550 lines
56 KiB
XML
550 lines
56 KiB
XML
<document ID-DOI="10.11609/JoTT.o2363.789-96" ID-ISSN="0974-7907" ID-Zenodo-Dep="4986617" LSID="urn:lsid:zoobank.org:act:6BED3074-F395-4F56-90BF-D6DE0D712385" approvalRequired="28" approvalRequired_for_document="1" approvalRequired_for_illustrations="1" approvalRequired_for_textStreams="26" checkinTime="1624036298780" checkinUser="carolina" docAuthor="Mirza, Zeeshan A. & Sanap, Rajesh V." docDate="2010" docId="B10435405972FFFEFEA6FCADFB82FAE5" docLanguage="en" docName="JThreatenedTaxa.2.4.789-796.pdf" docOrigin="Journal of Threatened Taxa 2 (4)" docSource="http://threatenedtaxa.org/index.php/JoTT/article/view/480" docStyle="DocumentStyle:AA2961C68A601FE2D2D9ECE51070E01E:JThreatenedTaxa.2009-2012.journal_article" docStyleId="AA2961C68A601FE2D2D9ECE51070E01E" docStyleName="JThreatenedTaxa.2009-2012.journal_article" docTitle="Lychas aareyensis Mirza & Sanap 2010, sp. nov." docType="treatment" docUuid="6BED3074-F395-4F56-90BF-D6DE0D712385" docUuidSource="ZooBank" docVersion="2" lastPageId="5" lastPageNumber="793" masterDocId="4D3D4D385973FFFBFF98FFC2FFA7FFA1" masterDocTitle="Description of a new species of scorpion of the genus Lychas C. L. Koch, 1845 (Scorpiones: Buthidae) from Maharashtra, India" masterLastPageNumber="796" masterPageNumber="789" pageId="1" pageNumber="790" updateTime="1624036618657" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of a new species of scorpion of the genus Lychas C. L. Koch, 1845 (Scorpiones: Buthidae) from Maharashtra, India</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mirza, Zeeshan A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sanap, Rajesh V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Threatened Taxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-04-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>789</mods:start>
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<mods:end>796</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://threatenedtaxa.org/index.php/JoTT/article/view/480</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11609/JoTT.o2363.789-96</mods:identifier>
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<mods:identifier type="ISSN">0974-7907</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4986617</mods:identifier>
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<mods:identifier type="ZooBank">CF0E2DE1-A65E-4A1B-8FBD-3D1042411FD9</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:6BED3074-F395-4F56-90BF-D6DE0D712385" httpUri="http://treatment.plazi.org/id/B10435405972FFFEFEA6FCADFB82FAE5" lastPageId="5" lastPageNumber="793" pageId="1" pageNumber="790">
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<subSubSection box="[318,627,879,902]" pageId="1" pageNumber="790" type="nomenclature">
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<paragraph blockId="1.[318,627,879,934]" box="[318,627,879,902]" pageId="1" pageNumber="790">
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<heading box="[318,627,879,902]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="790" reason="3">
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<emphasis bold="true" box="[318,627,879,902]" pageId="1" pageNumber="790">
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<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[318,531,879,902]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="790" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
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<emphasis bold="true" box="[318,531,879,902]" italics="true" pageId="1" pageNumber="790">Lychas aareyensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[538,627,879,902]" pageId="1" pageNumber="790" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[350,595,911,934]" pageId="1" pageNumber="790" type="description">
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<paragraph blockId="1.[318,627,879,934]" box="[350,595,911,934]" pageId="1" pageNumber="790">
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(
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<figureCitation box="[356,483,911,934]" captionStart-0="Images 1" captionStart-1="Images 10" captionStart-2="Image 13" captionStart-3="Images 14 & 15" captionStartId-0="2.[170,234,1300,1318]" captionStartId-1="2.[170,234,1904,1922]" captionStartId-2="4.[170,232,804,824]" captionStartId-3="5.[170,244,687,707]" captionTargetBox-0="[170,1418,238,1285]" captionTargetBox-1="[170,1419,1399,1887]" captionTargetBox-2="[170,1257,238,792]" captionTargetBox-3="[171,1418,237,669]" captionTargetId-0="figure-27@2.[598,992,238,788]" captionTargetId-1="figure-126@2.[598,988,1400,1756]" captionTargetId-2="figure-25@4.[167,1259,235,794]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Images 1 to 9. Lychas aareyensis sp. nov. female holotype (BNHS SC23) (scale 1.0mm) 1 - Carapace and Mesosoma; 2 - Genital operculum and pectinal teeth; 3 - Mesosomal tergites, dorsal; 4 - Mesosomal sternites, ventral; 5 - Chelicerae; 6 - Manus; 7 - Patella; 8 - Metasoma; 9 - Telson" captionText-1="Images 10 to 12. Lychas aareyensis sp. nov. male paratype (BNHS SC24) (scale 1.0mm) 10 - Carapace; 11 - Genital operculum and pectined; 12 - Mesosoma ventral (no scale)" captionText-2="Image 13. Lychas aareyensis sp. nov. female holotype BNHS SC23 in life" captionText-3="Images 14 & 15. Lychas aareyensis sp. nov. female with juveniles (not collected)" figureDoi-0="http://doi.org/10.5281/zenodo.4986621" figureDoi-1="http://doi.org/10.5281/zenodo.4986623" figureDoi-2="http://doi.org/10.5281/zenodo.4986627" figureDoi-3="http://doi.org/10.5281/zenodo.4986629" httpUri-0="https://zenodo.org/record/4986621/files/figure.png" httpUri-1="https://zenodo.org/record/4986623/files/figure.png" httpUri-2="https://zenodo.org/record/4986627/files/figure.png" httpUri-3="https://zenodo.org/record/4986629/files/figure.png" pageId="1" pageNumber="790">Image 1-15</figureCitation>
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,
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<figureCitation box="[493,586,911,934]" captionStart="Figures 1-3" captionStartId="1.[812,879,1880,1898]" captionTargetBox="[818,1415,1370,1864]" captionTargetId="figure-623@1.[809,1420,1365,1868]" captionTargetPageId="1" captionText="Figures 1-3. Lychas aareyensis sp. nov. female holotype" figureDoi="http://doi.org/10.5281/zenodo.4986619" httpUri="https://zenodo.org/record/4986619/files/figure.png" pageId="1" pageNumber="790">Figs. 1-3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="790" type="materials_examined">
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<paragraph blockId="1.[170,776,975,1189]" box="[170,378,975,998]" pageId="1" pageNumber="790">
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<emphasis bold="true" box="[170,378,975,998]" pageId="1" pageNumber="790">Material examined</emphasis>
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</paragraph>
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<paragraph blockId="1.[170,776,975,1189]" pageId="1" pageNumber="790">
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<materialsCitation collectingDate="2009-03-14" collectionCode="BNHS" collectorName="Rajesh Sanap & Zeeshan A. Mirza" country="India" elevation="103" latitude="19.145481" location="Maharashtra" longLatPrecision="1" longitude="72.88206" pageId="1" pageNumber="790" specimenCode="SC-23" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
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<typeStatus box="[208,307,1007,1030]" pageId="1" pageNumber="790">Holotype</typeStatus>
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: Female,
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<date box="[439,552,1007,1030]" pageId="1" pageNumber="790" value="2009-03-14">
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<collectingDate box="[439,552,1007,1030]" pageId="1" pageNumber="790" value="2009-03-14">14.iii.2009</collectingDate>
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</date>
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,
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<geoCoordinate box="[576,716,1007,1030]" degrees="19" direction="north" minutes="8" orientation="latitude" pageId="1" pageNumber="790" precision="1" seconds="43.73" value="19.145481">19°8’43.73”N</geoCoordinate>
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&
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<geoCoordinate box="[170,327,1039,1062]" degrees="72" direction="east" minutes="52" orientation="longitude" pageId="1" pageNumber="790" precision="1" seconds="55.39" value="72.88206">72°52’55.39”E</geoCoordinate>
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,
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<quantity box="[350,414,1039,1062]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="1" pageNumber="790" unit="m" value="103.0">
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<elevation box="[350,414,1039,1062]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="1" pageNumber="790" unit="m" value="103.0">103m</elevation>
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</quantity>
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, Aarey Milk Colony, Mumbai,
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<collectingRegion box="[170,309,1070,1093]" country="India" name="Maharashtra" pageId="1" pageNumber="790">Maharashtra</collectingRegion>
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,
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<collectingCountry box="[321,377,1070,1093]" name="India" pageId="1" pageNumber="790">India</collectingCountry>
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, coll.
