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<document ID-CLB-Dataset="7250" ID-DOI="http://dx.doi.org/10.3897/mycokeys.79.63363" ID-GBIF-Dataset="cec973b7-30eb-4570-afc1-159895cffa7c" ID-PMC="PMC8076164" ID-Pensoft-Pub="1314-4049-79-17" ID-Pensoft-UUID="CEDC3742C169540583072E7995018533" ID-PubMed="33958950" ModsDocID="1314-4049-79-17" checkinTime="1618878569608" checkinUser="pensoft" docAuthor="Eberhardt, Ursula, Beker, Henry J., Borgen, Torbjorn, Knudsen, Henning, Schuetz, Nicole &amp; Elborne, Steen A." docDate="2021" docId="14615E2CC0955AB7967B246FAB3F8AF3" docLanguage="en" docName="MycoKeys 79: 17-118" docOrigin="MycoKeys 79" docPubDate="2021-04-19" docSource="http://dx.doi.org/10.3897/mycokeys.79.63363" docTitle="Hebeloma grandisporum Beker, U. Eberh. &amp; A. Ronikier; Eberhardt, Ronikier, Schuetz &amp; Beker, Mycologia 107 (6): 1293 2015" docType="treatment" docVersion="6" id="CEDC3742C169540583072E7995018533" lastPageNumber="17" masterDocId="CEDC3742C169540583072E7995018533" masterDocTitle="A survey of Hebeloma (Hymenogastraceae) in Greenland" masterLastPageNumber="118" masterPageNumber="17" pageNumber="17" updateTime="1720462698628" updateUser="ExternalLinkService">
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<mods:title id="2D38BF5E85CA296C1E5186EFF47F287A">A survey of Hebeloma (Hymenogastraceae) in Greenland</mods:title>
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<mods:namePart id="1404CF2FDD88A1C64D7EB9D85954C0EF">Eberhardt, Ursula</mods:namePart>
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<mods:namePart id="DB726AF198E13A31773423B49C2A9E55">Beker, Henry J.</mods:namePart>
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<mods:affiliation id="7B63CF0FFBE8741DB175A34B93A796B1">Rue Pere de Deken 19, B- 1040 Bruxelles, Belgium &amp; Royal Holloway College, University of London, Egham, UK &amp; Plantentuin Meise, Nieuwelaan 38, B- 1860 Meise, Belgium</mods:affiliation>
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<mods:namePart id="90F7E9A940AEBCAEBF74E5410B90A065">Borgen, Torbjorn</mods:namePart>
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<mods:affiliation id="5A2396F158949CF48285813EBE81BAD7">Sensommervej 142, 8600 Silkeborg, Denmark</mods:affiliation>
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<mods:namePart id="16141D37A9EFF06A30A2B7A44DFEC585">Schuetz, Nicole</mods:namePart>
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<treatment id="14615E2CC0955AB7967B246FAB3F8AF3" ID-GBIF-Taxon="181518935" LSID="urn:lsid:plazi:treatment:14615E2CC0955AB7967B246FAB3F8AF3" httpUri="http://treatment.plazi.org/id/14615E2CC0955AB7967B246FAB3F8AF3" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection id="FD7B59EAD0B639B46659FD8E5959CE6D" pageId="0" pageNumber="17" type="nomenclature">
<paragraph id="E45C364F166CFE3FECF635822818D3F2" pageId="0" pageNumber="17">
<taxonomicName id="0C276C203B63089A42C1F6E822D250DF" ID-CoL="6LGZL" LSID="14615E2C-C095-5AB7-967B-246FAB3F8AF3" authority="Beker, U. Eberh. &amp; A. Ronikier; Eberhardt, Ronikier, Schuetz &amp; Beker, Mycologia 107 (6): 1293, 2015." authorityName="Beker, U. Eberh. &amp; A. Ronikier; Eberhardt, Ronikier, Schuetz &amp; Beker, Mycologia 107 (6): 1293" authorityYear="2015" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="treatment-meta" kingdom="Fungi" lsidName="Hebeloma grandisporum" order="Agaricales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="grandisporum">
Hebeloma grandisporum Beker, U. Eberh. &amp; A. Ronikier; Eberhardt, Ronikier,
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&amp; Beker, Mycologia 107(6): 1293, 2015.
