
<emphasis id="B96EEADEFFCD1F5DFCDEFC62FC748BEF" box="[774,909,987,1013]" italics="true" pageId="50" pageNumber="51">C. jimenezii</emphasis>
</taxonomicName>
but differs in its leaf segments that are densely indumentose abaxially. Specimens were previously included in
<taxonomicName id="4C1A4D4FFFCD1F5DFC88FC46FC2F8C03" authorityName="Mejia &amp; Garcia" authorityYear="2013" box="[848,982,1023,1049]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="jimenezii">
<emphasis id="B96EEADEFFCD1F5DFC88FC46FC2F8C03" box="[848,982,1023,1049]" italics="true" pageId="50" pageNumber="51">C. jimenezii</emphasis>
</taxonomicName>
, but
<bibRefCitation id="EF8B4B3DFFCD1F5DFBD6FC46FB0C8C03" author="Jestrow, B. &amp; Peguero, B. &amp; Jimenez, F. &amp; Cinea, W. &amp; Hass, M. &amp; Reeve, A. &amp; Meerow, A. &amp; Griffith, P. &amp; Maunder M. &amp; Francisco-Ortega, J." box="[1038,1269,1022,1049]" pageId="50" pageNumber="51" pagination="216 - 223" refId="ref62452" refString="Jestrow, B., Peguero, B., Jimenez, F., Cinea, W., Hass, M., Reeve, A., Meerow, A., Griffith, P., Maunder M. &amp; Francisco-Ortega, J. (2016) Genetic diversity and differentiation of the Critically Endangered Hispaniolan palm Coccothrinax jimenezii M. M. Mejia and R. G. Garcia based on novel SSR markers. Biochemical Systematics and Ecology 66: 216 - 223. https: // doi. org / 10.1016 / j. bse. 2016.04.013" type="journal article" year="2016">
Jestrow
<emphasis id="B96EEADEFFCD1F5DFBB3FC46FB5A8C03" box="[1131,1187,1022,1048]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
(2016)
</bibRefCitation>
showed that the two represented different taxa. Inflorescences of the
<typeStatus id="54A1886EFFCD1F5DFD09FB9AFCF88C26" box="[721,769,1059,1085]" pageId="50" pageNumber="51">type</typeStatus>
are branched to three orders.
<bibRefCitation id="EF8B4B3DFFCD1F5DFB93FB9AFACF8C26" author="Peguero, B. &amp; Jimenez, F. &amp; Joseph, P. &amp; Cinea, W. &amp; Griffith, P. &amp; Francisco-Ortega, J. &amp; Jestrow, B." box="[1099,1334,1058,1085]" pageId="50" pageNumber="51" pagination="145 - 153" refId="ref63894" refString="Peguero, B., Jimenez, F., Joseph, P., Cinea, W., Griffith, P., Francisco-Ortega, J. &amp; Jestrow, B. (2015) Coccothrinax jimenezii - a critically endangered palm from Hispaniola. Palms 59: 145 - 153." type="journal article" year="2015">
Peguero
<emphasis id="B96EEADEFFCD1F5DFB76FB9AFB1C8C27" box="[1198,1253,1058,1084]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
(2015)
</bibRefCitation>
noted that the small population consisted mostly of juveniles that were not reproductive.
</paragraph>
</subSubSection>
</treatment>
</document>