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<collectorName box="[440,593,1070,1093]" pageId="1" pageNumber="790">Rajesh Sanap</collectorName>
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and
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<collectorName pageId="1" pageNumber="790">Zeeshan A. Mirza</collectorName>
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(
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<collectionCode box="[241,310,1102,1125]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="1" pageNumber="790">BNHS</collectionCode>
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<specimenCode box="[317,391,1102,1125]" collectionCode="SC-" pageId="1" pageNumber="790">SC-23</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="1.[170,776,975,1189]" pageId="1" pageNumber="790">
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<materialsCitation box="[208,684,1134,1157]" collectionCode="BNHS" pageId="1" pageNumber="790" specimenCode="SC-25, SC-26" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
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<typeStatus box="[208,308,1134,1157]" pageId="1" pageNumber="790">Paratype</typeStatus>
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:
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<specimenCount box="[322,431,1134,1157]" pageId="1" pageNumber="790" type="female">2 females</specimenCount>
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(
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<collectionCode box="[448,517,1134,1157]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="1" pageNumber="790">BNHS</collectionCode>
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<specimenCode box="[529,600,1134,1157]" collectionCode="SC-" pageId="1" pageNumber="790">SC-25</specimenCode>
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/
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<specimenCode box="[607,678,1134,1157]" collectionCode="SC-" pageId="1" pageNumber="790">SC-26</specimenCode>
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)
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</materialsCitation>
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,
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<materialsCitation collectionCode="BNHS" pageId="1" pageNumber="790" specimenCode="SC-24" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount box="[698,775,1134,1157]" pageId="1" pageNumber="790" type="male">1 male</specimenCount>
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(
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<collectionCode box="[176,245,1166,1189]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="1" pageNumber="790">BNHS</collectionCode>
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<specimenCode box="[251,325,1166,1189]" collectionCode="SC-" pageId="1" pageNumber="790">SC-24</specimenCode>
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)
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</materialsCitation>
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,
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<materialsCitation box="[341,561,1166,1189]" collectionCode="BNHS" pageId="1" pageNumber="790" specimenCount="1" typeStatus="paratype">same data as above</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="790" type="etymology">
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<paragraph blockId="1.[170,775,1230,1317]" box="[170,293,1230,1253]" pageId="1" pageNumber="790">
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<emphasis bold="true" box="[170,293,1230,1253]" pageId="1" pageNumber="790">Etymology</emphasis>
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</paragraph>
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<paragraph blockId="1.[170,775,1230,1317]" pageId="1" pageNumber="790">
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The specific epithet refers to Aarey, the
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<typeStatus box="[631,676,1262,1285]" pageId="1" pageNumber="790">type</typeStatus>
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and only known locality for the new species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="790" type="diagnosis">
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<paragraph blockId="1.[170,775,1358,1669]" box="[170,285,1358,1381]" pageId="1" pageNumber="790">
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<emphasis bold="true" box="[170,285,1358,1381]" pageId="1" pageNumber="790">Diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="1.[170,775,1358,1669]" pageId="1" pageNumber="790">
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A small-sized scorpion which differs from other species of the genus as follows: median eye situated anteriorly in the ratio 1:1.6; 13-13 pectinal teeth; smaller size
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<quantity box="[675,775,1454,1477]" metricMagnitude="-2" metricUnit="m" metricValue="3.165" pageId="1" pageNumber="790" unit="mm" value="31.65">31.65mm</quantity>
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(maximum); external granule absent on immovable and present on movable finger of female; trichobothrium db between
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<emphasis box="[267,287,1550,1573]" italics="true" pageId="1" pageNumber="790">et</emphasis>
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and
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<emphasis box="[339,371,1550,1573]" italics="true" pageId="1" pageNumber="790">est</emphasis>
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situated closer to
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<emphasis box="[563,595,1550,1573]" italics="true" pageId="1" pageNumber="790">est</emphasis>
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; pedipalp patella in shade of orange with faint brown reticulate markings or spots; metasomal segment I and II with 10 keels and III with eight.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="793" pageId="1" pageNumber="790" type="description">
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<paragraph blockId="1.[170,776,1710,2021]" box="[170,302,1710,1733]" pageId="1" pageNumber="790">
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<emphasis bold="true" box="[170,302,1710,1733]" pageId="1" pageNumber="790">Description</emphasis>
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</paragraph>
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<paragraph blockId="1.[170,776,1710,2021]" lastBlockId="1.[813,1425,238,1317]" pageId="1" pageNumber="790">
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Measurements in
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<tableCitation box="[409,492,1742,1765]" captionStart="Table 1" captionStartId="3.[170,224,238,258]" captionText="Table 1. Detailed measurements of the types of Lychas aareyensis sp. nov. (in mm.)." pageId="1" pageNumber="790">Table 1</tableCitation>
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. Coloration: Basically in a shade of orange or yellow, symmetrically marbled with brown producing an overall spotted appearance (
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<figureCitation box="[176,287,1838,1861]" captionStart="Image 13" captionStartId="4.[170,232,804,824]" captionTargetBox="[170,1257,238,792]" captionTargetId="figure-25@4.[167,1259,235,794]" captionTargetPageId="4" captionText="Image 13. Lychas aareyensis sp. nov. female holotype BNHS SC23 in life" figureDoi="http://doi.org/10.5281/zenodo.4986627" httpUri="https://zenodo.org/record/4986627/files/figure.png" pageId="1" pageNumber="790">Image 13</figureCitation>
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). Carapace dark orange with random dark brown markings; eyes surrounded with black pigment. Mesosoma: brown with orange confluent spots on the distal regions of each tergites. Metasoma: segments I to III light brownish, segment IV more darker and segment V entirely black. Vesicle brownish-red. Venter yellowish with some diffuse brown spots on sternites V to VII. Chelicerae yellowish with dark variegated pigmentation; fingers dark brown with some light pigmentation. Pedipalp: yellowishorange with the femur and patella spotted or with reticulate markings; chela hand yellowish-orange; fingers darker and brownish. Legs yellowish with several diffuse spots
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</paragraph>
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<paragraph blockId="1.[813,1425,238,1317]" pageId="1" pageNumber="790">
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Surface of carapace coarsely granular with a few smooth patches (
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<figureCitation box="[1012,1108,462,485]" captionStart="Images 1" captionStartId="2.[170,234,1300,1318]" captionTargetBox="[170,1418,238,1285]" captionTargetId="figure-27@2.[598,992,238,788]" captionTargetPageId="2" captionText="Images 1 to 9. Lychas aareyensis sp. nov. female holotype (BNHS SC23) (scale 1.0mm) 1 - Carapace and Mesosoma; 2 - Genital operculum and pectinal teeth; 3 - Mesosomal tergites, dorsal; 4 - Mesosomal sternites, ventral; 5 - Chelicerae; 6 - Manus; 7 - Patella; 8 - Metasoma; 9 - Telson" figureDoi="http://doi.org/10.5281/zenodo.4986621" httpUri="https://zenodo.org/record/4986621/files/figure.png" pageId="1" pageNumber="790">Image 1</figureCitation>
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). A pair of median eyes situated anteriorly in the ratio 1:1.6. Median anterior and median central furrows shallow, the former wider; median posterior and post marginal furrows forming a deep triangular depression. Five pairs of lateral eyes present on anteriolateral portion with smooth lateral ocular tubercles of which two anterior pairs directed anterio-laterally. third pair posterio-laterally and last two pairs dorso-posteriorly and posteriorly respectively. Interocular portion black. Carinae absent, median lateral carinae poorly developed with a row of granules. Anterior margin introverted slightly and smooth; lateral margins finely crenulated and the posterior margin fairly smooth with some large granules on the periphery. Chelicerae smooth on basal segment (Image 5) with typical buthid dentition except only one tooth on ventral surface of immovable finger.