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<figureCitation id="A19735B2F29BE63505829019F8B12E48" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Hebeloma grandisporum A HK 17.101, photograph H. Knudsen B distribution of cited collections C spores x 1600 and D cheilocystidia x 1000 of HK 17.101 in Melzer's reagent. Scale bars: 5 µm; microphotographs H. K. J. Beker." figureDoi="10.3897/mycokeys.79.63363.figure13" httpUri="https://binary.pensoft.net/fig/533404" pageId="0" pageNumber="17">Fig. 13</figureCitation>
</paragraph>
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<subSubSection id="BA215B1005E6464734BC18BCA759F60E" pageId="0" pageNumber="17" type="description">
<paragraph id="BBD167535D661E64344C740E7F7DCB9E" pageId="0" pageNumber="17">Macroscopic description.</paragraph>
<paragraph id="AEA6A720A585C72A2D883B4E46349B44" pageId="0" pageNumber="17">
Cap up to 1.0 cm in diameter, convex to umbonate, margin usually smooth, but can be crenulate, becoming upturned with age, tacky when moist, with slight spotting, not hygrophanous, uniformly colored or bicolored, at center clay buff or pale yellowish brown or cinnamon, at margin paler, universal veil absent, partial veil present. Lamellae clay color, emarginate to adnate, maximum depth not recorded, number of lamellae {L}18-28; droplets absent, white fimbriate edge indistinct. Stem 1.0-2.0
<normalizedToken id="F678B22D7E33250DE62A2293664FF312" originalValue="×">x</normalizedToken>
0.2-0.3 cm, ochre-brown, paler in apex, cylindrical, stem Q 6.0-8.5, fibrillose, pruinose at apex. Context firm, stem interior stuffed, stem flesh not discoloring from base. Smell weakly raphanoid or insignificant. Taste not recorded. Spore deposit not recorded.
</paragraph>
<caption id="771184EB077AC22DF29178B1D95CABCF" doi="10.3897/mycokeys.79.63363.figure13" httpUri="https://binary.pensoft.net/fig/533404" pageId="0" pageNumber="17" start="Figure 13" startId="F13">
<paragraph id="9CD3B5A1A74B6E48358C033528CD9EC9" pageId="0" pageNumber="17">
<emphasis id="C93806E0B13CBE557A1E1D41377FBAFE" bold="true" pageId="0" pageNumber="17">Figure 13.</emphasis>
<taxonomicName id="BE82B8ABC32FD951E8C6AF8C4AC26F9B" authorityName="Beker, U. Eberh. &amp; A. Ronikier; Eberhardt, Ronikier, Schuetz &amp; Beker, Mycologia 107 (6): 1293" authorityYear="2015" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma grandisporum" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="grandisporum">
<emphasis id="60F2224058D3FE5DA41C2E3FE90F06BE" italics="true" pageId="0" pageNumber="17">Hebeloma grandisporum</emphasis>
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<emphasis id="F06C324736FCE56D07CCD98714CC3375" bold="true" pageId="0" pageNumber="17">A</emphasis>
HK17.101, photograph H. Knudsen
<emphasis id="F31C9DBD40D3C24D66BBBD41BE8E4E81" bold="true" pageId="0" pageNumber="17">B</emphasis>
distribution of cited collections
<emphasis id="8319C0AC130AFEB9323FE25200D4A3F1" bold="true" pageId="0" pageNumber="17">C</emphasis>
spores
<normalizedToken id="0D85F7057BEDD7EB36BD3F31F9E11BFC" originalValue="×">x</normalizedToken>
1600 and
<emphasis id="EDA701A2057493EA6A1C17A5BF6A4C1C" bold="true" pageId="0" pageNumber="17">D</emphasis>
cheilocystidia
<normalizedToken id="F50F279C7254CC74FD0816612614BE6D" originalValue="×">x</normalizedToken>
1000 of HK17.101 in
<normalizedToken id="909A4F2DC406337EA6A881B0B4139081" originalValue="Melzers">Melzer's</normalizedToken>
reagent. Scale bars: 5
<normalizedToken id="4E93763C5F846FBFB84AA798D4A174BD" originalValue="µm">µm</normalizedToken>
; microphotographs H.K.J. Beker.
