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:
<treatmentCitation id="0AA98F2B9E7D3A3ACC6E42504BBEFCE7" author="Tirira, D. &amp; J. Brito &amp; S. F. Burneo &amp; J. P. Carrera-Estupinan &amp; Comision de Diversidad de la AEM" box="[269,496,854,878]" page="33" pageId="10" pageNumber="11" year="2022">
<bibRefCitation id="EF99D4CB9E7D3A3ACC6E42504BBEFCE7" author="Tirira" box="[269,496,854,878]" etAl="et al." firstAuthor="Tirira" page="33" pageId="10" pageNumber="11" refId="ref8593" refString="Tirira, D., J. Brito, S. F. Burneo, J. P. Carrera-Estupinan, and Comision de Diversidad de la AEM. 2022. Mammals of Ecuador: official updated species checklist. Version 2022.1. Quito: Asociacion Ecuatoriana de Mastozoologia. Internet resource (http: // aem. mamiferosdelecuador. com), accessed June, 3 2022." type="url" year="2022">Tirira et al., 2022: 33</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C312FAB19E7D3A3ACDFF428A4EDEFB22" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BB7A93A9E7D3A3ACDFF428A4906FC78" blockId="10.[108,1214,908,1756]" pageId="10" pageNumber="11">
<materialsCitation id="3B60A3679E7D3A3ACDFF428A4906FC78" collectingDate="2009-03-12" collectionCode="QCAZ" collectorName="Carlos Boada" country="Ecuador" elevation="1340" latitude="-4.258" location="Las Orquideas" longLatPrecision="13" longitude="-78.6765" pageId="10" pageNumber="11" specimenCode="QCAZ 11122" specimenCount="1" specimenCount-female="1" stateProvince="Zamora Chinchipe province" typeStatus="holotype">
<typeStatus id="54B317989E7D3A3ACDFF428A4B55FC2F" box="[156,283,908,934]" pageId="10" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
: An
<specimenCount id="9D0E62B39E7D3A3ACC30428A4BADFC2F" box="[339,483,908,934]" pageId="10" pageNumber="11" type="adult">adult female</specimenCount>
(
<specimenCode id="DBAE01419E7D3A3ACC92428A48DDFC2F" box="[497,659,908,934]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ 11122</specimenCode>
) collected on
<date id="FFB68FFA9E7D3A3ACE56428A49A2FC2F" box="[821,1004,908,934]" pageId="10" pageNumber="11" value="2009-03-12">
<collectingDate id="EFF276129E7D3A3ACE56428A49A2FC2F" box="[821,1004,908,934]" pageId="10" pageNumber="11" value="2009-03-12">March 12, 2009</collectingDate>
</date>
, by
<collectorName id="26FDCCEC9E7D3A3AC97A428A4EF7FC2F" box="[1049,1209,908,934]" pageId="10" pageNumber="11">Carlos Boada</collectorName>
. The body is preserved in 75% ethanol, with the skull removed and cleaned. Muscle and liver tissues preserved in 95% ethanol are also deposited at
<collectionCode id="ED1931FF9E7D3A3ACF9242D1490AFC78" box="[753,836,983,1009]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB7A93A9E7D3A3ACDFF42FB4996FBB5" blockId="10.[108,1214,908,1756]" pageId="10" pageNumber="11">
TYPE LOCALITY:
<location id="8ED7FFE19E7D3A3ACC3E42FB484BFB9F" LSID="urn:lsid:plazi:treatment:03A1182C9E7D3A3FCF6340FF49FAFA25:8ED7FFE19E7D3A3ACC3E42FB484BFB9F" box="[349,517,1020,1047]" country="Ecuador" county="Nangaritza River" latitude="-4.258" longLatPrecision="13" longitude="-78.6765" municipality="Miazi Alto" name="Las Orquideas" pageId="10" pageNumber="11" stateProvince="Zamora Chinchipe province">Las Orquideas</location>
,
<locationDeviation id="9A05B0189E7D3A3ACF7342FA499BFB9F" box="[528,981,1020,1047]" pageId="10" pageNumber="11">Miazi Alto near Nangaritza River basin</locationDeviation>
,
<collectingRegion id="49CC67D89E7D3A3ACE8342FB4A9BFBB2" country="Ecuador" name="Zamora-Chinchipe" pageId="10" pageNumber="11">Zamora Chinchipe province</collectingRegion>
,
<collectingCountry id="F31FE9AA9E7D3A3ACD8245274B0BFBB2" box="[225,325,1057,1083]" name="Ecuador" pageId="10" pageNumber="11">Ecuador</collectingCountry>
,
<geoCoordinate id="EE3CCFFD9E7D3A3ACC3345274B9EFBB2" box="[336,464,1057,1084]" degrees="04" direction="south" minutes="15.48" orientation="latitude" pageId="10" pageNumber="11" precision="9" value="-4.258">04°15.48S</geoCoordinate>
,
<geoCoordinate id="EE3CCFFD9E7D3A3ACCBF45274828FBB5" box="[476,614,1057,1084]" degrees="78" direction="west" minutes="40.59" orientation="longitude" pageId="10" pageNumber="11" precision="9" value="-78.6765">78°40.59W</geoCoordinate>
, between
<quantity id="4CF004DF9E7D3A3ACFBF45274932FBB5" box="[732,892,1057,1084]" metricMagnitude="3" metricUnit="m" metricValue="1.34" metricValueMax="1.43" metricValueMin="1.25" pageId="10" pageNumber="11" unit="m" value="1340.0" valueMax="1430.0" valueMin="1250.0">
<elevation id="00254E099E7D3A3ACFBF45274932FBB5" box="[732,892,1057,1084]" metricMagnitude="3" metricUnit="m" metricValue="1.34" metricValueMax="1.43" metricValueMin="1.25" pageId="10" pageNumber="11" unit="m" value="1340.0" valueMax="1430.0" valueMin="1250.0">12501430 m</elevation>
</quantity>
(
<figureCitation id="1333B5BF9E7D3A3ACEEF45274984FBB5" box="[908,970,1057,1084]" captionStart="FIGURE 5" captionStartId="12.[108,194,1499,1520]" captionTargetBox="[108,1212,224,1473]" captionTargetId="figure-51@12.[108,1212,224,1473]" captionTargetPageId="12" captionText="FIGURE 5. Map of NW South America showing the collection localities of Sturnira bidens (circles), S. nana (diamonds), and Sturnira boadai, sp. nov. (stars), analyzed in this study. The collection points of Sturnira nana overlap, so only two diamonds are observed." figureDoi="http://doi.org/10.5281/zenodo.10261227" httpUri="https://zenodo.org/record/10261227/files/figure.png" pageId="10" pageNumber="11">fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BB7A93A9E7D3A3ACDFF45414EDEFB22" blockId="10.[108,1214,908,1756]" pageId="10" pageNumber="11">
<materialsCitation id="3B60A3679E7D3A3ACDFF45414EDEFB22" collectingDate="2009-03-12" collectionCode="QCAZ" collectorName="Carlos Boada" country="Ecuador" elevation="1340" latitude="-4.258" location="Las Orquideas" longLatPrecision="13" longitude="-78.6765" pageId="10" pageNumber="11" specimenCode="QCAZ11120, QCAZ11116, QCAZ11119, QCAZ11121, QCAZ11123" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Zamora Chinchipe province" typeStatus="paratype">
<typeStatus id="54B317989E7D3A3ACDFF45414B6CFBE8" box="[156,290,1095,1121]" pageId="10" pageNumber="11" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D0E62B39E7D3A3ACC5445414B87FBE8" box="[311,457,1095,1121]" count="1" pageId="10" pageNumber="11" type="female">One female</specimenCount>
(
<specimenCode id="DBAE01419E7D3A3ACCBC45414833FBE8" box="[479,637,1095,1121]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ11120</specimenCode>
) and
<specimenCount id="9D0E62B39E7D3A3ACFAD4541491AFBE8" box="[718,852,1095,1121]" count="4" pageId="10" pageNumber="11" type="male">four males</specimenCount>
(
<specimenCode id="DBAE01419E7D3A3ACE0845414E47FBE8" box="[875,1033,1095,1121]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ11116</specimenCode>
,
<specimenCode id="DBAE01419E7D3A3AC97F45414EF6FBE8" box="[1052,1208,1095,1121]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ11119</specimenCode>
,
<specimenCode id="DBAE01419E7D3A3ACD0F456A4B4AFB0F" box="[108,260,1132,1158]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ11121</specimenCode>
,
<specimenCode id="DBAE01419E7D3A3ACC6D456A4BEBFB0F" box="[270,421,1132,1158]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="10" pageNumber="11" type="Museum">QCAZ11123</specimenCode>
) were also collected at the type locality on
<date id="FFB68FFA9E7D3A3ACEF0456A4E09FB0F" box="[915,1095,1132,1158]" pageId="10" pageNumber="11" value="2009-03-12">
<collectingDate id="EFF276129E7D3A3ACEF0456A4E09FB0F" box="[915,1095,1132,1158]" pageId="10" pageNumber="11" value="2009-03-12">March 12, 2009</collectingDate>
</date>
by
<collectorName id="26FDCCEC9E7D3A3AC913456A4AF7FB22" pageId="10" pageNumber="11">Carlos Boada</collectorName>
. All specimens are preserved in 75% ethanol, with the skulls removed and cleaned.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C312FAB19E7D3A3ACDFF45B14B85FAEE" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BB7A93A9E7D3A3ACDFF45B14B85FAEE" blockId="10.[108,1214,908,1756]" pageId="10" pageNumber="11">
DISTRIBUTION: Known from two confirmed localities in
<collectingRegion id="49CC67D89E7D3A3ACE5B45B14EC4FB58" box="[824,1162,1207,1233]" country="Ecuador" name="Zamora-Chinchipe" pageId="10" pageNumber="11">Zamora Chinchipe Province</collectingRegion>
: Las Orquideas, Miazi Alto near
<location id="8ED7FFE19E7D3A3ACCD045DA4837FB7F" LSID="urn:lsid:plazi:treatment:03A1182C9E7D3A3FCF6340FF49FAFA25:8ED7FFE19E7D3A3ACCD045DA4837FB7F" box="[435,633,1244,1270]" country="Ecuador" latitude="-3.6355555" longLatPrecision="21" longitude="-78.38944" name="Nangaritza River" pageId="10" pageNumber="11" stateProvince="Zamora-Chinchipe">Nangaritza River</location>
basin (
<geoCoordinate id="EE3CCFFD9E7D3A3ACFAE45DA4937FB7F" box="[717,889,1243,1271]" degrees="04" direction="south" minutes="15" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="29.30" value="-4.2581387">04°1529.30″ S</geoCoordinate>
;
<geoCoordinate id="EE3CCFFD9E7D3A3ACEE645DA4E70FB7F" box="[901,1086,1243,1271]" degrees="78" direction="west" minutes="40" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="53.40" value="-78.681496">78°4053.40″ W</geoCoordinate>