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</paragraph>
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<paragraph blockId="1.[813,1425,238,1317]" pageId="1" pageNumber="790">
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Femur carinae crenulated; the length of femur is shorter than carapace and patella. Patella carinated and dorso-inferior carinae evenly crenulated (Image 7). Manus smooth, without carinae (Image 6) and length of chelae twice the length of femur. Fingers as long as patella. smooth. Dentition on fingers overlapping. External granule absent on the immovable finger but a single external granule present on the movable finger and a strong apical tooth present in both sexes. Trichobothrial pattern on pedipalp as in
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<figureCitation box="[1092,1182,1262,1285]" captionStart="Figures 1-3" captionStartId="1.[812,879,1880,1898]" captionTargetBox="[818,1415,1370,1864]" captionTargetId="figure-623@1.[809,1420,1365,1868]" captionTargetPageId="1" captionText="Figures 1-3. Lychas aareyensis sp. nov. female holotype" figureDoi="http://doi.org/10.5281/zenodo.4986619" httpUri="https://zenodo.org/record/4986619/files/figure.png" pageId="1" pageNumber="790">Figs 1-3</figureCitation>
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. Legs finely granular and carinated on femur and patella. Tibia smooth with
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4986619" ID-Zenodo-Dep="4986619" box="[812,1310,1880,1898]" httpUri="https://zenodo.org/record/4986619/files/figure.png" pageId="1" pageNumber="790" startId="1.[812,879,1880,1898]" targetBox="[818,1415,1370,1864]" targetPageId="1">
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<paragraph blockId="1.[812,1401,1880,2010]" box="[812,1310,1880,1898]" pageId="1" pageNumber="790">
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Figures 1-3.
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<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[922,1088,1880,1898]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="790" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
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<emphasis bold="true" box="[922,1088,1880,1898]" italics="true" pageId="1" pageNumber="790">Lychas aareyensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1093,1163,1880,1898]" pageId="1" pageNumber="790" rank="species">sp. nov.</taxonomicNameLabel>
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female holotype
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</paragraph>
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</caption>
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||
<paragraph blockId="1.[812,1401,1880,2010]" box="[812,1110,1903,1921]" pageId="1" pageNumber="790">
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||
<emphasis bold="true" box="[812,989,1903,1921]" pageId="1" pageNumber="790">tricobothrial pattern</emphasis>
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. Not on scale.
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</paragraph>
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<paragraph blockId="1.[812,1401,1880,2010]" box="[812,1202,1925,1943]" pageId="1" pageNumber="790">1 - Manus; 2 - Patella, dorsal; 3 - Femur, dorsal</paragraph>
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<paragraph blockId="1.[812,1401,1880,2010]" pageId="1" pageNumber="790">d – dorsal; e – external; dt – dorsal terminal; db – dorsal basal; et – external terminal; est – external sub-terminal; esb – external sub-basal; eb – external basal</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4986621" ID-Zenodo-Dep="4986621" httpUri="https://zenodo.org/record/4986621/files/figure.png" pageId="2" pageNumber="791" startId="2.[170,234,1300,1318]" targetBox="[170,1418,238,1285]" targetPageId="2">
|
||
<paragraph blockId="2.[170,1419,1300,1430]" pageId="2" pageNumber="791">
|
||
<emphasis bold="true" box="[170,937,1300,1318]" pageId="2" pageNumber="791">
|
||
Images 1 to 9.
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[299,465,1300,1318]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="791" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[299,465,1300,1318]" italics="true" pageId="2" pageNumber="791">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[470,540,1300,1318]" pageId="2" pageNumber="791" rank="species">sp. nov.</taxonomicNameLabel>
|
||
female holotype (BNHS SC23) (scale 1.0mm)
|
||
</emphasis>
|
||
1 - Carapace and Mesosoma; 2 - Genital operculum and pectinal teeth; 3 - Mesosomal tergites, dorsal; 4 - Mesosomal sternites, ventral; 5 - Chelicerae; 6 - Manus; 7 - Patella; 8 - Metasoma; 9 - Telson
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4986623" ID-Zenodo-Dep="4986623" httpUri="https://zenodo.org/record/4986623/files/figure.png" pageId="2" pageNumber="791" startId="2.[170,234,1904,1922]" targetBox="[170,1419,1399,1887]" targetPageId="2">
|
||
<paragraph blockId="2.[170,941,1904,1944]" pageId="2" pageNumber="791">
|
||
<emphasis bold="true" box="[170,941,1904,1922]" pageId="2" pageNumber="791">
|
||
Images 10 to 12.