</paragraph>
</caption>
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<subSubSection id="33BFC5B557A65C80AE14F2544DC3AF7A" pageId="0" pageNumber="17" type="description">
<paragraph id="E9D4930DDB25FA572E212A7B8AA34777" pageId="0" pageNumber="17">Microscopic description.</paragraph>
<paragraph id="2B03284EA9072385721913FB2279F976" pageId="0" pageNumber="17">
Spores amygdaloid or limoniform, very strongly papillate, on ave. 14-16
<normalizedToken id="8EB9CF7C45D0FC5BE777E2BC18A857C6" originalValue="×">x</normalizedToken>
8.5-9.5
<normalizedToken id="C57B6BABE0842E5707ACAA0BDDBED9DE" originalValue="µm">µm</normalizedToken>
, ave. Q = 1.6-1.8, pale brown to yellow brown, weakly guttulate, almost smooth to weakly ornamented (O1O2), perispore not loosening (P0), distinctly to rather strongly dextrinoid (D2D3). Basidia 23-44
<normalizedToken id="F586017C9777B4A776257E654DDAF899" originalValue="×">x</normalizedToken>
6-10
<normalizedToken id="3F081FE44639C5BBF9E2FFD705A56C2D" originalValue="µm">µm</normalizedToken>
, ave. Q = 3.1-4.3, characteristically 2-spored. Cheilocystidia lageniform or ventricose, occasionally clavate-ventricose, characteristically geniculate and septate, on ave. 40-70
<normalizedToken id="F261D26B2A051CB25FE9D19CB1179347" originalValue="×">x</normalizedToken>
5.5-6.5 (apex)
<normalizedToken id="CA52585AA0F143A575D0E15A72629754" originalValue="×">x</normalizedToken>
4.5-5.5 (middle)
<normalizedToken id="AFDD10DA9B702BAF66FAFB5BB343EA9E" originalValue="×">x</normalizedToken>
8-10.5 (base)
<normalizedToken id="B06A2768F179BE9BEB2552C0154060BB" originalValue="µm">µm</normalizedToken>
, ratios A/M = 1.02-1.44, A/B = 0.53-0.75, B/M = 1.48-2.03. Epicutis an ixocutis, up to 130
<normalizedToken id="9CDADD2AE4E9E9642736C9D69180E919" originalValue="µm">µm</normalizedToken>
thick (measured from exsiccata), maximum hyphae width 6
<normalizedToken id="1EA98512FAE999F49B5254237D2C62B8" originalValue="µm">µm</normalizedToken>
, ixocutis hyphae sometimes encrusted, shape of trama elements beneath subcutis isodiametric. Caulocystidia similar to cheilocystidia, up to 120
<normalizedToken id="EE4369D4FE724DC7FBB9C0C51383174D" originalValue="µm">µm</normalizedToken>
long.
</paragraph>
</subSubSection>
<subSubSection id="CE2922450A74C06E4D4AC81402C7EAA6" pageId="0" pageNumber="17" type="collections examined">
<paragraph id="99A9DA6E938ED81F85077A2B2554325A" pageId="0" pageNumber="17">Collections examined.</paragraph>
<paragraph id="48DF222D10860352EB34BF1E3E1D0000" pageId="0" pageNumber="17">
<emphasis id="D4835FD34D0F01C7F04143F4832AA0FD" bold="true" pageId="0" pageNumber="17">N-Greenland</emphasis>
: Zackenberg,
<normalizedToken id="3C21DE1D6E3E1A595BF4993E5B2C2B6E" originalValue="Sydkæret">Sydkaeret</normalizedToken>
,
<geoCoordinate id="D1D53EA715E4D33F1ACDDF487895764F" degrees="74.50" direction="north" orientation="latitude" precision="555" value="74.5">74.50°N</geoCoordinate>
,
<geoCoordinate id="7E139D878AFAFE6089F96D24B296A303" degrees="20.75" direction="west" orientation="longitude" precision="555" value="-20.75">20.75°W</geoCoordinate>
, 19 Aug 1999, T. Borgen (TB99.376, C-F-104295), 30 m, with
<taxonomicName id="2AB0D12916DD278F471F6584D3EE5F70" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix arctica" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arctica">
<emphasis id="8802BD90905815A32075E260F0D1EE27" italics="true" pageId="0" pageNumber="17">Salix arctica</emphasis>
</taxonomicName>
.
<emphasis id="63BC404B1D664C9E00087494E1752D85" bold="true" pageId="0" pageNumber="17">E-Greenland</emphasis>
: Jameson Land, Nerlerit Inaat/Constable Pynt, west side of Nathorst Fjeld,
<geoCoordinate id="44B8735843F5CEC11A68B9DE9B49E358" degrees="70.76" direction="north" orientation="latitude" precision="555" value="70.76">70.76°N</geoCoordinate>
,
<geoCoordinate id="B6A069C383CC1B7EFC6A86B552D1DE22" degrees="22.64" direction="west" orientation="longitude" precision="555" value="-22.64">22.64°W</geoCoordinate>
, 5 Aug 2017, H. Knudsen (HK17.101, C-F-104997), 40 m, with
<taxonomicName id="FF2BA00DA260B5D004B8C3AF02D4BEF2" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="genus">
<emphasis id="4A1221D7512BC25DB57299A92AD005A7" italics="true" pageId="0" pageNumber="17">Salix</emphasis>
</taxonomicName>
sp. in tundra.