) and Military Detachment Cóndor Mirador,
<collectorName id="26FDCCEC9E7D3A3ACF77440748C8FA92" box="[532,646,1281,1307]" pageId="10" pageNumber="11">El Pangui</collectorName>
(
<geoCoordinate id="EE3CCFFD9E7D3A3ACFFB4407496BFA92" box="[664,805,1281,1308]" degrees="03" direction="south" minutes="38" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="08" value="-3.6355555">03°3808″ S</geoCoordinate>
;
<geoCoordinate id="EE3CCFFD9E7D3A3ACE5044074981FA92" box="[819,975,1281,1308]" degrees="78" direction="west" minutes="23" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="22" value="-78.38944">78°2322″ W</geoCoordinate>
) near the border of
<collectingCountry id="F31FE9AA9E7D3A3ACD0F44214A81FAC8" box="[108,207,1319,1345]" name="Ecuador" pageId="10" pageNumber="11">Ecuador</collectingCountry>
and
<collectingCountry id="F31FE9AA9E7D3A3ACC6B44214B71FAC8" box="[264,319,1319,1345]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
(
<figureCitation id="1333B5BF9E7D3A3ACC2D44214BC5FAC8" box="[334,395,1318,1345]" captionStart="FIGURE 5" captionStartId="12.[108,194,1499,1520]" captionTargetBox="[108,1212,224,1473]" captionTargetId="figure-51@12.[108,1212,224,1473]" captionTargetPageId="12" captionText="FIGURE 5. Map of NW South America showing the collection localities of Sturnira bidens (circles), S. nana (diamonds), and Sturnira boadai, sp. nov. (stars), analyzed in this study. The collection points of Sturnira nana overlap, so only two diamonds are observed." figureDoi="http://doi.org/10.5281/zenodo.10261227" httpUri="https://zenodo.org/record/10261227/files/figure.png" pageId="10" pageNumber="11">fig. 5</figureCitation>
). We expect that
<taxonomicName id="4C08D2B99E7D3A3ACF364421488EFAC8" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[597,704,1319,1345]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E7D3A3ACF364421488EFAC8" box="[597,704,1319,1345]" italics="true" pageId="10" pageNumber="11">S. boadai</emphasis>
</taxonomicName>
also occurs in adjacent, climatically similar habitats of northeastern
<collectingCountry id="F31FE9AA9E7D3A3ACCED444B4B86FAEE" box="[398,456,1357,1383]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C312FAB19E7D3A3BCDFF44774BFBFB58" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BB7A93A9E7D3A3BCDFF44774BFBFB58" blockId="10.[108,1214,908,1756]" lastBlockId="11.[108,1214,1020,1756]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
DIAGNOSIS:
<taxonomicName id="4C08D2B99E7D3A3ACC4F44744BACFA02" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[300,482,1393,1420]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E7D3A3ACC4F44744BACFA02" box="[300,482,1393,1420]" italics="true" pageId="10" pageNumber="11">Sturnira boadai</emphasis>
</taxonomicName>
is a small species (FA =
<quantity id="4CF004DF9E7D3A3ACE67447749E1FA05" box="[772,943,1393,1420]" metricMagnitude="-2" metricUnit="m" metricValue="3.315" metricValueMax="3.38" metricValueMin="3.25" pageId="10" pageNumber="11" unit="mm" value="33.15" valueMax="33.8" valueMin="32.5">32.533.8 mm</quantity>
, GLS =
<quantity id="4CF004DF9E7D3A3AC96D44774EF6FA05" box="[1038,1208,1393,1420]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" metricValueMax="1.92" metricValueMin="1.83" pageId="10" pageNumber="11" unit="mm" value="18.75" valueMax="19.2" valueMin="18.3">18.319.2 mm</quantity>
;
<tableCitation id="C68A9C819E7D3A3ACD0E44914A89FA38" box="[109,199,1431,1457]" captionStart="TABLE 1" captionStartId="6.[108,182,224,248]" captionTargetPageId="6" captionText="TABLE 1. Measurements (in mm) of Sturnira nana and Sturnira EC. Tabulated sample statistics include the observed range and the mean plus or minus one standard deviation. N = sample size." httpUri="http://table.plazi.org/id/DF77F9B29E713A36CD0F41E64998FE9F" pageId="10" pageNumber="11" tableUuid="DF77F9B29E713A36CD0F41E64998FE9F">tables 1</tableCitation>
,
<tableCitation id="C68A9C819E7D3A3ACDBB44914AA9FA38" box="[216,231,1431,1457]" captionStart="TABLE 6" captionStartId="15.[108,182,224,248]" captionTargetPageId="15" captionText="TABLE 6. Forearm and craniodental measurements (mm) analyzed for the type series of Sturnira boadai." httpUri="http://table.plazi.org/id/DF77F9B29E783A3FCD0F41E64ED0FF71" pageId="10" pageNumber="11" tableUuid="DF77F9B29E783A3FCD0F41E64ED0FF71">6</tableCitation>
) that is externally distinguished from other congeners by lacking shoulder glands (epaulettes), and by its sparsely haired hind feet, interfemoral membrane, and forearm. The braincase is globular, and the zygomatic arches are incomplete. The foramen magnum has an acutely angled anteroventral margin. The inner upper incisors protrude notably from the skull profile, and in rostral view, the distal third of their medial surfaces are in contact. Two to four lower incisors are present, of which the inner incisors are trilobed and subtriangular. The outer lower incisors (when present) are small, bilobed, with blunt edges, and inclined towards the inner incisors. In the series of specimens collected in Miazi Alto (N = 9), some intraspecific variation is observed in relation to the number and presence of the outer lower incisors: most specimens exhibit four lower incisors (e.g., QCAZ11122) and, in specimens where the outer lower incisors are absent (e.g., QCAZ11121), superficial alveoli or diastemata are observed.
<specimenCount id="9D0E62B39E7C3A3BCD0F45414B5BFBE8" box="[108,277,1095,1121]" count="1" pageId="11" pageNumber="12" type="generic">One specimen</specimenCount>
(QCAZ11119) has no outer lower incisors, alveoli, or spaces between i1 and the canine. Lastly,
<specimenCount id="9D0E62B39E7C3A3BCC78456B4B87FB0F" box="[283,457,1132,1159]" count="2" pageId="11" pageNumber="12" type="generic">two specimens</specimenCount>
(QCAZ11120 and QCAZ11123) exhibit only one outer incisor. In one of the latter cases (QCAZ 11123), the external incisor is minuscule and difficult to observe with the naked eye.
</paragraph>
</subSubSection>
<caption id="DF77F9B29E7C3A3BCD0F42AD4B9BFC56" ID-DOI="http://doi.org/10.5281/zenodo.10261222" ID-Zenodo-Dep="10261222" httpUri="https://zenodo.org/record/10261222/files/figure.png" pageId="11" pageNumber="12" startId="11.[108,195,939,960]" targetBox="[126,1198,229,912]" targetPageId="11" targetType="figure">
<paragraph id="8BB7A93A9E7C3A3BCD0F42AD4B9BFC56" blockId="11.[108,1213,937,991]" pageId="11" pageNumber="12">
FIGURE 4. Climatic PCA based on the climatic variables from the collection localities of
<taxonomicName id="4C08D2B99E7C3A3BC94842AC4EF3FC48" authorityName="Gardner and O'Neil" authorityYear="1971" box="[1067,1213,938,961]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E7C3A3BC94842AC4EF3FC48" box="[1067,1213,938,961]" italics="true" pageId="11" pageNumber="12">Sturnira nana</emphasis>
</taxonomicName>
(squares) and
<taxonomicName id="4C08D2B99E7C3A3BCD9D42C14B38FC56" authority="EC" authorityName="EC" box="[254,374,967,991]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B97C75289E7C3A3BCD9D42C14B1FFC57" box="[254,337,967,990]" italics="true" pageId="11" pageNumber="12">Sturnira</emphasis>
EC
</taxonomicName>
(circles).
</paragraph>
</caption>
<subSubSection id="C312FAB19E7C3A3ECDFF45DA4B2EFAEA" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
<paragraph id="8BB7A93A9E7C3A3BCDFF45DA4923F9CE" blockId="11.[108,1214,1020,1756]" pageId="11" pageNumber="12">
DESCRIPTION:
<taxonomicName id="4C08D2B99E7C3A3BCC2A45DA4BB0FB7F" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[329,510,1244,1270]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E7C3A3BCC2A45DA4BB0FB7F" box="[329,510,1244,1270]" italics="true" pageId="11" pageNumber="12">Sturnira boadai</emphasis>
</taxonomicName>
is one of the two smallest species of yellow-shouldered bats, with most measurements overlapping those of
<taxonomicName id="4C08D2B99E7C3A3BCFE8440448AFFA92" authorityName="Gardner and O'Neil" authorityYear="1971" box="[651,737,1282,1308]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E7C3A3BCFE8440448AFFA92" box="[651,737,1282,1308]" italics="true" pageId="11" pageNumber="12">S. nana</emphasis>
</taxonomicName>
. The dorsal fur is dense and dark brown, with long (
<quantity id="4CF004DF9E7C3A3BCD9244214B14FAC8" box="[241,346,1319,1345]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="11" pageNumber="12" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
) hairs. Dorsal hairs are tetracolored with a narrow white basal band of around 10% of the hair length, an epibasal brown band of about 40% of the hair length, a subterminal light-brown band of about 30% of the hair length, and a dark-brown apical band that covers 20% of the hair length. The ventral fur and underparts are lighter than the dorsal fur. Ventral hairs are tricolored due to lacking the terminal dark brown tip of the dorsal hairs. The fur is sparsely distributed at the dorsal surface of the femur, tibia, hind feet, interfemoral membrane, and upper forearm. The wing membranes are grayish to blackish brown. Shoulder glands are absent. The nose leaf is dark brown, long, and narrow.