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[319,486,1904,1922]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="791" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[319,486,1904,1922]" italics="true" pageId="2" pageNumber="791">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[491,561,1904,1922]" pageId="2" pageNumber="791" rank="species">sp. nov.</taxonomicNameLabel>
|
||
male paratype (BNHS SC24) (scale 1.0mm)
|
||
</emphasis>
|
||
10 - Carapace; 11 - Genital operculum and pectined; 12 - Mesosoma ventral (no scale)
|
||
</paragraph>
|
||
</caption>
|
||
<caption box="[170,1021,238,258]" pageId="3" pageNumber="792" startId="3.[170,224,238,258]">
|
||
<paragraph blockId="3.[170,1021,238,258]" box="[170,1021,238,258]" pageId="3" pageNumber="792">
|
||
Table 1. Detailed measurements of the types of
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[651,840,238,258]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="792" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[651,840,238,258]" italics="true" pageId="3" pageNumber="792">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[847,926,238,258]" pageId="3" pageNumber="792" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(in mm.).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="792">
|
||
<table box="[186,1037,290,1429]" gridcols="5" gridrows="18" pageId="3" pageNumber="792">
|
||
<tr box="[186,1037,290,308]" gridrow="0" pageId="3" pageNumber="792" rowspan-0="1" rowspan-3="1" rowspan-4="1">
|
||
<td box="[494,764,290,308]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="3" pageNumber="792">
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[539,693,290,308]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="792" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis box="[539,693,290,308]" italics="true" pageId="3" pageNumber="792">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[698,764,290,308]" pageId="3" pageNumber="792" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</td>
|
||
</tr>
|
||
<tr box="[186,1037,327,390]" gridrow="1" pageId="3" pageNumber="792" rowspan-0="1">
|
||
<td box="[494,598,327,390]" gridcol="1" gridrow="1" pageId="3" pageNumber="792">Holotype Female BNHS SC23</td>
|
||
<td box="[640,764,327,390]" gridcol="2" gridrow="1" pageId="3" pageNumber="792">Paratype Male BNHS SC24</td>
|
||
<td box="[787,891,327,390]" gridcol="3" gridrow="1" pageId="3" pageNumber="792">Paratype Female BNHS SC25</td>
|
||
<td box="[933,1037,327,390]" gridcol="4" gridrow="1" pageId="3" pageNumber="792">Paratype Female BNHS SC26</td>
|
||
</tr>
|
||
<tr box="[186,1037,409,427]" gridrow="2" pageId="3" pageNumber="792">
|
||
<td box="[186,442,409,427]" gridcol="0" gridrow="2" pageId="3" pageNumber="792">Body length</td>
|
||
<td box="[494,598,409,427]" gridcol="1" gridrow="2" pageId="3" pageNumber="792">31.65</td>
|
||
<td box="[640,764,409,427]" gridcol="2" gridrow="2" pageId="3" pageNumber="792">*</td>
|
||
<td box="[787,891,409,427]" gridcol="3" gridrow="2" pageId="3" pageNumber="792">27.19</td>
|
||
<td box="[933,1037,409,427]" gridcol="4" gridrow="2" pageId="3" pageNumber="792">31.36</td>
|
||
</tr>
|
||
<tr box="[186,1037,455,518]" gridrow="3" pageId="3" pageNumber="792">
|
||
<td box="[186,442,455,518]" gridcol="0" gridrow="3" pageId="3" pageNumber="792">Carapace Length Carapace Anterior Width Carapace Posterior Width</td>
|
||
<td box="[494,598,455,518]" gridcol="1" gridrow="3" pageId="3" pageNumber="792">3.63 2.29 3.71</td>
|
||
<td box="[640,764,455,518]" gridcol="2" gridrow="3" pageId="3" pageNumber="792">3.55 1.78 3.75</td>
|
||
<td box="[787,891,455,518]" gridcol="3" gridrow="3" pageId="3" pageNumber="792">3.25 1.82 3.06</td>
|
||
<td box="[933,1037,455,518]" gridcol="4" gridrow="3" pageId="3" pageNumber="792">3.78 2.89 3.36</td>
|
||
</tr>
|
||
<tr box="[186,1037,549,567]" gridrow="4" pageId="3" pageNumber="792">
|
||
<td box="[186,442,549,567]" gridcol="0" gridrow="4" pageId="3" pageNumber="792">Mesosoma Length</td>
|
||
<td box="[494,598,549,567]" gridcol="1" gridrow="4" pageId="3" pageNumber="792">9.61</td>
|
||
<td box="[640,764,549,567]" gridcol="2" gridrow="4" pageId="3" pageNumber="792">7.50</td>
|
||
<td box="[787,891,549,567]" gridcol="3" gridrow="4" pageId="3" pageNumber="792">7.65</td>
|
||
<td box="[933,1037,549,567]" gridcol="4" gridrow="4" pageId="3" pageNumber="792">10.08</td>
|
||
</tr>
|
||
<tr box="[186,1037,591,609]" gridrow="5" pageId="3" pageNumber="792">
|
||
<td box="[186,442,591,609]" gridcol="0" gridrow="5" pageId="3" pageNumber="792">Metasoma Length</td>
|
||
<td box="[494,598,591,609]" gridcol="1" gridrow="5" pageId="3" pageNumber="792">18.41</td>
|
||
<td box="[640,764,591,609]" gridcol="2" gridrow="5" pageId="3" pageNumber="792">*</td>
|
||
<td box="[787,891,591,609]" gridcol="3" gridrow="5" pageId="3" pageNumber="792">16.29</td>
|
||
<td box="[933,1037,591,609]" gridcol="4" gridrow="5" pageId="3" pageNumber="792">17.5</td>
|
||
</tr>
|
||
<tr box="[186,1037,633,674]" gridrow="6" pageId="3" pageNumber="792">
|
||
<td box="[186,442,633,674]" gridcol="0" gridrow="6" pageId="3" pageNumber="792">Metasomal Segment I Length Width</td>
|
||
<td box="[494,598,633,674]" gridcol="1" gridrow="6" pageId="3" pageNumber="792">1.97 1.90</td>
|
||
<td box="[640,764,633,674]" gridcol="2" gridrow="6" pageId="3" pageNumber="792">2.03 1.59</td>
|
||
<td box="[787,891,633,674]" gridcol="3" gridrow="6" pageId="3" pageNumber="792">1.55 1.66</td>
|
||
<td box="[933,1037,633,674]" gridcol="4" gridrow="6" pageId="3" pageNumber="792">2.19 1.87</td>
|
||
</tr>
|
||
<tr box="[186,1037,702,743]" gridrow="7" pageId="3" pageNumber="792">
|
||
<td box="[186,442,702,743]" gridcol="0" gridrow="7" pageId="3" pageNumber="792">Metasomal Segment II Length Width</td>
|
||
<td box="[494,598,702,743]" gridcol="1" gridrow="7" pageId="3" pageNumber="792">2.47 1.72</td>
|
||
<td box="[640,764,702,743]" gridcol="2" gridrow="7" pageId="3" pageNumber="792">2.55 1.33</td>
|
||
<td box="[787,891,702,743]" gridcol="3" gridrow="7" pageId="3" pageNumber="792">2.16 1.54</td>
|
||
<td box="[933,1037,702,743]" gridcol="4" gridrow="7" pageId="3" pageNumber="792">2.48 1.76</td>
|
||
</tr>
|
||
<tr box="[186,1037,773,813]" gridrow="8" pageId="3" pageNumber="792">
|
||
<td box="[186,442,773,813]" gridcol="0" gridrow="8" pageId="3" pageNumber="792">Metasomal Segment III Length Width</td>
|
||
<td box="[494,598,773,813]" gridcol="1" gridrow="8" pageId="3" pageNumber="792">2.74 1.68</td>
|
||
<td box="[640,764,773,813]" gridcol="2" gridrow="8" pageId="3" pageNumber="792">* *</td>
|
||
<td box="[787,891,773,813]" gridcol="3" gridrow="8" pageId="3" pageNumber="792">2.10 1.52</td>
|
||
<td box="[933,1037,773,813]" gridcol="4" gridrow="8" pageId="3" pageNumber="792">2.77 1.62</td>
|
||
</tr>
|
||