</paragraph>
</subSubSection>
<subSubSection id="1446EEDF530FAE523BFC14AB07D9E626" pageId="0" pageNumber="17" type="distribution">
<paragraph id="D4C0584DECCEC5F35DD6A0C94269E348" pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph id="530755663C90D85F214556B3BCE50627" pageId="0" pageNumber="17">Recently described from the southern Carpathians in Romania from a single collection. The two Greenland collections are the first recorded since the type has been published. Both are from the High Arctic zone in Greenland at 70° and 74°N, and it is thus in the northernmost group among the 28 Greenland species. The records are the first from North America and the first outside Europe.</paragraph>
</subSubSection>
<subSubSection id="41ACEDDB5A627BD65968CCBA794DAB89" pageId="0" pageNumber="17" type="habitat">
<paragraph id="FB8A27F84CF5FAAD404B42F9C1DE05BF" pageId="0" pageNumber="17">Habitat and ecology.</paragraph>
<paragraph id="B242944A2B41BC599787EDF40E3963F6" pageId="0" pageNumber="17">
Only two records, both most likely with
<taxonomicName id="8F5C91252B1BD90C030CD1A6F3FCBEA0" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix arctica" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arctica">
<emphasis id="C14DD914F7F360545A0EF4222710179D" italics="true" pageId="0" pageNumber="17">Salix arctica</emphasis>
</taxonomicName>
and both on calcareous soil. The only previous collection, the type, grew with
<taxonomicName id="1065E1A5BE50C77F6BCA327311AD553A" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="S. retusa" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="retusa">
<emphasis id="65F674F69976E3670E0D267AA2F99EF7" italics="true" pageId="0" pageNumber="17">S. retusa</emphasis>
</taxonomicName>
L. at 2270 m a.s.l. in the Carpathians (
<bibRefCitation id="D30894D247D170480C9E480B298F5B9D" DOI="https://doi.org/10.3852/15-097" author="Eberhardt, U" journalOrPublisher="Mycologia" pageId="0" pageNumber="17" pagination="1285 - 1303" refId="B11" refString="Eberhardt, U, Ronikier, A, Schuetz, N, Beker, HJ, 2015a. The genus Hebeloma in the alpine belt of the Carpathians including two new species. Mycologia 107: 1285 - 1303, DOI: https://doi.org/10.3852/15-097" title="The genus Hebeloma in the alpine belt of the Carpathians including two new species." url="https://doi.org/10.3852/15-097" volume="107" year="2015 a">Eberhardt et al. 2015a</bibRefCitation>
).
<bibRefCitation id="B85CDCBC3756DE1195990058F94CFE60" DOI="https://doi.org/10.1134/S1995425518050086" author="Shiryaev, AG" journalOrPublisher="Contemporary Problems of Ecology" pageId="0" pageNumber="17" pagination="458 - 471" refId="B44" refString="Shiryaev, AG, Zmitrovich, IV, Ezhov, ON, 2018. Taxonomic and ecological structure of basidial macromycetes biota in polar deserts of the northern hemisphere. Contemporary Problems of Ecology 11: 458 - 471, DOI: https://doi.org/10.1134/S1995425518050086" title="Taxonomic and ecological structure of basidial macromycetes biota in polar deserts of the northern hemisphere." url="https://doi.org/10.1134/S1995425518050086" volume="11" year="2018">Shiryaev et al. (2018)</bibRefCitation>
reported from
<bibRefCitation id="3C2B99F1AEF5C8C0B54A97D955F3AADF" author="Zmitrovich, IV" editor="Matveeva, NV" journalOrPublisher="Marafon, St. Petersburg" pageId="0" pageNumber="17" pagination="211 - 225" refId="B54" refString="Zmitrovich, IV, Ezhov, ON, 2015. Agaricoid fungi (Basidiomycota, Agaricomycetes). In: Matveeva, NV, Ed., Rasteniya i griby polyarnykh pustyn' severnogo polushariya (The Plants andFungi of Polar Deserts of Northern Hemisphere). Marafon, St. Petersburg: 211 - 225" title="Agaricoid fungi (Basidiomycota, Agaricomycetes)." volumeTitle="Rasteniya i griby polyarnykh pustyn' severnogo polushariya (The Plants andFungi of Polar Deserts of Northern Hemisphere)." year="2015">Zmitrovich and Ezhov (2015)</bibRefCitation>
the presence of
<taxonomicName id="18555FF24E1492CB1767DC3564C1FD89" authorityName="Groger &amp; Zschieschang" authorityYear="1981" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma gigaspermum" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="gigaspermum">