</paragraph>
<paragraph id="8BB7A93A9E7C3A3BCDFF47574941F952" blockId="11.[108,1214,1020,1756]" pageId="11" pageNumber="12">
The skull of
<taxonomicName id="4C08D2B99E7C3A3BCC4F47544BACF9E2" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[300,482,1617,1644]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E7C3A3BCC4F47544BACF9E2" box="[300,482,1617,1644]" italics="true" pageId="11" pageNumber="12">Sturnira boadai</emphasis>
</taxonomicName>
has a globular braincase with a flattened rostrum by comparison with other members of the subgenus
<taxonomicName id="4C08D2B99E7C3A3BCF3A477148FFF918" authorityName="Thomas" authorityYear="1915" box="[601,689,1655,1681]" class="Mammalia" family="Phyllostomidae" genus="Corvira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B97C75289E7C3A3BCF3A477148FFF918" box="[601,689,1655,1681]" italics="true" pageId="11" pageNumber="12">Corvira</emphasis>
</taxonomicName>
. A sagittal crest is not developed. The zygomatic arches are always incomplete. The basisphenoid pits and septum are shallow. The anteroventral margin of the foramen magnum is angular.
</paragraph>
<caption id="DF77F9B29E7B3A3CCD0F44DD487AF9A5" ID-DOI="http://doi.org/10.5281/zenodo.10261227" ID-Zenodo-Dep="10261227" httpUri="https://zenodo.org/record/10261227/files/figure.png" pageId="12" pageNumber="13" startId="12.[108,194,1499,1520]" targetBox="[108,1212,224,1473]" targetPageId="12" targetType="figure">
<paragraph id="8BB7A93A9E7B3A3CCD0F44DD487AF9A5" blockId="12.[108,1213,1497,1580]" pageId="12" pageNumber="13">
FIGURE 5. Map of NW South America showing the collection localities of
<taxonomicName id="4C08D2B99E7B3A3CCE1144DC4E42FA78" box="[882,1036,1497,1521]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="bidens">
<emphasis id="B97C75289E7B3A3CCE1144DC4E42FA78" box="[882,1036,1497,1521]" italics="true" pageId="12" pageNumber="13">Sturnira bidens</emphasis>
</taxonomicName>
(circles),
<taxonomicName id="4C08D2B99E7B3A3CC91244DC4EF3FA78" authorityName="Gardner and O'Neil" authorityYear="1971" box="[1137,1213,1498,1521]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E7B3A3CC91244DC4EF3FA78" box="[1137,1213,1498,1521]" italics="true" pageId="12" pageNumber="13">S. nana</emphasis>
</taxonomicName>
(diamonds), and
<taxonomicName id="4C08D2B99E7B3A3CCC7A44F14BFBF986" authority="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho, 2023" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[281,437,1527,1551]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="boadai" status="sp. nov.">
<emphasis id="B97C75289E7B3A3CCC7A44F14BFBF986" box="[281,437,1527,1551]" italics="true" pageId="12" pageNumber="13">Sturnira boadai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24FC8539E7B3A3CCCA344F14842F987" box="[448,524,1527,1550]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
(stars), analyzed in this study. The collection points of
<taxonomicName id="4C08D2B99E7B3A3CC95244F14EF2F987" authorityName="Gardner and O'Neil" authorityYear="1971" box="[1073,1212,1527,1550]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E7B3A3CC95244F14EF2F987" box="[1073,1212,1527,1550]" italics="true" pageId="12" pageNumber="13">Sturnira nana</emphasis>
</taxonomicName>
overlap, so only two diamonds are observed.
</paragraph>
</caption>
<caption id="DF77F9B29E7A3A3DCD0F41E649B6FEBB" ID-Table-UUID="DF77F9B29E7A3A3DCD0F41E649B6FEBB" httpUri="http://table.plazi.org/id/DF77F9B29E7A3A3DCD0F41E649B6FEBB" pageId="13" pageNumber="14" startId="13.[108,182,224,248]" targetBox="[238,1063,341,1205]" targetIsTable="true" targetPageId="13" targetType="table">
<paragraph id="8BB7A93A9E7A3A3DCD0F41E649B6FEBB" blockId="13.[108,1205,224,307]" pageId="13" pageNumber="14">
TABLE 5. Component loadings of the principal components analysis based on a correlation matrix of the 19 bioclimatic variables collection localities of
<taxonomicName id="4C08D2B99E7A3A3DCF2941F8489BFE9C" authorityName="Gardner and O'Neil" authorityYear="1971" box="[586,725,254,277]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E7A3A3DCF2941F8489BFE9C" box="[586,725,254,277]" italics="true" pageId="13" pageNumber="14">Sturnira nana</emphasis>
</taxonomicName>
, and
<emphasis id="B97C75289E7A3A3DCE6C41F8496FFE9C" box="[783,801,254,277]" italics="true" pageId="13" pageNumber="14">S.</emphasis>
EC (N = 7). The table includes the first two principal components and the proportion of variance explained by each component.
</paragraph>
</caption>
<paragraph id="8BB7A93A9E7A3A3DCE3E40534E55FB3C" pageId="13" pageNumber="14">
<table id="F9085B9A9E7AC5CFCD8D40534E69FB3C" box="[238,1063,341,1205]" gridcols="3" gridrows="22" pageId="13" pageNumber="14">
<tr id="3538AB789E7AC5CFCD8D40534E69FEE2" box="[238,1063,341,363]" gridrow="0" pageId="13" pageNumber="14" rowspan-0="1">
<th id="76E9C2049E7AC5CFCE4B40534E69FEE2" box="[808,1063,341,363]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="13" pageNumber="14">
<emphasis id="B97C75289E7A3A3DCE3E40534999FEE2" bold="true" box="[861,983,341,363]" pageId="13" pageNumber="14">Components</emphasis>
</th>
</tr>
<tr id="3538AB789E7AC5CFCD8D40784E69FE1A" box="[238,1063,382,403]" gridrow="1" pageId="13" pageNumber="14" rowspan-0="1">
<td id="76E9C2049E7AC5CFCE4B40784939FE1A" box="[808,887,382,403]" gridcol="1" gridrow="1" pageId="13" pageNumber="14">PC 1</td>
<td id="76E9C2049E7AC5CFCEAE40784E69FE1A" box="[973,1063,382,403]" gridcol="2" gridrow="1" pageId="13" pageNumber="14">PC 2</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D40A34E69FE32" box="[238,1063,421,443]" gridrow="2" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D40A348F4FE32" box="[238,698,421,443]" gridcol="0" gridrow="2" pageId="13" pageNumber="14">Annual mean temperature -BIO1</th>
<td id="76E9C2049E7AC5CFCE4B40A34939FE32" box="[808,887,421,443]" gridcol="1" gridrow="2" pageId="13" pageNumber="14">0.94767</td>
<td id="76E9C2049E7AC5CFCEAE40A34E69FE32" box="[973,1063,421,443]" gridcol="2" gridrow="2" pageId="13" pageNumber="14">0.24599</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D40CB4E69FE6A" box="[238,1063,461,483]" gridrow="3" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D40CB48F4FE6A" box="[238,698,461,483]" gridcol="0" gridrow="3" pageId="13" pageNumber="14">Mean diurnal range - BIO2</th>
<td id="76E9C2049E7AC5CFCE4B40CB4939FE6A" box="[808,887,461,483]" gridcol="1" gridrow="3" pageId="13" pageNumber="14">0.93301</td>
<td id="76E9C2049E7AC5CFCEAE40CB4E69FE6A" box="[973,1063,461,483]" gridcol="2" gridrow="3" pageId="13" pageNumber="14">0.28858</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D40F34E69FD82" box="[238,1063,501,523]" gridrow="4" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D40F348F4FD82" box="[238,698,501,523]" gridcol="0" gridrow="4" pageId="13" pageNumber="14">Isothermality - BIO3</th>
<td id="76E9C2049E7AC5CFCE4B40F34939FD82" box="[808,887,501,523]" gridcol="1" gridrow="4" pageId="13" pageNumber="14">0.93906</td>
<td id="76E9C2049E7AC5CFCEAE40F34E69FD82" box="[973,1063,501,523]" gridcol="2" gridrow="4" pageId="13" pageNumber="14">0.24825</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D431B4E69FDBA" box="[238,1063,541,563]" gridrow="5" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D431B48F4FDBA" box="[238,698,541,563]" gridcol="0" gridrow="5" pageId="13" pageNumber="14">Temperature seasonality - BIO4</th>
<td id="76E9C2049E7AC5CFCE4B431B4939FDBA" box="[808,887,541,563]" gridcol="1" gridrow="5" pageId="13" pageNumber="14">-0.87055</td>
<td id="76E9C2049E7AC5CFCEAE431B4E69FDBA" box="[973,1063,541,563]" gridcol="2" gridrow="5" pageId="13" pageNumber="14">0.45631</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D43434E69FDD2" box="[238,1063,581,603]" gridrow="6" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D434348F4FDD2" box="[238,698,581,603]" gridcol="0" gridrow="6" pageId="13" pageNumber="14">Maximum temperature of warmest month BIO5</th>
<td id="76E9C2049E7AC5CFCE4B43434939FDD2" box="[808,887,581,603]" gridcol="1" gridrow="6" pageId="13" pageNumber="14">-0.20329</td>
<td id="76E9C2049E7AC5CFCEAE43434E69FDD2" box="[973,1063,581,603]" gridcol="2" gridrow="6" pageId="13" pageNumber="14">0.78056</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D436B4E69FD0A" box="[238,1063,621,643]" gridrow="7" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D436B48F4FD0A" box="[238,698,621,643]" gridcol="0" gridrow="7" pageId="13" pageNumber="14">Minimum temperature of coldest month BIO6</th>
<td id="76E9C2049E7AC5CFCE4B436B4939FD0A" box="[808,887,621,643]" gridcol="1" gridrow="7" pageId="13" pageNumber="14">-0.96516</td>
<td id="76E9C2049E7AC5CFCEAE436B4E69FD0A" box="[973,1063,621,643]" gridcol="2" gridrow="7" pageId="13" pageNumber="14">0.17681</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D43934E69FD22" box="[238,1063,661,683]" gridrow="8" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D439348F4FD22" box="[238,698,661,683]" gridcol="0" gridrow="8" pageId="13" pageNumber="14">Temperature annual range BIO7</th>
<td id="76E9C2049E7AC5CFCE4B43934939FD22" box="[808,887,661,683]" gridcol="1" gridrow="8" pageId="13" pageNumber="14">0.94094</td>
<td id="76E9C2049E7AC5CFCEAE43934E69FD22" box="[973,1063,661,683]" gridcol="2" gridrow="8" pageId="13" pageNumber="14">-0.11996</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D43BB4E69FD5A" box="[238,1063,701,723]" gridrow="9" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D43BB48F4FD5A" box="[238,698,701,723]" gridcol="0" gridrow="9" pageId="13" pageNumber="14">Mean temperature of wettest quarter BIO8</th>