<tr box="[186,1037,839,880]" gridrow="9" pageId="3" pageNumber="792">
|
||
<td box="[186,442,839,880]" gridcol="0" gridrow="9" pageId="3" pageNumber="792">Metasomal Segment IV Length Width</td>
|
||
<td box="[494,598,839,880]" gridcol="1" gridrow="9" pageId="3" pageNumber="792">3.25 1.64</td>
|
||
<td box="[640,764,839,880]" gridcol="2" gridrow="9" pageId="3" pageNumber="792">3.37 1.41</td>
|
||
<td box="[787,891,839,880]" gridcol="3" gridrow="9" pageId="3" pageNumber="792">2.80 1.44</td>
|
||
<td box="[933,1037,839,880]" gridcol="4" gridrow="9" pageId="3" pageNumber="792">3.16 1.65</td>
|
||
</tr>
|
||
<tr box="[186,1037,904,967]" gridrow="10" pageId="3" pageNumber="792">
|
||
<td box="[186,442,904,967]" gridcol="0" gridrow="10" pageId="3" pageNumber="792">Metasomal Segment V Length Width Height</td>
|
||
<td box="[494,598,904,967]" gridcol="1" gridrow="10" pageId="3" pageNumber="792">3.75 1.49 1.60</td>
|
||
<td box="[640,764,904,967]" gridcol="2" gridrow="10" pageId="3" pageNumber="792">3.84 1.34 1.42</td>
|
||
<td box="[787,891,904,967]" gridcol="3" gridrow="10" pageId="3" pageNumber="792">3.61 1.35 1.49</td>
|
||
<td box="[933,1037,904,967]" gridcol="4" gridrow="10" pageId="3" pageNumber="792">3.67 1.58 1.62</td>
|
||
</tr>
|
||
<tr box="[186,1037,993,1056]" gridrow="11" pageId="3" pageNumber="792">
|
||
<td box="[186,442,993,1056]" gridcol="0" gridrow="11" pageId="3" pageNumber="792">Telson Length Width Height</td>
|
||
<td box="[494,598,993,1056]" gridcol="1" gridrow="11" pageId="3" pageNumber="792">3.09 1.22 1.26</td>
|
||
<td box="[640,764,993,1056]" gridcol="2" gridrow="11" pageId="3" pageNumber="792">3.05 1.08 1.19</td>
|
||
<td box="[787,891,993,1056]" gridcol="3" gridrow="11" pageId="3" pageNumber="792">3.05 1.94 1.06</td>
|
||
<td box="[933,1037,993,1056]" gridcol="4" gridrow="11" pageId="3" pageNumber="792">2.12 1.27 1.29</td>
|
||
</tr>
|
||
<tr box="[186,1037,1081,1099]" gridrow="12" pageId="3" pageNumber="792">
|
||
<td box="[186,442,1081,1099]" gridcol="0" gridrow="12" pageId="3" pageNumber="792">Aculeus Length</td>
|
||
<td box="[494,598,1081,1099]" gridcol="1" gridrow="12" pageId="3" pageNumber="792">1.14</td>
|
||
<td box="[640,764,1081,1099]" gridcol="2" gridrow="12" pageId="3" pageNumber="792">1.15</td>
|
||
<td box="[787,891,1081,1099]" gridcol="3" gridrow="12" pageId="3" pageNumber="792">1.02</td>
|
||
<td box="[933,1037,1081,1099]" gridcol="4" gridrow="12" pageId="3" pageNumber="792">1.11</td>
|
||
</tr>
|
||
<tr box="[186,1037,1123,1141]" gridrow="13" pageId="3" pageNumber="792">
|
||
<td box="[186,442,1123,1141]" gridcol="0" gridrow="13" pageId="3" pageNumber="792">Pedipalp Total Length</td>
|
||
<td box="[494,598,1123,1141]" gridcol="1" gridrow="13" pageId="3" pageNumber="792">12.03</td>
|
||
<td box="[640,764,1123,1141]" gridcol="2" gridrow="13" pageId="3" pageNumber="792">11.82</td>
|
||
<td box="[787,891,1123,1141]" gridcol="3" gridrow="13" pageId="3" pageNumber="792">10.82</td>
|
||
<td box="[933,1037,1123,1141]" gridcol="4" gridrow="13" pageId="3" pageNumber="792">11.70</td>
|
||
</tr>
|
||
<tr box="[186,1037,1167,1207]" gridrow="14" pageId="3" pageNumber="792">
|
||
<td box="[186,442,1167,1207]" gridcol="0" gridrow="14" pageId="3" pageNumber="792">Pedipalp femur Length Width</td>
|
||
<td box="[494,598,1167,1207]" gridcol="1" gridrow="14" pageId="3" pageNumber="792">2.94 1.08</td>
|
||
<td box="[640,764,1167,1207]" gridcol="2" gridrow="14" pageId="3" pageNumber="792">3.11 086</td>
|
||
<td box="[787,891,1167,1207]" gridcol="3" gridrow="14" pageId="3" pageNumber="792">2.60 0.91</td>
|
||
<td box="[933,1037,1167,1207]" gridcol="4" gridrow="14" pageId="3" pageNumber="792">3.18 1.11</td>
|
||
</tr>
|
||
<tr box="[186,1037,1234,1275]" gridrow="15" pageId="3" pageNumber="792">
|
||
<td box="[186,442,1234,1275]" gridcol="0" gridrow="15" pageId="3" pageNumber="792">Pedipalp Patella Length Width</td>
|
||
<td box="[494,598,1234,1275]" gridcol="1" gridrow="15" pageId="3" pageNumber="792">3.44 1.33</td>
|
||
<td box="[640,764,1234,1275]" gridcol="2" gridrow="15" pageId="3" pageNumber="792">3.42 1.13</td>
|
||
<td box="[787,891,1234,1275]" gridcol="3" gridrow="15" pageId="3" pageNumber="792">3.32 1.21</td>
|
||
<td box="[933,1037,1234,1275]" gridcol="4" gridrow="15" pageId="3" pageNumber="792">3.45 1.15</td>
|
||
</tr>
|
||
<tr box="[186,1037,1312,1375]" gridrow="16" pageId="3" pageNumber="792">
|
||
<td box="[186,442,1312,1375]" gridcol="0" gridrow="16" pageId="3" pageNumber="792">Pedipalp Chelae Length Width Height</td>
|
||
<td box="[494,598,1312,1375]" gridcol="1" gridrow="16" pageId="3" pageNumber="792">5.65 1.33 1.19</td>
|
||
<td box="[640,764,1312,1375]" gridcol="2" gridrow="16" pageId="3" pageNumber="792">5.29 1.14 1.07</td>
|
||
<td box="[787,891,1312,1375]" gridcol="3" gridrow="16" pageId="3" pageNumber="792">4.90 1.15 1.05</td>
|
||
<td box="[933,1037,1312,1375]" gridcol="4" gridrow="16" pageId="3" pageNumber="792">5.07 1.30 1.32</td>
|
||
</tr>
|
||
<tr box="[186,1037,1411,1429]" gridrow="17" pageId="3" pageNumber="792">
|
||
<td box="[186,442,1411,1429]" gridcol="0" gridrow="17" pageId="3" pageNumber="792">Movable Finger Length</td>
|
||
<td box="[494,598,1411,1429]" gridcol="1" gridrow="17" pageId="3" pageNumber="792">3.38</td>
|
||
<td box="[640,764,1411,1429]" gridcol="2" gridrow="17" pageId="3" pageNumber="792">2.92</td>
|
||
<td box="[787,891,1411,1429]" gridcol="3" gridrow="17" pageId="3" pageNumber="792">2.98</td>
|
||
<td box="[933,1037,1411,1429]" gridcol="4" gridrow="17" pageId="3" pageNumber="792">3.02</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="3.[1073,1355,1392,1432]" pageId="3" pageNumber="792">‘*’ not provided as the specimen is damaged.</paragraph>
|
||
<paragraph blockId="3.[170,782,1505,2008]" pageId="3" pageNumber="792">
|
||
very short and inconspicuous tibial spur on III & IV pairs. Tarsomere I almost as long as tibia but slender. Smooth and clothed with bristles on ventral portion. Tarsomere II delicate, thin and smooth covered thickly with paired bristles on ventral portion. A pair of claws present. Pectines well developed and more than four times as long as wide, middle lamellae divided into 8-9 small digits. Fulcra distinct. Lamellae and fulcra covered with setae. Pectinal teeth well developed; 13/
|
||
<quantity box="[528,580,1761,1784]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="3" pageNumber="792" unit="in" value="14.0">14 in</quantity>
|
||
female (Image 2). Basal piece smooth not much sclerotized and invaginated on anterior margin forming a distinctive anterior notch as if composed of two halves fused medially. Genital operculum wider than long; paired genital papillae visible at the base of the genital operculum in the male. Cephalothoracic sternum triangular possessing a distinct median furrow and a small triangular basal pit.