<emphasis id="EBE8E95DBF165453B26100EF75884768" italics="true" pageId="0" pageNumber="17">Hebeloma gigaspermum</emphasis>
</taxonomicName>
<normalizedToken id="4C6EA890D0B58B97A64A85C5B85CB87D" originalValue="Gröger">Groeger</normalizedToken>
&amp; Zschiesch. from Franz Josefs Land (Russia). According to
<bibRefCitation id="04782AC41FA9B3D322F51FEF13BA899D" author="Beker, HJ" journalOrPublisher="Fungi Europaei Edizioni Tecnografica, Lomazzo" pageId="0" pageNumber="17" refId="B4" refString="Beker, HJ, Eberhardt, U, Vesterholt, J, 2016. Hebeloma (Fr.) P. Kumm. Fungi Europaei Edizioni Tecnografica, Lomazzo" title="Hebeloma (Fr.) P. Kumm." year="2016">Beker et al. (2016)</bibRefCitation>
,
<taxonomicName id="2F24F2F5E645EE103DBB387763DB127A" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. gigaspermum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="gigaspermum">
<emphasis id="D44027BA8F3EB50EFD380DA48A2E2F13" italics="true" pageId="0" pageNumber="17">H. gigaspermum</emphasis>
</taxonomicName>
is most likely a synonym of
<taxonomicName id="66E6E030ED63767481D3FCC1744D61A3" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. nauseosum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="nauseosum">
<emphasis id="CD7C2B452ACA5DA7CDEED18476D94BB5" italics="true" pageId="0" pageNumber="17">H. nauseosum</emphasis>
</taxonomicName>
Sacc. Since neither
<taxonomicName id="60794AE5176474B6E7C9C8398A9564D6" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. nauseosum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="nauseosum">
<emphasis id="95D32FFED246CACAA7522FC08BD843DC" italics="true" pageId="0" pageNumber="17">H. nauseosum</emphasis>
</taxonomicName>
nor any other species of
<taxonomicName id="B5E26F3A4FE16A779AFF303ACC082DAA" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. subsp. sect." order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="section" section="Sacchariolentia">H. sect. Sacchariolentia</taxonomicName>
occurs in arctic areas, it may be that the large spores and the occurrence in the High Arctic zone rather point to
<taxonomicName id="C3E50F8CD403C4D9C9007F11D2FBC5FB" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. grandisporum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="grandisporum">
<emphasis id="E1E058D2BDFEAFA32D41ED04BAC102C8" italics="true" pageId="0" pageNumber="17">H. grandisporum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B75DAABEEFC8C45EECFA499F7B830B2C" pageId="0" pageNumber="17" type="notes">
<paragraph id="7E21B65AA54FAE94A18B62F1EB7555C1" pageId="0" pageNumber="17">Notes.</paragraph>
<paragraph id="FBE127D91DEBC34CE00173A5EE13A95D" pageId="0" pageNumber="17">
<taxonomicName id="84055DAE8AC29903A47ECF2DB2AD74E3" authorityName="Beker, U. Eberh. &amp; A. Ronikier; Eberhardt, Ronikier, Schuetz &amp; Beker, Mycologia 107 (6): 1293" authorityYear="2015" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma grandisporum" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="grandisporum">
<emphasis id="5891C669B472047B4649E12D7E3F5FE5" italics="true" pageId="0" pageNumber="17">Hebeloma grandisporum</emphasis>
</taxonomicName>
is one of the most easily recognized among the arctic-alpine
<taxonomicName id="9560E6A1515AD05EA37F2DB2A72B98D9" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="genus">
<emphasis id="C3AC3DCCF094B27D89C1528B062DD4ED" italics="true" pageId="0" pageNumber="17">Hebeloma</emphasis>
</taxonomicName>
species by the small basidiomes, the large spores, the relatively few, distant lamellae and large, 2-spored basidia. To date, it is the only
<taxonomicName id="EFDF77760592D5C204D8B2F464300942" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="genus">
<emphasis id="A8DCF94D47620A447C4D0E097FCBA376" italics="true" pageId="0" pageNumber="17">Hebeloma</emphasis>
</taxonomicName>
, of which we are aware, that has consistently 2-spored basidia.
</paragraph>
</subSubSection>
</treatment>
</document>