<td id="76E9C2049E7AC5CFCE4B43BB4939FD5A" box="[808,887,701,723]" gridcol="1" gridrow="9" pageId="13" pageNumber="14">-0.13235</td>
<td id="76E9C2049E7AC5CFCEAE43BB4E69FD5A" box="[973,1063,701,723]" gridcol="2" gridrow="9" pageId="13" pageNumber="14">0.78517</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D43E34E69FD72" box="[238,1063,741,763]" gridrow="10" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D43E348F4FD72" box="[238,698,741,763]" gridcol="0" gridrow="10" pageId="13" pageNumber="14">Mean temperature of driest quarter BIO9</th>
<td id="76E9C2049E7AC5CFCE4B43E34939FD72" box="[808,887,741,763]" gridcol="1" gridrow="10" pageId="13" pageNumber="14">-0.96462</td>
<td id="76E9C2049E7AC5CFCEAE43E34E69FD72" box="[973,1063,741,763]" gridcol="2" gridrow="10" pageId="13" pageNumber="14">0.19795</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D420B4E69FCAA" box="[238,1063,781,803]" gridrow="11" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D420B48F4FCAA" box="[238,698,781,803]" gridcol="0" gridrow="11" pageId="13" pageNumber="14">Mean temperature of warmest quarter BIO10</th>
<td id="76E9C2049E7AC5CFCE4B420B4939FCAA" box="[808,887,781,803]" gridcol="1" gridrow="11" pageId="13" pageNumber="14">-0.21603</td>
<td id="76E9C2049E7AC5CFCEAE420B4E69FCAA" box="[973,1063,781,803]" gridcol="2" gridrow="11" pageId="13" pageNumber="14">0.80937</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D42334E69FCC2" box="[238,1063,821,843]" gridrow="12" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D423348F4FCC2" box="[238,698,821,843]" gridcol="0" gridrow="12" pageId="13" pageNumber="14">Mean temperature of coldest quarter BIO11</th>
<td id="76E9C2049E7AC5CFCE4B42334939FCC2" box="[808,887,821,843]" gridcol="1" gridrow="12" pageId="13" pageNumber="14">-0.965</td>
<td id="76E9C2049E7AC5CFCEAE42334E69FCC2" box="[973,1063,821,843]" gridcol="2" gridrow="12" pageId="13" pageNumber="14">0.23675</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D425B4E69FCFA" box="[238,1063,861,883]" gridrow="13" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D425B48F4FCFA" box="[238,698,861,883]" gridcol="0" gridrow="13" pageId="13" pageNumber="14">Annual precipitation BIO12</th>
<td id="76E9C2049E7AC5CFCE4B425B4939FCFA" box="[808,887,861,883]" gridcol="1" gridrow="13" pageId="13" pageNumber="14">0.97492</td>
<td id="76E9C2049E7AC5CFCEAE425B4E69FCFA" box="[973,1063,861,883]" gridcol="2" gridrow="13" pageId="13" pageNumber="14">-0.1052</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D42834E69FC12" box="[238,1063,901,923]" gridrow="14" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D428348F4FC12" box="[238,698,901,923]" gridcol="0" gridrow="14" pageId="13" pageNumber="14">Precipitation of wettest month BIO13</th>
<td id="76E9C2049E7AC5CFCE4B42834939FC12" box="[808,887,901,923]" gridcol="1" gridrow="14" pageId="13" pageNumber="14">-0.93227</td>
<td id="76E9C2049E7AC5CFCEAE42834E69FC12" box="[973,1063,901,923]" gridcol="2" gridrow="14" pageId="13" pageNumber="14">-0.25005</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D42AB4E69FC4A" box="[238,1063,941,963]" gridrow="15" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D42AB48F4FC4A" box="[238,698,941,963]" gridcol="0" gridrow="15" pageId="13" pageNumber="14">Precipitation of driest month BIO14</th>
<td id="76E9C2049E7AC5CFCE4B42AB4939FC4A" box="[808,887,941,963]" gridcol="1" gridrow="15" pageId="13" pageNumber="14">0.97346</td>
<td id="76E9C2049E7AC5CFCEAE42AB4E69FC4A" box="[973,1063,941,963]" gridcol="2" gridrow="15" pageId="13" pageNumber="14">0.19309</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D42D34E69FC62" box="[238,1063,981,1003]" gridrow="16" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D42D348F4FC62" box="[238,698,981,1003]" gridcol="0" gridrow="16" pageId="13" pageNumber="14">Precipitation seasonality BIO15</th>
<td id="76E9C2049E7AC5CFCE4B42D34939FC62" box="[808,887,981,1003]" gridcol="1" gridrow="16" pageId="13" pageNumber="14">0.97448</td>
<td id="76E9C2049E7AC5CFCEAE42D34E69FC62" box="[973,1063,981,1003]" gridcol="2" gridrow="16" pageId="13" pageNumber="14">0.18217</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D42FB4E69FB9A" box="[238,1063,1021,1043]" gridrow="17" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D42FB48F4FB9A" box="[238,698,1021,1043]" gridcol="0" gridrow="17" pageId="13" pageNumber="14">Precipitation of wettest quarter BIO16</th>
<td id="76E9C2049E7AC5CFCE4B42FB4939FB9A" box="[808,887,1021,1043]" gridcol="1" gridrow="17" pageId="13" pageNumber="14">-0.4039</td>
<td id="76E9C2049E7AC5CFCEAE42FB4E69FB9A" box="[973,1063,1021,1043]" gridcol="2" gridrow="17" pageId="13" pageNumber="14">0.54498</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D45234E69FBB2" box="[238,1063,1061,1083]" gridrow="18" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D452348F4FBB2" box="[238,698,1061,1083]" gridcol="0" gridrow="18" pageId="13" pageNumber="14">Precipitation of driest quarter BIO17</th>
<td id="76E9C2049E7AC5CFCE4B45234939FBB2" box="[808,887,1061,1083]" gridcol="1" gridrow="18" pageId="13" pageNumber="14">0.97769</td>
<td id="76E9C2049E7AC5CFCEAE45234E69FBB2" box="[973,1063,1061,1083]" gridcol="2" gridrow="18" pageId="13" pageNumber="14">-0.049589</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D454B4E69FBEA" box="[238,1063,1101,1123]" gridrow="19" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D454B48F4FBEA" box="[238,698,1101,1123]" gridcol="0" gridrow="19" pageId="13" pageNumber="14">Precipitation of warmest quarter BIO18</th>
<td id="76E9C2049E7AC5CFCE4B454B4939FBEA" box="[808,887,1101,1123]" gridcol="1" gridrow="19" pageId="13" pageNumber="14">0.94914</td>
<td id="76E9C2049E7AC5CFCEAE454B4E69FBEA" box="[973,1063,1101,1123]" gridcol="2" gridrow="19" pageId="13" pageNumber="14">0.1972</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D45734E69FB02" box="[238,1063,1141,1163]" gridrow="20" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D457348F4FB02" box="[238,698,1141,1163]" gridcol="0" gridrow="20" pageId="13" pageNumber="14">Precipitation of coldest quarter BIO19</th>
<td id="76E9C2049E7AC5CFCE4B45734939FB02" box="[808,887,1141,1163]" gridcol="1" gridrow="20" pageId="13" pageNumber="14">0.6817</td>
<td id="76E9C2049E7AC5CFCEAE45734E69FB02" box="[973,1063,1141,1163]" gridcol="2" gridrow="20" pageId="13" pageNumber="14">0.56847</td>
</tr>
<tr id="3538AB789E7AC5CFCD8D45994E69FB3C" box="[238,1063,1183,1205]" gridrow="21" pageId="13" pageNumber="14">
<th id="76E9C2049E7AC5CFCD8D459948F4FB3C" box="[238,698,1183,1205]" gridcol="0" gridrow="21" pageId="13" pageNumber="14">Proportion of variance</th>
<td id="76E9C2049E7AC5CFCE4B45994939FB3C" box="[808,887,1183,1205]" gridcol="1" gridrow="21" pageId="13" pageNumber="14">70.49%</td>
<td id="76E9C2049E7AC5CFCEAE45994E69FB3C" box="[973,1063,1183,1205]" gridcol="2" gridrow="21" pageId="13" pageNumber="14">17.06%</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB7A93A9E7A3A3DCDFF44224B86F950" blockId="13.[108,1213,1316,1753]" pageId="13" pageNumber="14">
The dental formula is I2/12, C1/1, P2/2, M3/3 x 2 = 3032. The inner upper incisor (I1) is proodont, with a straight occlusal edge and a well-developed posterolateral cusp. The outer upper incisors (I2) are small and opisthodont. I2 is close to but not in contact with the posterolateral cusp of the I1 (
<figureCitation id="1333B5BF9E7A3A3DCCC84492484DFA27" box="[427,515,1428,1454]" captionStart="FIGURE 6" captionStartId="14.[109,195,773,794]" captionTargetBox="[216,1104,224,747]" captionTargetId="figure-401@14.[216,1104,224,747]" captionTargetPageId="14" captionText="FIGURE 6. A. Upper and lower incisors of the holotype of Sturnira boadai, sp. nov. (QCAZ 11122). B. Detail of incisors. Photographs by Rubén D. Jarrín." figureDoi="http://doi.org/10.5281/zenodo.10261230" httpUri="https://zenodo.org/record/10261230/files/figure.png" pageId="13" pageNumber="14">fig. 6A</figureCitation>
). I1 is more than three times the height of I2. The upper canine (C1) is long and robust. The first upper premolar (P3) is small, narrow, and half the height of the second upper premolar (P4). P4 is broad, with a blunt distal cusp that is more noticeable in some specimens than in others, but it is always present. Diastemata are present between P3 and C1, and between P3 and P4. M1 and M2 are broad. The anteroposterior length of the first upper molar (M1) is greater than that of M2 (
<figureCitation id="1333B5BF9E7A3A3DCE00474949FAF9E0" box="[867,948,1615,1641]" captionStart="FIGURE 1" captionStartId="5.[109,195,1281,1302]" captionTargetBox="[108,1212,224,1255]" captionTargetId="figure-149@5.[108,1212,224,1255]" captionTargetPageId="5" captionText="FIGURE 1. A, B. Lateral, C, D. dorsal, and E, F. ventral views of the skulls and G, H. lateral view of the mandibles of Sturnira nana (AMNH 219172, left column) and Sturnira EC (QCAZ 11122, right column). Red arrows show the differences between the two skulls." figureDoi="http://doi.org/10.5281/zenodo.10261214" httpUri="https://zenodo.org/record/10261214/files/figure.png" pageId="13" pageNumber="14">fig. 1B</figureCitation>