|
||
</paragraph>
|
||
<paragraph blockId="3.[813,1419,1505,2008]" pageId="3" pageNumber="792">
|
||
Mesosoma: All tergites sparsely granular, more granular posteriorly (
|
||
<figureCitation box="[1052,1164,1537,1560]" captionStart="Images 1" captionStartId="2.[170,234,1300,1318]" captionTargetBox="[170,1418,238,1285]" captionTargetId="figure-27@2.[598,992,238,788]" captionTargetPageId="2" captionText="Images 1 to 9. Lychas aareyensis sp. nov. female holotype (BNHS SC23) (scale 1.0mm) 1 - Carapace and Mesosoma; 2 - Genital operculum and pectinal teeth; 3 - Mesosomal tergites, dorsal; 4 - Mesosomal sternites, ventral; 5 - Chelicerae; 6 - Manus; 7 - Patella; 8 - Metasoma; 9 - Telson" figureDoi="http://doi.org/10.5281/zenodo.4986621" httpUri="https://zenodo.org/record/4986621/files/figure.png" pageId="3" pageNumber="792">Images 1</figureCitation>
|
||
& 3). Tergites I-VI monocarinated. Lateral and posterior margins crenulated. Pretergal portion finely granular. Tergite VIl more granular and with four crenulated carinae. Lateral and posterior margins granular. Sternites III-VI smooth; each sternite with a pair of book lungs (Image 4). Last sternite granular and carinated. Carinae weakly crenulated and granules obsolete. Lateral margins smooth. Posterior margin uneven. Presternal portion short and smooth.
|
||
</paragraph>
|
||
<paragraph blockId="3.[813,1419,1505,2008]" lastBlockId="4.[170,783,868,1371]" lastPageId="4" lastPageNumber="793" pageId="3" pageNumber="792">Metasoma slightly more than five times as long as carapace (Image 8). Basal segment longer than wide. All segments carinated. Segments I and II with all ten carinae (paired ventral, ventrolateral, intermedian, dorsolateral, and dorsal carinae); all carinae crenulated. Segments III & IV with eight carinae (intermedian carinae absent), all carinae crenulated and lateral carinae developed only on half of anterior portion of segment III; segment V with five (axial, paired ventrolateral and dorsolateral carinae). Dorsal carinae on segments I-IV weakly tuberculate posteriorly. Intercarinal portion weakly and much sparsely granular. Anterior and posterior margins of each segment smooth. Segment V almost as long as carapace; only inferior lateral and single inferior median carinae poorly granular and weakly crenulated. Intercarinal space poorly granular. Anal rim of this segment smooth. Telson (Image 9) slightly shorter than segment V. Vesicle as long as segment IV, weakly granular, ventral median crest ending posteriorly into a subaculear tubercle, provided with a pair of minute teeth on inner margin. Aculeus long, as long as the width of the vesicle; sharp, not much curved and dark on distal portion.</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4986627" ID-Zenodo-Dep="4986627" box="[170,904,804,824]" httpUri="https://zenodo.org/record/4986627/files/figure.png" pageId="4" pageNumber="793" startId="4.[170,232,804,824]" targetBox="[170,1257,238,792]" targetPageId="4">
|
||
<paragraph blockId="4.[170,904,804,824]" box="[170,904,804,824]" pageId="4" pageNumber="793">
|
||
Image 13.
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[273,463,804,824]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="793" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[273,463,804,824]" italics="true" pageId="4" pageNumber="793">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[469,549,804,824]" pageId="4" pageNumber="793" rank="species">sp. nov.</taxonomicNameLabel>
|
||
female holotype BNHS SC23 in life
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="4.[170,775,1412,1787]" box="[170,285,1412,1435]" pageId="4" pageNumber="793">
|
||
<emphasis bold="true" box="[170,285,1412,1435]" pageId="4" pageNumber="793">Variations</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="4.[170,775,1412,1787]" pageId="4" pageNumber="793">
|
||
The
|
||
<typeStatus box="[258,363,1444,1467]" pageId="4" pageNumber="793">paratypes</typeStatus>
|
||
match the description of the
|
||
<typeStatus box="[684,775,1444,1467]" pageId="4" pageNumber="793">holotype</typeStatus>
|
||
but the male
|
||
<typeStatus box="[328,421,1476,1499]" pageId="4" pageNumber="793">paratype</typeStatus>
|
||
(BNHS SC24) differs from the
|
||
<typeStatus box="[170,261,1508,1531]" pageId="4" pageNumber="793">holotype</typeStatus>
|
||
as follows: carapace of male less granular than the female (
|
||
<figureCitation box="[302,412,1540,1563]" captionStart="Images 10" captionStartId="2.[170,234,1904,1922]" captionTargetBox="[170,1419,1399,1887]" captionTargetId="figure-126@2.[598,988,1400,1756]" captionTargetPageId="2" captionText="Images 10 to 12. Lychas aareyensis sp. nov. male paratype (BNHS SC24) (scale 1.0mm) 10 - Carapace; 11 - Genital operculum and pectined; 12 - Mesosoma ventral (no scale)" figureDoi="http://doi.org/10.5281/zenodo.4986623" httpUri="https://zenodo.org/record/4986623/files/figure.png" pageId="4" pageNumber="793">Image 10</figureCitation>
|
||
); median eyes situated anteriorly in the ratio 1:1.7; a single external granule present on the immovable finger; pectines 17/
|
||
<quantity box="[546,601,1604,1627]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="4" pageNumber="793" unit="in" value="18.0">18 in</quantity>
|
||
number (Image 11); brightly coloured and with more confluent markings (
|
||
<figureCitation box="[176,292,1668,1691]" captionStart="Images 10" captionStartId="2.[170,234,1904,1922]" captionTargetBox="[170,1419,1399,1887]" captionTargetId="figure-126@2.[598,988,1400,1756]" captionTargetPageId="2" captionText="Images 10 to 12. Lychas aareyensis sp. nov. male paratype (BNHS SC24) (scale 1.0mm) 10 - Carapace; 11 - Genital operculum and pectined; 12 - Mesosoma ventral (no scale)" figureDoi="http://doi.org/10.5281/zenodo.4986623" httpUri="https://zenodo.org/record/4986623/files/figure.png" pageId="4" pageNumber="793">Images 10</figureCitation>
|
||
& 12). The type series range from
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.942" metricValueMax="3.165" metricValueMin="2.719" pageId="4" pageNumber="793" unit="mm" value="29.42" valueMax="31.65" valueMin="27.19">27.19 to 31.65 mm</quantity>
|
||
in total length. The granules on the movable and immovable finger is variable presence and absence of external in the type series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="793" type="discussion">
|
||
<paragraph blockId="4.[170,775,1828,2011]" box="[170,271,1828,1851]" pageId="4" pageNumber="793">
|
||
<emphasis bold="true" box="[170,271,1828,1851]" pageId="4" pageNumber="793">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="4.[170,775,1828,2011]" lastBlockId="4.[813,1418,868,1562]" pageId="4" pageNumber="793">
|
||
A small sized scorpion (average
|
||
<quantity box="[560,699,1860,1883]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="4" pageNumber="793" unit="mm" value="29.0" valueMax="31.0" valueMin="27.0">27 to 31 mm</quantity>
|
||
) which closely resembles
|
||
<emphasis box="[372,572,1892,1915]" italics="true" pageId="4" pageNumber="793">
|
||
<taxonomicName authorityName="C. L. Koch" authorityYear="1845" box="[372,447,1892,1915]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="793" phylum="Arthropoda" rank="genus">Lychas</taxonomicName>
|
||
nigristernis
|
||
</emphasis>
|
||
, but can be easily distinguished (
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[332,537,1924,1947]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="793" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis box="[332,537,1924,1947]" italics="true" pageId="4" pageNumber="793">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[551,642,1924,1947]" pageId="4" pageNumber="793" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs.
|
||
<emphasis italics="true" pageId="4" pageNumber="793">
|
||
<taxonomicName authorityName="C. L. Koch" authorityYear="1845" box="[700,775,1924,1947]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="793" phylum="Arthropoda" rank="genus">Lychas</taxonomicName>
|
||
nigristernis
|
||
</emphasis>
|
||
) as follows- Median eye situated anteriorly in the ratio 1:1.6 or 1:1.7 vs. 1:
|
||
<quantity box="[528,601,1988,2011]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="4" pageNumber="793" unit="in" value="1.2">1.2 in</quantity>
|
||
<emphasis box="[613,768,1988,2011]" italics="true" pageId="4" pageNumber="793">L. nigristermis</emphasis>
|
||
; female with 13-13 pectinal teeth vs.
|
||
<quantity box="[1212,1307,868,891]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="4" pageNumber="793" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14–16 in</quantity>
|
||
female
|
||
<emphasis italics="true" pageId="4" pageNumber="793">L. nigristernis</emphasis>
|
||
; smaller size
|
||
<quantity box="[1068,1168,900,923]" metricMagnitude="-2" metricUnit="m" metricValue="3.165" pageId="4" pageNumber="793" unit="mm" value="31.65">31.65mm</quantity>
|
||
(maximum) vs.
|
||
<quantity box="[1331,1417,900,923]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="4" pageNumber="793" unit="mm" value="45.2">45.2mm</quantity>
|
||
in
|
||
<emphasis box="[842,988,932,955]" italics="true" pageId="4" pageNumber="793">L. nigristernis</emphasis>
|
||
; trichobothrium db between et and est situated closer to est vs. trichobothrium db between dt and et situated closer to et; pedipalp patella in shade of orange with faint brown reticulate markings or spots vs. pedipalp patella entirely black (
|
||
<bibRefCitation author="Kovarik, F." box="[1151,1300,1059,1083]" pageId="4" pageNumber="793" pagination="311 - 371" refId="ref4124" refString="Kovarik, F. (1997). Revision of the genera Lychas and Hemilychas, with description of six new species (Scorpiones: Buthidae). Acta Societas Zoologicae Bohemicae 61: 311 - 371." type="journal article" year="1997">Kovařik 1997</bibRefCitation>
|
||
; Tikader & Bastawade 1983).
|
||
</paragraph>
|
||
<paragraph blockId="4.[813,1418,868,1562]" pageId="4" pageNumber="793">
|
||
<taxonomicName authority="Mirza & Sanap, 2010" authorityName="Mirza & Sanap" authorityYear="2010" box="[850,1047,1124,1147]" class="Arachnida" family="Buthidae" genus="Lychas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="793" phylum="Arthropoda" rank="species" species="aareyensis" status="sp. nov.">
|
||
<emphasis box="[850,1047,1124,1147]" italics="true" pageId="4" pageNumber="793">Lychas aareyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1053,1136,1124,1147]" pageId="4" pageNumber="793" rank="species">sp. nov.</taxonomicNameLabel>
|
||
may be distinguished from all other mainland Indian congeners on the basis of (nonoverlapping character): Median eye situated anteriorly in the ratio 1:1.6 or 1:1.7 (1:
|
||
<quantity box="[1090,1137,1220,1243]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="793" unit="in" value="2.0">2 in</quantity>
|
||
<emphasis box="[1146,1291,1220,1243]" italics="true" pageId="4" pageNumber="793">L. nigristernis</emphasis>
|
||
; 1:
|
||
<quantity box="[1318,1388,1220,1243]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="4" pageNumber="793" unit="in" value="2.5">2.5 in</quantity>
|
||
<emphasis italics="true" pageId="4" pageNumber="793">L. gravelyi</emphasis>
|
||
; 1: to 1:
|
||
<quantity box="[971,1036,1252,1275]" metricMagnitude="-2" metricUnit="m" metricValue="5.334" pageId="4" pageNumber="793" unit="in" value="2.1">2.1 in</quantity>
|
||
<emphasis box="[1041,1191,1252,1275]" italics="true" pageId="4" pageNumber="793">L. mucronatus</emphasis>
|
||
; 1:
|
||
<quantity box="[1214,1295,1252,1275]" metricMagnitude="-2" metricUnit="m" metricValue="5.715" pageId="4" pageNumber="793" unit="in" value="2.25">2.25 in</quantity>
|
||
<emphasis box="[1300,1410,1252,1275]" italics="true" pageId="4" pageNumber="793">L. rugosus</emphasis>
|
||
; 1:
|
||
<quantity box="[823,914,1284,1307]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" pageId="4" pageNumber="793" unit="in" value="2.75">2.75 in</quantity>
|
||
<emphasis box="[930,1085,1284,1307]" italics="true" pageId="4" pageNumber="793">L. hendersoni</emphasis>
|
||
), mesosomal tergites monocarinated (tri-carinated in
|
||
<emphasis box="[1114,1263,1316,1339]" italics="true" pageId="4" pageNumber="793">L. tricarinatus</emphasis>
|
||
,
|
||
<emphasis box="[1282,1418,1315,1339]" italics="true" pageId="4" pageNumber="793">L. laeviforns</emphasis>
|
||
and
|
||
<emphasis box="[867,974,1348,1371]" italics="true" pageId="4" pageNumber="793">L. scaber</emphasis>
|
||
), 13-18 pectines (
|
||
<quantity box="[1191,1252,1348,1371]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="4" pageNumber="793" unit="in" value="21.0">21 in</quantity>
|
||
<emphasis box="[1267,1418,1348,1371]" italics="true" pageId="4" pageNumber="793">L. albimanus;</emphasis>
|
||
<quantity box="[813,877,1380,1403]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="4" pageNumber="793" unit="in" value="24.0">24 in</quantity>
|
||
<emphasis box="[895,1043,1380,1403]" italics="true" pageId="4" pageNumber="793">L. biharensis</emphasis>
|
||
;
|
||
<quantity box="[1069,1133,1380,1403]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="793" unit="in" value="20.0">20 in</quantity>
|
||
<emphasis box="[1152,1337,1380,1403]" italics="true" pageId="4" pageNumber="793">L. kamshetensis</emphasis>
|
||
), third metasomal segment with eight keels (six in
|
||
<emphasis box="[1283,1403,1412,1435]" italics="true" pageId="4" pageNumber="793">L. kharpadi</emphasis>
|
||
), trichobothrium db between et and est situated closer to est (trichobothrium db between et and est situated closer to et in
|
||
<emphasis box="[901,1002,1508,1531]" italics="true" pageId="4" pageNumber="793">L. rackae</emphasis>
|
||
; trichobothrium db between dt and et situated closer to et in
|
||
<emphasis box="[1053,1160,1539,1562]" italics="true" pageId="4" pageNumber="793">L. hillyardi</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="794" pageId="4" pageNumber="793" type="distribution">
|
||
<paragraph blockId="4.[813,1418,1604,2011]" box="[813,1176,1604,1627]" pageId="4" pageNumber="793">
|
||
<emphasis bold="true" box="[813,1176,1604,1627]" pageId="4" pageNumber="793">Distribution and Natural History</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="4.[813,1418,1604,2011]" lastBlockId="5.[813,1419,1229,1348]" lastPageId="5" lastPageNumber="794" pageId="4" pageNumber="793">