). In occlusal view, the paracone of M1 is shorter than the metacone. The second upper molar (M2) is ovoid and has a broad crown. The third upper molar (M3) is small, with a crown area approximately one-half that of M2 (
<figureCitation id="1333B5BF9E7A3A3DCC0947B94BF4F950" box="[362,442,1727,1753]" captionStart="FIGURE 1" captionStartId="5.[109,195,1281,1302]" captionTargetBox="[108,1212,224,1255]" captionTargetId="figure-149@5.[108,1212,224,1255]" captionTargetPageId="5" captionText="FIGURE 1. A, B. Lateral, C, D. dorsal, and E, F. ventral views of the skulls and G, H. lateral view of the mandibles of Sturnira nana (AMNH 219172, left column) and Sturnira EC (QCAZ 11122, right column). Red arrows show the differences between the two skulls." figureDoi="http://doi.org/10.5281/zenodo.10261214" httpUri="https://zenodo.org/record/10261214/files/figure.png" pageId="13" pageNumber="14">fig. 1F</figureCitation>
).
</paragraph>
<caption id="DF77F9B29E793A3ECD0E4203487CFCB0" ID-DOI="http://doi.org/10.5281/zenodo.10261230" ID-Zenodo-Dep="10261230" httpUri="https://zenodo.org/record/10261230/files/figure.png" pageId="14" pageNumber="15" startId="14.[109,195,773,794]" targetBox="[216,1104,224,747]" targetPageId="14" targetType="figure">
<paragraph id="8BB7A93A9E793A3ECD0E4203487CFCB0" blockId="14.[109,1212,771,825]" pageId="14" pageNumber="15">
FIGURE 6.
<emphasis id="B97C75289E793A3ECD8242054AB4FC92" bold="true" box="[225,250,771,795]" pageId="14" pageNumber="15">A.</emphasis>
Upper and lower incisors of the holotype of
<taxonomicName id="4C08D2B99E793A3ECFA242024910FC92" authority="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho, 2023" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[705,862,771,795]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai" status="sp. nov.">
<emphasis id="B97C75289E793A3ECFA242024910FC92" box="[705,862,771,795]" italics="true" pageId="14" pageNumber="15">Sturnira boadai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24FC8539E793A3ECE09420249F9FC92" box="[874,951,772,795]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(QCAZ 11122).
<emphasis id="B97C75289E793A3EC90142024E37FC95" bold="true" box="[1122,1145,772,796]" pageId="14" pageNumber="15">B.</emphasis>
Detail of incisors. Photographs by Rubén D. Jarrín.
</paragraph>
</caption>
<paragraph id="8BB7A93A9E793A3ECDFF42624B2EFAEA" blockId="14.[108,1215,867,1753]" pageId="14" pageNumber="15">
Two to four lower incisors are present. The inner lower incisor (i1) is trilobed. When present, the outer lower incisor (i2s) is minute, bilobed, and approximately one-third the size of i1. The anterior surface of the lower canine(c1) is in contact with the entire posterior surface of the i2 (
<figureCitation id="1333B5BF9E793A3ECDA342D54B5EFC64" box="[192,272,979,1005]" captionStart="FIGURE 6" captionStartId="14.[109,195,773,794]" captionTargetBox="[216,1104,224,747]" captionTargetId="figure-401@14.[216,1104,224,747]" captionTargetPageId="14" captionText="FIGURE 6. A. Upper and lower incisors of the holotype of Sturnira boadai, sp. nov. (QCAZ 11122). B. Detail of incisors. Photographs by Rubén D. Jarrín." figureDoi="http://doi.org/10.5281/zenodo.10261230" httpUri="https://zenodo.org/record/10261230/files/figure.png" pageId="14" pageNumber="15">fig. 6B</figureCitation>
), but when i2 is missing, the c1 is in contact with i1. The lower canines are long, narrow, robust, and laterally divergent, with their shafts slanted outward. The anteroposterior length of the first lower premolar (p2) is greater than that of the second lower premolar (p4). In lateral view, p2 exhibits an irregular, tricuspid border that is wider (anteroposterior dimension) than the crown of the tooth is tall. By contrast, p4 is taller than it is wide, and it has a well-developed main cusp. Both lower premolars are separated by a diastema (
<figureCitation id="1333B5BF9E793A3EC94045884E34FB21" box="[1059,1146,1166,1192]" captionStart="FIGURE 1" captionStartId="5.[109,195,1281,1302]" captionTargetBox="[108,1212,224,1255]" captionTargetId="figure-149@5.[108,1212,224,1255]" captionTargetPageId="5" captionText="FIGURE 1. A, B. Lateral, C, D. dorsal, and E, F. ventral views of the skulls and G, H. lateral view of the mandibles of Sturnira nana (AMNH 219172, left column) and Sturnira EC (QCAZ 11122, right column). Red arrows show the differences between the two skulls." figureDoi="http://doi.org/10.5281/zenodo.10261214" httpUri="https://zenodo.org/record/10261214/files/figure.png" pageId="14" pageNumber="15">fig. 1H</figureCitation>
). The first lower molar (m1) is broad. The anteroposterior length of m1 is larger than that of the second lower molar (m2). The third lower molar (m3) is small, with two well-defined lobes separated by a notch between the metaconid and entoconid. The metaconid and entoconid of m1 and m2 are moderately defined. Adjacent upper and lower teeth are separated by narrow diastemata (
<figureCitation id="1333B5BF9E793A3ECD9F444F4B1FFAEA" box="[252,337,1353,1379]" captionStart="FIGURE 1" captionStartId="5.[109,195,1281,1302]" captionTargetBox="[108,1212,224,1255]" captionTargetId="figure-149@5.[108,1212,224,1255]" captionTargetPageId="5" captionText="FIGURE 1. A, B. Lateral, C, D. dorsal, and E, F. ventral views of the skulls and G, H. lateral view of the mandibles of Sturnira nana (AMNH 219172, left column) and Sturnira EC (QCAZ 11122, right column). Red arrows show the differences between the two skulls." figureDoi="http://doi.org/10.5281/zenodo.10261214" httpUri="https://zenodo.org/record/10261214/files/figure.png" pageId="14" pageNumber="15">fig. 1H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C312FAB19E793A3FCDFF44684ABFFB45" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BB7A93A9E793A3ECDFF44684EFFF9E0" blockId="14.[108,1215,867,1753]" pageId="14" pageNumber="15">
COMPARISONS: Among other species traditionally referred to the subgenus
<taxonomicName id="4C08D2B99E793A3EC97444694E3AFA00" box="[1047,1140,1391,1417]" class="Actinopterygii" family="Sciaenidae" genus="Corvina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="B97C75289E793A3EC97444694E3AFA00" box="[1047,1140,1391,1417]" italics="true" pageId="14" pageNumber="15">Corvina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C08D2B99E793A3EC9E044694ABFFA24" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E793A3EC9E044694ABFFA24" italics="true" pageId="14" pageNumber="15">Sturnira boadai</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C08D2B99E793A3ECCE944924BA1FA24" box="[394,495,1427,1454]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="bidens">
<emphasis id="B97C75289E793A3ECCE944924BA1FA24" box="[394,495,1427,1454]" italics="true" pageId="14" pageNumber="15">S. bidens</emphasis>
</taxonomicName>
in size, being notably smaller. Morphologically,
<taxonomicName id="4C08D2B99E793A3EC94744924EC0FA24" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[1060,1166,1427,1454]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E793A3EC94744924EC0FA24" box="[1060,1166,1427,1454]" italics="true" pageId="14" pageNumber="15">S. boadai</emphasis>
</taxonomicName>
has sparsely haired hind feet and uropatagium, whereas both structures are densely haired in
<taxonomicName id="4C08D2B99E793A3EC9C444BF4AF8FA71" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="bidens">
<emphasis id="B97C75289E793A3EC9C444BF4AF8FA71" italics="true" pageId="14" pageNumber="15">S. bidens</emphasis>
</taxonomicName>
. Additionally,
<taxonomicName id="4C08D2B99E793A3ECC0544D94B9DFA71" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[358,467,1502,1529]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E793A3ECC0544D94B9DFA71" box="[358,467,1502,1529]" italics="true" pageId="14" pageNumber="15">S. boadai</emphasis>
</taxonomicName>
possesses one or two pairs of lower incisors, whereas
<taxonomicName id="4C08D2B99E793A3EC93444D94EF0FA71" box="[1111,1214,1502,1529]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="bidens">
<emphasis id="B97C75289E793A3EC93444D94EF0FA71" box="[1111,1214,1502,1529]" italics="true" pageId="14" pageNumber="15">S. bidens</emphasis>
</taxonomicName>
always has a single pair of lower incisors. The anterior margin of the foramen magnum in
<taxonomicName id="4C08D2B99E793A3EC9C947024AF5F9CA" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E793A3EC9C947024AF5F9CA" italics="true" pageId="14" pageNumber="15">S. boadai</emphasis>
</taxonomicName>
is angular, whereas it is rounded in most
<taxonomicName id="4C08D2B99E793A3ECFA2472F4964F9CA" box="[705,810,1577,1603]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="bidens">
<emphasis id="B97C75289E793A3ECFA2472F4964F9CA" box="[705,810,1577,1603]" italics="true" pageId="14" pageNumber="15">S. bidens</emphasis>
</taxonomicName>
specimens. Lastly, the zygomatic arches are incomplete in
<taxonomicName id="4C08D2B99E793A3ECCF647494BB1F9E1" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[405,511,1614,1641]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E793A3ECCF647494BB1F9E1" box="[405,511,1614,1641]" italics="true" pageId="14" pageNumber="15">S. boadai</emphasis>
</taxonomicName>
, whereas they can be complete or incomplete in
<taxonomicName id="4C08D2B99E793A3EC92047484EE4F9E1" box="[1091,1194,1614,1640]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="17" phylum="Chordata" rank="species" species="bidens">
S.
<emphasis id="B97C75289E793A3EC90247484EE4F9E1" box="[1121,1194,1614,1640]" italics="true" pageId="14" pageNumber="15">bidens</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB7A93A9E793A3FCDFF47754ABFFB45" blockId="14.[108,1215,867,1753]" lastBlockId="15.[108,1213,903,1452]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Externally,
<taxonomicName id="4C08D2B99E793A3ECC4E47724BBCF904" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[301,498,1651,1678]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E793A3ECC4E47724BBCF904" box="[301,498,1651,1678]" italics="true" pageId="14" pageNumber="15">Sturnira boadai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C08D2B99E793A3ECF5B477248D9F904" authorityName="Gardner and O'Neil" authorityYear="1971" box="[568,663,1652,1678]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E793A3ECF5B477248D9F904" box="[568,663,1652,1678]" italics="true" pageId="14" pageNumber="15">S. nana</emphasis>
</taxonomicName>
are similar. Epaulettes (patches of stained shoulder hairs) are not evident in either species. The dorsal fur is long and tetracolored in both species, and the forearm, legs, feet, proximal segments of the wings, and uropatagium are sparsely covered with long hairs. However, both species can be distinguished by several craniodental characteristics. The skull has a globular braincase and a less elongated rostrum in
<taxonomicName id="4C08D2B99E783A3FCDB642D44B0BFC65" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[213,325,978,1004]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E783A3FCDB642D44B0BFC65" box="[213,325,978,1004]" italics="true" pageId="15" pageNumber="16">S. boadai</emphasis>
</taxonomicName>
, whereas the braincase is relatively long with a narrow, sloping rostrum in
<taxonomicName id="4C08D2B99E783A3FCDED42FE4AA7FB98" authorityName="Gardner and O'Neil" authorityYear="1971" box="[142,233,1016,1042]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E783A3FCDED42FE4AA7FB98" box="[142,233,1016,1042]" italics="true" pageId="15" pageNumber="16">S. nana</emphasis>
</taxonomicName>
. The anterior margin of the foramen magnum is angular in
<taxonomicName id="4C08D2B99E783A3FCEB842FE4E07FB98" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[987,1097,1015,1042]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E783A3FCEB842FE4E07FB98" box="[987,1097,1015,1042]" italics="true" pageId="15" pageNumber="16">S. boadai</emphasis>
</taxonomicName>
, whereas it is rounded
<taxonomicName id="4C08D2B99E783A3FCC75451B4B3CFBBF" authorityName="Gardner and O'Neil" authorityYear="1971" box="[278,370,1053,1079]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E783A3FCC75451B4B3CFBBF" box="[278,370,1053,1079]" italics="true" pageId="15" pageNumber="16">S. nana</emphasis>
</taxonomicName>
. The zygomatic arches are always incomplete in
<taxonomicName id="4C08D2B99E783A3FCEB4451B4E07FBBF" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[983,1097,1052,1079]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E783A3FCEB4451B4E07FBBF" box="[983,1097,1052,1079]" italics="true" pageId="15" pageNumber="16">S. boadai</emphasis>
</taxonomicName>
, whereas they can be complete or incomplete in
<taxonomicName id="4C08D2B99E783A3FCF3B454448FBFBD5" authorityName="Gardner and O'Neil" authorityYear="1971" box="[600,693,1090,1116]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E783A3FCF3B454448FBFBD5" box="[600,693,1090,1116]" italics="true" pageId="15" pageNumber="16">S. nana</emphasis>
</taxonomicName>
. P3 is small, narrow, sharp crowned, and not in contact with either C1 or P
<quantity id="4CF004DF9E783A3FCF6945614877FB08" box="[522,569,1127,1153]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="16" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C08D2B99E783A3FCF20456E48E4FB08" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[579,682,1127,1154]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E783A3FCF20456E48E4FB08" box="[579,682,1127,1154]" italics="true" pageId="15" pageNumber="16">S boadai</emphasis>
</taxonomicName>
, whereas P3 is broader and in contact with both C1 and P
<quantity id="4CF004DF9E783A3FCC41458B4B1CFB2F" box="[290,338,1164,1191]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="16" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C08D2B99E783A3FCC3E458B4BF8FB2F" authorityName="Gardner and O'Neil" authorityYear="1971" box="[349,438,1165,1191]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E783A3FCC3E458B4BF8FB2F" box="[349,438,1165,1191]" italics="true" pageId="15" pageNumber="16">S. nana</emphasis>
</taxonomicName>
. Lastly, M3 is less wide in lateral view, whereas M3 is broader in
<taxonomicName id="4C08D2B99E783A3FCDED45B44AA7FB45" authorityName="Gardner and O'Neil" authorityYear="1971" box="[142,233,1202,1228]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nana">
<emphasis id="B97C75289E783A3FCDED45B44AA7FB45" box="[142,233,1202,1228]" italics="true" pageId="15" pageNumber="16">S. nana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF77F9B29E783A3FCD0F41E64ED0FF71" ID-Table-UUID="DF77F9B29E783A3FCD0F41E64ED0FF71" box="[108,1182,224,249]" httpUri="http://table.plazi.org/id/DF77F9B29E783A3FCD0F41E64ED0FF71" pageId="15" pageNumber="16" startId="15.[108,182,224,248]" targetBox="[119,1195,281,824]" targetIsTable="true" targetPageId="15" targetType="table">
<paragraph id="8BB7A93A9E783A3FCD0F41E64ED0FF71" blockId="15.[108,1182,224,249]" box="[108,1182,224,249]" pageId="15" pageNumber="16">
TABLE 6. Forearm and craniodental measurements (mm) analyzed for the type series of
<taxonomicName id="4C08D2B99E783A3FCE9F41E74ED7FF71" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[1020,1177,224,248]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E783A3FCE9F41E74ED7FF71" box="[1020,1177,224,248]" italics="true" pageId="15" pageNumber="16">Sturnira boadai</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BB7A93A9E783A3FCD14401C4EC7FCBE" pageId="15" pageNumber="16">
<table id="F9085B9A9E78C5CFCD14401F4EE5FCB1" box="[119,1195,281,824]" gridcols="7" gridrows="14" pageId="15" pageNumber="16">
<tr id="3538AB789E78C5CFCD14401F4EE5FEB9" box="[119,1195,281,304]" gridrow="0" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD14401F4AB2FEB9" box="[119,252,281,304]" gridcol="0" gridrow="0" pageId="15" pageNumber="16">Measurements</th>
<th id="76E9C2049E78C5CFCC7F401F4BD9FEB9" box="[284,407,281,304]" gridcol="1" gridrow="0" pageId="15" pageNumber="16">QCAZ11122a</th>
<th id="76E9C2049E78C5CFCCDD401F487CFEB9" box="[446,562,281,304]" gridcol="2" gridrow="0" pageId="15" pageNumber="16">QCAZ11120</th>
<th id="76E9C2049E78C5CFCF3D401F489FFEB9" box="[606,721,281,304]" gridcol="3" gridrow="0" pageId="15" pageNumber="16">QCAZ11116</th>
<th id="76E9C2049E78C5CFCF9E401F493FFEB9" box="[765,881,281,304]" gridcol="4" gridrow="0" pageId="15" pageNumber="16">QCAZ11119</th>
<th id="76E9C2049E78C5CFCEFF401F4E5EFEB9" box="[924,1040,281,304]" gridcol="5" gridrow="0" pageId="15" pageNumber="16">QCAZ11121</th>
<th id="76E9C2049E78C5CFC954401F4EE5FEB9" box="[1079,1195,281,304]" gridcol="6" gridrow="0" pageId="15" pageNumber="16">QCAZ11123</th>
</tr>
<tr id="3538AB789E78C5CFCD1440474EE5FED1" box="[119,1195,321,344]" gridrow="1" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1440474AB2FED1" box="[119,252,321,344]" gridcol="0" gridrow="1" pageId="15" pageNumber="16">FA</th>
<td id="76E9C2049E78C5CFCC7F40474BD9FED1" box="[284,407,321,344]" gridcol="1" gridrow="1" pageId="15" pageNumber="16">33.67</td>
<td id="76E9C2049E78C5CFCCDD4047487CFED1" box="[446,562,321,344]" gridcol="2" gridrow="1" pageId="15" pageNumber="16">33.79</td>
<td id="76E9C2049E78C5CFCF3D4047489FFED1" box="[606,721,321,344]" gridcol="3" gridrow="1" pageId="15" pageNumber="16">33.36</td>
<td id="76E9C2049E78C5CFCF9E4047493FFED1" box="[765,881,321,344]" gridcol="4" gridrow="1" pageId="15" pageNumber="16">33.80</td>
<td id="76E9C2049E78C5CFCEFF40474E5EFED1" box="[924,1040,321,344]" gridcol="5" gridrow="1" pageId="15" pageNumber="16">32.52</td>
<td id="76E9C2049E78C5CFC95440474EE5FED1" box="[1079,1195,321,344]" gridcol="6" gridrow="1" pageId="15" pageNumber="16">33.35</td>
</tr>
<tr id="3538AB789E78C5CFCD14406F4EE5FE09" box="[119,1195,361,384]" gridrow="2" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD14406F4AB2FE09" box="[119,252,361,384]" gridcol="0" gridrow="2" pageId="15" pageNumber="16">CC</th>
<td id="76E9C2049E78C5CFCC7F406F4BD9FE09" box="[284,407,361,384]" gridcol="1" gridrow="2" pageId="15" pageNumber="16">4.64</td>
<td id="76E9C2049E78C5CFCCDD406F487CFE09" box="[446,562,361,384]" gridcol="2" gridrow="2" pageId="15" pageNumber="16">4.47</td>
<td id="76E9C2049E78C5CFCF3D406F489FFE09" box="[606,721,361,384]" gridcol="3" gridrow="2" pageId="15" pageNumber="16">4.46</td>
<td id="76E9C2049E78C5CFCF9E406F493FFE09" box="[765,881,361,384]" gridcol="4" gridrow="2" pageId="15" pageNumber="16">4.45</td>
<td id="76E9C2049E78C5CFCEFF406F4E5EFE09" box="[924,1040,361,384]" gridcol="5" gridrow="2" pageId="15" pageNumber="16">4.76</td>
<td id="76E9C2049E78C5CFC954406F4EE5FE09" box="[1079,1195,361,384]" gridcol="6" gridrow="2" pageId="15" pageNumber="16">4.43</td>
</tr>
<tr id="3538AB789E78C5CFCD1440974EE5FE21" box="[119,1195,401,424]" gridrow="3" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1440974AB2FE21" box="[119,252,401,424]" gridcol="0" gridrow="3" pageId="15" pageNumber="16">M2-M2</th>
<td id="76E9C2049E78C5CFCC7F40974BD9FE21" box="[284,407,401,424]" gridcol="1" gridrow="3" pageId="15" pageNumber="16">6.03</td>
<td id="76E9C2049E78C5CFCCDD4097487CFE21" box="[446,562,401,424]" gridcol="2" gridrow="3" pageId="15" pageNumber="16">6.09</td>
<td id="76E9C2049E78C5CFCF3D4097489FFE21" box="[606,721,401,424]" gridcol="3" gridrow="3" pageId="15" pageNumber="16">5.89</td>
<td id="76E9C2049E78C5CFCF9E4097493FFE21" box="[765,881,401,424]" gridcol="4" gridrow="3" pageId="15" pageNumber="16">6.04</td>
<td id="76E9C2049E78C5CFCEFF40974E5EFE21" box="[924,1040,401,424]" gridcol="5" gridrow="3" pageId="15" pageNumber="16">5.72</td>
<td id="76E9C2049E78C5CFC95440974EE5FE21" box="[1079,1195,401,424]" gridcol="6" gridrow="3" pageId="15" pageNumber="16">5.97</td>
</tr>
<tr id="3538AB789E78C5CFCD1440BF4EE5FE46" box="[119,1195,441,463]" gridrow="4" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1440BF4AB2FE46" box="[119,252,441,463]" gridcol="0" gridrow="4" pageId="15" pageNumber="16">BB</th>
<td id="76E9C2049E78C5CFCC7F40BF4BD9FE46" box="[284,407,441,463]" gridcol="1" gridrow="4" pageId="15" pageNumber="16">9.03</td>
<td id="76E9C2049E78C5CFCCDD40BF487CFE46" box="[446,562,441,463]" gridcol="2" gridrow="4" pageId="15" pageNumber="16">8.85</td>
<td id="76E9C2049E78C5CFCF3D40BF489FFE46" box="[606,721,441,463]" gridcol="3" gridrow="4" pageId="15" pageNumber="16">8.94</td>
<td id="76E9C2049E78C5CFCF9E40BF493FFE46" box="[765,881,441,463]" gridcol="4" gridrow="4" pageId="15" pageNumber="16">9.09</td>
<td id="76E9C2049E78C5CFCEFF40BF4E5EFE46" box="[924,1040,441,463]" gridcol="5" gridrow="4" pageId="15" pageNumber="16">8.78</td>
<td id="76E9C2049E78C5CFC95440BF4EE5FE46" box="[1079,1195,441,463]" gridcol="6" gridrow="4" pageId="15" pageNumber="16">8.84</td>
</tr>
<tr id="3538AB789E78C5CFCD1440E44EE5FE71" box="[119,1195,482,504]" gridrow="5" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1440E44AB2FE71" box="[119,252,482,504]" gridcol="0" gridrow="5" pageId="15" pageNumber="16">CIL</th>
<td id="76E9C2049E78C5CFCC7F40E44BD9FE71" box="[284,407,482,504]" gridcol="1" gridrow="5" pageId="15" pageNumber="16">16.97</td>
<td id="76E9C2049E78C5CFCCDD40E4487CFE71" box="[446,562,482,504]" gridcol="2" gridrow="5" pageId="15" pageNumber="16">16.61</td>
<td id="76E9C2049E78C5CFCF3D40E4489FFE71" box="[606,721,482,504]" gridcol="3" gridrow="5" pageId="15" pageNumber="16">16.10</td>
<td id="76E9C2049E78C5CFCF9E40E4493FFE71" box="[765,881,482,504]" gridcol="4" gridrow="5" pageId="15" pageNumber="16">16.69</td>
<td id="76E9C2049E78C5CFCEFF40E44E5EFE71" box="[924,1040,482,504]" gridcol="5" gridrow="5" pageId="15" pageNumber="16">17.06</td>
<td id="76E9C2049E78C5CFC95440E44EE5FE71" box="[1079,1195,482,504]" gridcol="6" gridrow="5" pageId="15" pageNumber="16">16.83</td>
</tr>
<tr id="3538AB789E78C5CFCD14430F4EE5FDA9" box="[119,1195,521,544]" gridrow="6" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD14430F4AB2FDA9" box="[119,252,521,544]" gridcol="0" gridrow="6" pageId="15" pageNumber="16">CCL</th>
<td id="76E9C2049E78C5CFCC7F430F4BD9FDA9" box="[284,407,521,544]" gridcol="1" gridrow="6" pageId="15" pageNumber="16">16.14</td>
<td id="76E9C2049E78C5CFCCDD430F487CFDA9" box="[446,562,521,544]" gridcol="2" gridrow="6" pageId="15" pageNumber="16">15.85</td>
<td id="76E9C2049E78C5CFCF3D430F489FFDA9" box="[606,721,521,544]" gridcol="3" gridrow="6" pageId="15" pageNumber="16">15.73</td>
<td id="76E9C2049E78C5CFCF9E430F493FFDA9" box="[765,881,521,544]" gridcol="4" gridrow="6" pageId="15" pageNumber="16">16.00</td>
<td id="76E9C2049E78C5CFCEFF430F4E5EFDA9" box="[924,1040,521,544]" gridcol="5" gridrow="6" pageId="15" pageNumber="16">16.32</td>
<td id="76E9C2049E78C5CFC954430F4EE5FDA9" box="[1079,1195,521,544]" gridcol="6" gridrow="6" pageId="15" pageNumber="16">16.30</td>
</tr>
<tr id="3538AB789E78C5CFCD1443374EE5FDC1" box="[119,1195,561,584]" gridrow="7" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1443374AB2FDC1" box="[119,252,561,584]" gridcol="0" gridrow="7" pageId="15" pageNumber="16">DENL</th>
<td id="76E9C2049E78C5CFCC7F43374BD9FDC1" box="[284,407,561,584]" gridcol="1" gridrow="7" pageId="15" pageNumber="16">11.62</td>
<td id="76E9C2049E78C5CFCCDD4337487CFDC1" box="[446,562,561,584]" gridcol="2" gridrow="7" pageId="15" pageNumber="16">11.33</td>
<td id="76E9C2049E78C5CFCF3D4337489FFDC1" box="[606,721,561,584]" gridcol="3" gridrow="7" pageId="15" pageNumber="16">11.51</td>
<td id="76E9C2049E78C5CFCF9E4337493FFDC1" box="[765,881,561,584]" gridcol="4" gridrow="7" pageId="15" pageNumber="16">11.68</td>
<td id="76E9C2049E78C5CFCEFF43374E5EFDC1" box="[924,1040,561,584]" gridcol="5" gridrow="7" pageId="15" pageNumber="16">11.70</td>
<td id="76E9C2049E78C5CFC95443374EE5FDC1" box="[1079,1195,561,584]" gridcol="6" gridrow="7" pageId="15" pageNumber="16">11.75</td>
</tr>
<tr id="3538AB789E78C5CFCD14435F4EE5FDE6" box="[119,1195,601,623]" gridrow="8" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD14435F4AB2FDE6" box="[119,252,601,623]" gridcol="0" gridrow="8" pageId="15" pageNumber="16">GLS</th>
<td id="76E9C2049E78C5CFCC7F435F4BD9FDE6" box="[284,407,601,623]" gridcol="1" gridrow="8" pageId="15" pageNumber="16">19.17</td>
<td id="76E9C2049E78C5CFCCDD435F487CFDE6" box="[446,562,601,623]" gridcol="2" gridrow="8" pageId="15" pageNumber="16">18.72</td>
<td id="76E9C2049E78C5CFCF3D435F489FFDE6" box="[606,721,601,623]" gridcol="3" gridrow="8" pageId="15" pageNumber="16">18.54</td>
<td id="76E9C2049E78C5CFCF9E435F493FFDE6" box="[765,881,601,623]" gridcol="4" gridrow="8" pageId="15" pageNumber="16">18.81</td>
<td id="76E9C2049E78C5CFCEFF435F4E5EFDE6" box="[924,1040,601,623]" gridcol="5" gridrow="8" pageId="15" pageNumber="16">19.04</td>
<td id="76E9C2049E78C5CFC954435F4EE5FDE6" box="[1079,1195,601,623]" gridcol="6" gridrow="8" pageId="15" pageNumber="16">18.85</td>
</tr>
<tr id="3538AB789E78C5CFCD1443874EE5FD11" box="[119,1195,641,664]" gridrow="9" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1443874AB2FD11" box="[119,252,641,664]" gridcol="0" gridrow="9" pageId="15" pageNumber="16">PB</th>
<td id="76E9C2049E78C5CFCC7F43874BD9FD11" box="[284,407,641,664]" gridcol="1" gridrow="9" pageId="15" pageNumber="16">4.92</td>
<td id="76E9C2049E78C5CFCCDD4387487CFD11" box="[446,562,641,664]" gridcol="2" gridrow="9" pageId="15" pageNumber="16">5.00</td>
<td id="76E9C2049E78C5CFCF3D4387489FFD11" box="[606,721,641,664]" gridcol="3" gridrow="9" pageId="15" pageNumber="16">4.72</td>
<td id="76E9C2049E78C5CFCF9E4387493FFD11" box="[765,881,641,664]" gridcol="4" gridrow="9" pageId="15" pageNumber="16">5.03</td>
<td id="76E9C2049E78C5CFCEFF43874E5EFD11" box="[924,1040,641,664]" gridcol="5" gridrow="9" pageId="15" pageNumber="16">4.85</td>
<td id="76E9C2049E78C5CFC95443874EE5FD11" box="[1079,1195,641,664]" gridcol="6" gridrow="9" pageId="15" pageNumber="16">4.69</td>
</tr>
<tr id="3538AB789E78C5CFCD1443AF4EE5FD49" box="[119,1195,681,704]" gridrow="10" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1443AF4AB2FD49" box="[119,252,681,704]" gridcol="0" gridrow="10" pageId="15" pageNumber="16">MANDL</th>
<td id="76E9C2049E78C5CFCC7F43AF4BD9FD49" box="[284,407,681,704]" gridcol="1" gridrow="10" pageId="15" pageNumber="16">5.73</td>
<td id="76E9C2049E78C5CFCCDD43AF487CFD49" box="[446,562,681,704]" gridcol="2" gridrow="10" pageId="15" pageNumber="16">6.11</td>
<td id="76E9C2049E78C5CFCF3D43AF489FFD49" box="[606,721,681,704]" gridcol="3" gridrow="10" pageId="15" pageNumber="16">5.74</td>
<td id="76E9C2049E78C5CFCF9E43AF493FFD49" box="[765,881,681,704]" gridcol="4" gridrow="10" pageId="15" pageNumber="16">5.63</td>
<td id="76E9C2049E78C5CFCEFF43AF4E5EFD49" box="[924,1040,681,704]" gridcol="5" gridrow="10" pageId="15" pageNumber="16">6.24</td>
<td id="76E9C2049E78C5CFC95443AF4EE5FD49" box="[1079,1195,681,704]" gridcol="6" gridrow="10" pageId="15" pageNumber="16">5.62</td>
</tr>
<tr id="3538AB789E78C5CFCD1443D74EE5FD61" box="[119,1195,721,744]" gridrow="11" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1443D74AB2FD61" box="[119,252,721,744]" gridcol="0" gridrow="11" pageId="15" pageNumber="16">MTRL</th>
<td id="76E9C2049E78C5CFCC7F43D74BD9FD61" box="[284,407,721,744]" gridcol="1" gridrow="11" pageId="15" pageNumber="16">5.00</td>
<td id="76E9C2049E78C5CFCCDD43D7487CFD61" box="[446,562,721,744]" gridcol="2" gridrow="11" pageId="15" pageNumber="16">5.00</td>
<td id="76E9C2049E78C5CFCF3D43D7489FFD61" box="[606,721,721,744]" gridcol="3" gridrow="11" pageId="15" pageNumber="16">5.20</td>
<td id="76E9C2049E78C5CFCF9E43D7493FFD61" box="[765,881,721,744]" gridcol="4" gridrow="11" pageId="15" pageNumber="16">4.73</td>
<td id="76E9C2049E78C5CFCEFF43D74E5EFD61" box="[924,1040,721,744]" gridcol="5" gridrow="11" pageId="15" pageNumber="16">5.00</td>
<td id="76E9C2049E78C5CFC95443D74EE5FD61" box="[1079,1195,721,744]" gridcol="6" gridrow="11" pageId="15" pageNumber="16">4.96</td>
</tr>
<tr id="3538AB789E78C5CFCD1443FF4EE5FC99" box="[119,1195,761,784]" gridrow="12" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1443FF4AB2FC99" box="[119,252,761,784]" gridcol="0" gridrow="12" pageId="15" pageNumber="16">PL</th>
<td id="76E9C2049E78C5CFCC7F43FF4BD9FC99" box="[284,407,761,784]" gridcol="1" gridrow="12" pageId="15" pageNumber="16">7.45</td>
<td id="76E9C2049E78C5CFCCDD43FF487CFC99" box="[446,562,761,784]" gridcol="2" gridrow="12" pageId="15" pageNumber="16">7.13</td>
<td id="76E9C2049E78C5CFCF3D43FF489FFC99" box="[606,721,761,784]" gridcol="3" gridrow="12" pageId="15" pageNumber="16">7.46</td>
<td id="76E9C2049E78C5CFCF9E43FF493FFC99" box="[765,881,761,784]" gridcol="4" gridrow="12" pageId="15" pageNumber="16">7.37</td>
<td id="76E9C2049E78C5CFCEFF43FF4E5EFC99" box="[924,1040,761,784]" gridcol="5" gridrow="12" pageId="15" pageNumber="16">8.50</td>
<td id="76E9C2049E78C5CFC95443FF4EE5FC99" box="[1079,1195,761,784]" gridcol="6" gridrow="12" pageId="15" pageNumber="16">7.40</td>
</tr>
<tr id="3538AB789E78C5CFCD1442274EE5FCB1" box="[119,1195,801,824]" gridrow="13" pageId="15" pageNumber="16">
<th id="76E9C2049E78C5CFCD1442274AB2FCB1" box="[119,252,801,824]" gridcol="0" gridrow="13" pageId="15" pageNumber="16">ZB</th>
<td id="76E9C2049E78C5CFCC7F42274BD9FCB1" box="[284,407,801,824]" gridcol="1" gridrow="13" pageId="15" pageNumber="16">10.93</td>
<td id="76E9C2049E78C5CFCCDD4227487CFCB1" box="[446,562,801,824]" gridcol="2" gridrow="13" pageId="15" pageNumber="16">10.46</td>
<td id="76E9C2049E78C5CFCF3D4227489FFCB1" box="[606,721,801,824]" gridcol="3" gridrow="13" pageId="15" pageNumber="16">10.55</td>
<td id="76E9C2049E78C5CFCF9E4227493FFCB1" box="[765,881,801,824]" gridcol="4" gridrow="13" pageId="15" pageNumber="16">11.04</td>
<td id="76E9C2049E78C5CFCEFF42274E5EFCB1" box="[924,1040,801,824]" gridcol="5" gridrow="13" pageId="15" pageNumber="16">10.78</td>
<td id="76E9C2049E78C5CFC95442274EE5FCB1" box="[1079,1195,801,824]" gridcol="6" gridrow="13" pageId="15" pageNumber="16">10.38</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB7A93A9E783A3FCD0F424B4A9DFCED" blockId="15.[108,211,845,868]" box="[108,211,845,868]" pageId="15" pageNumber="16">
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<superScript id="7C7D04729E783A3FCD0F424B4A3DFCD2" attach="left" box="[108,115,845,859]" fontSize="6" pageId="15" pageNumber="16">a</superScript>
Holotype.
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</paragraph>
<subSubSection id="C312FAB19E783A3FCDFF45D149FAFA25" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BB7A93A9E783A3FCDFF45D149FAFA25" blockId="15.[108,1213,903,1452]" pageId="15" pageNumber="16">
ETYMOLOGY: The epithet
<taxonomicName id="4C08D2B99E783A3FCCA545D1485BFB78" authorityName="Yánez-Fernández &amp; Marchán-Rivadeneira &amp; Velazco &amp; Burneo &amp; Tinoco &amp; Camacho" authorityYear="2023" box="[454,533,1239,1265]" class="Mammalia" family="Phyllostomidae" genus="Sturnira" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="boadai">
<emphasis id="B97C75289E783A3FCCA545D1485BFB78" box="[454,533,1239,1265]" italics="true" pageId="15" pageNumber="16">boadai</emphasis>
</taxonomicName>
is dedicated to the memory of the Ecuadorian mammalogist Carlos Boada (19732015). Carlos was passionate about studying small mammals, especially bats and rodents. His academic contributions to the knowledge of Ecuadorian mammals were remarkable and primarily included taxonomic assessments and biological inventories. Carlos trained an extensive group of young mammalogists in the country, and herein we commemorate his early departure by naming this new species in his honor.
</paragraph>
</subSubSection>
</treatment>
</document>