|
||
The type locality Aarey Milk Colony (
|
||
<superScript attach="none" box="[1252,1301,1635,1659]" fontSize="5" pageId="4" pageNumber="793">1908</superScript>
|
||
’43.73”N &
|
||
<superScript attach="none" box="[813,876,1668,1691]" fontSize="5" pageId="4" pageNumber="793">72052</superScript>
|
||
’55.39”E,
|
||
<quantity box="[980,1041,1668,1691]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="793" unit="m" value="103.0">103m</quantity>
|
||
) is located to the south of Sanjay Gandhi National Park (
|
||
<figureCitation box="[1056,1176,1700,1723]" captionStart="Image 18" captionStartId="6.[170,232,787,807]" captionTargetBox="[170,1416,239,769]" captionTargetId="figure-282@6.[167,1419,237,772]" captionTargetPageId="6" captionText="Image 18. Aarey Milk Colony, the type locality in Mumbai." figureDoi="http://doi.org/10.5281/zenodo.4986633" httpUri="https://zenodo.org/record/4986633/files/figure.png" pageId="4" pageNumber="793">Images 18</figureCitation>
|
||
). The habitat in which this species has been found is varied, from scrub to dense deciduous forest (
|
||
<figureCitation box="[1009,1192,1764,1787]" captionStart="Images 16 & 17" captionStartId="5.[170,244,1166,1186]" captionTargetBox="[172,1417,749,1151]" captionTargetPageId="5" captionText="Images 16 & 17. Habitat in which specimens of Lychas aareyensis sp. nov. were found." figureDoi="http://doi.org/10.5281/zenodo.4986631" httpUri="https://zenodo.org/record/4986631/files/figure.png" pageId="4" pageNumber="793">Images 16 & 17</figureCitation>
|
||
) to open patches in the forest. As the forest in the Sanjay Gandhi National Park and adjoining protected areas share a similar biotype, it is quite possible that this species may yet be found in those areas as well as other parts of the northern Western Ghats. The scorpions were found mostly under boulders and on roads during the night. The
|
||
<typeStatus box="[1327,1418,1956,1979]" pageId="4" pageNumber="793">holotype</typeStatus>
|
||
was found under a boulder from under a tree and all the
|
||
<typeStatus box="[170,275,1229,1252]" pageId="5" pageNumber="794">paratypes</typeStatus>
|
||
were found on a road in a single night within ca.
|
||
<specimenCount box="[214,281,1261,1284]" pageId="5" pageNumber="794" type="generic">
|
||
<quantity box="[214,277,1261,1284]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="794" unit="m" value="500.0">500m</quantity>
|
||
.
|
||
</specimenCount>
|
||
A female specimen collected on
|
||
<date pageId="5" pageNumber="794" value="2008-03-31">31 March 2008</date>
|
||
was retained in captivity for observations and on the
|
||
<date box="[170,312,1325,1348]" pageId="5" pageNumber="794" value="2008-05-14">14 May 2008</date>
|
||
it gave birth
|
||
<specimenCount box="[456,584,1325,1348]" pageId="5" pageNumber="794" type="juvenile">13 juveniles</specimenCount>
|
||
(
|
||
<figureCitation box="[599,707,1325,1348]" captionStart="Images 14 & 15" captionStartId="5.[170,244,687,707]" captionTargetBox="[171,1418,237,669]" captionTargetPageId="5" captionText="Images 14 & 15. Lychas aareyensis sp. nov. female with juveniles (not collected)" figureDoi="http://doi.org/10.5281/zenodo.4986629" httpUri="https://zenodo.org/record/4986629/files/figure.png" pageId="5" pageNumber="794">Image 14</figureCitation>
|
||
). The female started giving birth around 1100hr and continued till 1237hr. The female while giving birth had spread its pedipalp wide and open, Metasoma raised, front two pair of legs were held together to support the juvenile which would emerge from the genital operculum and climb on its back. The female would sway its pedipalp and metasoma as the juvenile emerged and would take a pause of 20 to 30 second after giving birth to a single juvenile. The juveniles stayed with the female until their third molt and later abandoned their mother. The juveniles just at birth were light colored and turned gradually darker as they grew older. While in captivity, the female was fed on a diet of cultured beetle larvae which were readily accepted. The scorpion would hold the larvae in its pedipalp far away from its body; sting it repeatedly until the prey ceased all signs of life and initiate the process of devouring it. We recorded other females (not collected) with six (
|
||
<figureCitation box="[269,376,1901,1924]" captionStart="Images 14 & 15" captionStartId="5.[170,244,687,707]" captionTargetBox="[171,1418,237,669]" captionTargetPageId="5" captionText="Images 14 & 15. Lychas aareyensis sp. nov. female with juveniles (not collected)" figureDoi="http://doi.org/10.5281/zenodo.4986629" httpUri="https://zenodo.org/record/4986629/files/figure.png" pageId="5" pageNumber="794">Image 15</figureCitation>
|
||
) and another with
|
||
<specimenCount box="[590,745,1901,1924]" pageId="5" pageNumber="794" type="juvenile">eight juveniles</specimenCount>
|
||
in the field in the month of April. Members of this genus are also referred to as bark scorpions. Although numerous
|
||
<taxonomicName box="[170,337,1997,2020]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="794" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[170,245,1997,2020]" italics="true" pageId="5" pageNumber="794">Lychas</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have been found mostly under bark of trees, this species has only been found on the ground. The sympatric species observed at the type locality were
|
||
<emphasis box="[813,1019,1293,1316]" italics="true" pageId="5" pageNumber="794">
|
||
<taxonomicName box="[813,930,1293,1316]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="794" phylum="Arthropoda" rank="genus">Hottentotta</taxonomicName>
|
||
tumulus
|
||
</emphasis>
|
||
,
|
||
<emphasis box="[1032,1255,1293,1316]" italics="true" pageId="5" pageNumber="794">
|
||
<taxonomicName box="[1032,1149,1293,1316]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="794" phylum="Arthropoda" rank="genus">Hottentotta</taxonomicName>
|
||
rugiscutis
|
||
</emphasis>
|
||
,
|
||
<taxonomicName box="[1268,1415,1293,1316]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="794" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[1268,1373,1293,1316]" italics="true" pageId="5" pageNumber="794">Isometrus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<emphasis box="[813,1054,1325,1348]" italics="true" pageId="5" pageNumber="794">
|
||
<taxonomicName box="[813,958,1325,1348]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="794" phylum="Arthropoda" rank="genus">Heterometrus</taxonomicName>
|
||
phipsoni
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |