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<document id="850976C1A301E7EE3566A88C1A0EB73B" ID-CLB-Dataset="46041" ID-DOI="10.5281/zenodo.279058" ID-GBIF-Dataset="aadbfc66-59f3-4ecb-af49-5788c22fa7e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="279058" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460407254231" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="03EE87BA5A3BFFEA3C9078F5FC51FE2A" docLanguage="en" docName="zt03093p034.pdf" docOrigin="Zootaxa 3093" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochthebius Perkins, 2011, new species" docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="FFD7FFC25A35FFE53C077B5AFF96FFE5" masterDocTitle="New species and new collection records of aquatic beetles in the genus Ochthebius Leach from southern Africa (Coleoptera: Hydraenidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="15" updateTime="1698299945171" updateUser="ExternalLinkService">
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<mods:title id="4F3CAA895C72F27681EE0B3E49C28782">New species and new collection records of aquatic beetles in the genus Ochthebius Leach from southern Africa (Coleoptera: Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="ECB46EE02891257A242CBC62D0FE1024" type="personal">
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<mods:namePart id="794EA9E3559BF2B921DBD7F94E970796">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="09FD6A6206589C234981A0099B0CBC3D">text</mods:typeOfResource>
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<mods:part id="CCE1DD9765CDB90A93A384EBD6F45472">
<mods:date id="B915735176193CBFF9899EBA1150E24F">2011</mods:date>
<mods:detail id="7D4E9B3269FA46C9787CE4D8E9388AEE" type="volume">
<mods:number id="4AD5977F6246AB7A404018D523A2CBDC">3093</mods:number>
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<mods:classification id="CCA6F7EC1244C08CD02718B5C07112B4">journal article</mods:classification>
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<mods:identifier id="D73C9617E40AD82C56582014A0DEEFCF" type="Zenodo-Dep">279058</mods:identifier>
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<treatment id="03EE87BA5A3BFFEA3C9078F5FC51FE2A" ID-DOI="http://doi.org/10.5281/zenodo.6194041" ID-GBIF-Taxon="119422227" ID-Zenodo-Dep="6194041" LSID="urn:lsid:plazi:treatment:03EE87BA5A3BFFEA3C9078F5FC51FE2A" httpUri="http://treatment.plazi.org/id/03EE87BA5A3BFFEA3C9078F5FC51FE2A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="C35D65275A3BFFEB3C9078F5FEDCFC0F" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BF836AC5A3BFFEB3C9078F5FD17FC2C" blockId="14.[151,641,943,1002]" box="[151,641,943,969]" pageId="14" pageNumber="15">
<heading id="D0B081C05A3BFFEB3C9078F5FD17FC2C" bold="true" box="[151,641,943,969]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B933EABE5A3BFFEB3C9078F5FD17FC2C" bold="true" box="[151,641,943,969]" pageId="14" pageNumber="15">
<taxonomicName id="4C474D2F5A3BFFEB3C9078F5FEB4FC2C" ID-CoL="8VRWR" box="[151,290,943,969]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B933EABE5A3BFFEB3C9078F5FEB4FC2C" bold="true" box="[151,290,943,969]" italics="true" pageId="14" pageNumber="15">Ochthebius</emphasis>
</taxonomicName>
(
<emphasis id="B933EABE5A3BFFEB3D3578F5FECEFC2C" bold="true" box="[306,344,943,969]" italics="true" pageId="14" pageNumber="15">s.s.</emphasis>
)
<taxonomicName id="4C474D2F5A3BFFEB3D6F78F5FE74FC2C" box="[360,482,943,969]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bicomicus" status="sp. nov.">
<emphasis id="B933EABE5A3BFFEB3D6F78F5FE74FC2C" bold="true" box="[360,482,943,969]" italics="true" pageId="14" pageNumber="15">bicomicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20057C55A3BFFEB3DF578F5FD17FC2C" box="[498,641,943,969]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF836AC5A3BFFEB3C907888FEDCFC0F" blockId="14.[151,641,943,1002]" box="[151,330,978,1002]" pageId="14" pageNumber="15">
(
<figureCitation id="137C2A295A3BFFEB3C987888FF78FC0F" box="[159,238,978,1002]" captionStart="FIGURE 9" captionStartId="15.[151,250,1977,2000]" captionTargetBox="[263,1343,534,1927]" captionTargetId="figure@15.[233,1353,519,1955]" captionTargetPageId="15" captionText="FIGURE 9. Ochthebius bicomicus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279064/files/figure.png" pageId="14" pageNumber="15">Figs. 9</figureCitation>
,
<figureCitation id="137C2A295A3BFFEB3CFD7888FE8EFC0F" box="[250,280,978,1002]" captionStart="FIGURE 10" captionStartId="16.[151,250,1605,1628]" captionText="FIGURE 10. Ochthebius bicomicus, aedeagus of holotype." pageId="14" pageNumber="15">10</figureCitation>
,
<figureCitation id="137C2A295A3BFFEB3D237888FED7FC0F" box="[292,321,978,1002]" captionStart="FIGURES 19 20" captionStartId="29.[151,264,1650,1673]" captionTargetBox="[269,1320,202,1620]" captionTargetId="figure@29.[258,1328,193,1629]" captionTargetPageId="29" captionText="FIGURES 19 20. Geographical distributions of Ochthebius. 19. O. endroedyi. 20. O. anchorus, O. bicomicus, O. granulinus, O. involatus, and O. zulu." httpUri="https://zenodo.org/record/279071/files/figure.png" pageId="14" pageNumber="15">20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35D65275A3BFFEB3C907F41FB57FB27" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BF836AC5A3BFFEB3C907F41FB10FBB3" blockId="14.[151,1437,1050,2010]" pageId="14" pageNumber="15">
<emphasis id="B933EABE5A3BFFEB3C907F41FE2DFBD6" bold="true" box="[151,443,1050,1075]" pageId="14" pageNumber="15">
<typeStatus id="54FC880E5A3BFFEB3C907F41FF46FBD6" box="[151,208,1051,1075]" pageId="14" pageNumber="15">Type</typeStatus>
Material.
<typeStatus id="54FC880E5A3BFFEB3D577F41FE2DFBD6" box="[336,443,1051,1075]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<emphasis id="B933EABE5A3BFFEB3E217F40FC6BFBD6" bold="true" box="[550,1021,1050,1075]" pageId="14" pageNumber="15">
<collectingCountry id="F350763C5A3BFFEB3E217F40FD53FBD6" box="[550,709,1050,1075]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
: Western Cape Province:
</emphasis>
<quantity id="4CBF9B495A3BFFEB380A7F40FBC4FBD7" box="[1037,1106,1050,1074]" metricMagnitude="4" metricUnit="m" metricValue="3.5405568" pageId="14" pageNumber="15" unit="mi" value="22.0">22 mi</quantity>
. N. Nelspoort,
<geoCoordinate id="EE73506B5A3BFFEB38F97F40FAF1FBD7" box="[1278,1383,1050,1074]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-31.85">31° 51' S</geoCoordinate>
,
<geoCoordinate id="EE73506B5A3BFFEB39727F40FF45FBB3" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="22.95">22° 57' E</geoCoordinate>
,
<date id="FFF9106C5A3BFFEB3CD97F64FEDAFBB3" box="[222,332,1086,1110]" pageId="14" pageNumber="15" value="1968-03-03">3 iii 1968</date>
, P. J. Spangler (
<collectionCode id="ED56AE695A3BFFEB3DFA7F64FDC2FBB3" LSID="urn:lsid:biocol.org:col:34871" box="[509,596,1086,1110]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15">USNM</collectionCode>
).
<emphasis id="B933EABE5A3BFFEB3E607F65FD7EFBB2" bold="true" box="[615,744,1087,1111]" pageId="14" pageNumber="15">
<typeStatus id="54FC880E5A3BFFEB3E607F65FD75FBB2" box="[615,739,1087,1111]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="54FC880E5A3BFFEB3F887F64FC67FBB3" box="[911,1009,1086,1110]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(12
<collectionCode id="ED56AE695A3BFFEB38237F64FBECFBB3" LSID="urn:lsid:biocol.org:col:34871" box="[1060,1146,1086,1110]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15">USNM</collectionCode>
).
</paragraph>
<paragraph id="8BF836AC5A3BFFEB3CC07F38FB57FB27" blockId="14.[151,1437,1050,2010]" pageId="14" pageNumber="15">
<emphasis id="B933EABE5A3BFFEB3CC07F38FE45FB9E" bold="true" box="[199,467,1122,1147]" pageId="14" pageNumber="15">Differential Diagnosis.</emphasis>
This species is quite similar in dorsal habitus to
<taxonomicName id="4C474D2F5A3BFFEB38077F39FBE8FB9E" box="[1024,1150,1123,1147]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sitiensis">
<emphasis id="B933EABE5A3BFFEB38077F39FBE8FB9E" box="[1024,1150,1123,1147]" italics="true" pageId="14" pageNumber="15">O. sitiensis</emphasis>
</taxonomicName>
; both have very coarsely and confluently punctate pronotal foveae (
<figureCitation id="137C2A295A3BFFEB3E737FDCFD55FB7B" box="[628,707,1158,1182]" captionStart="FIGURE 3" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[242,1328,493,1855]" captionTargetId="figure@7.[232,1355,448,1884]" captionTargetPageId="7" captionText="FIGURE 3. Ochthebius sitiensis, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279061/files/figure.png" pageId="14" pageNumber="15">Figs. 3</figureCitation>
,
<figureCitation id="137C2A295A3BFFEB3ED77FDCFD76FB7B" box="[720,736,1158,1182]" captionStart="FIGURE 9" captionStartId="15.[151,250,1977,2000]" captionTargetBox="[263,1343,534,1927]" captionTargetId="figure@15.[233,1353,519,1955]" captionTargetPageId="15" captionText="FIGURE 9. Ochthebius bicomicus, dorsal and lateral habitus of holotype." httpUri="https://zenodo.org/record/279064/files/figure.png" pageId="14" pageNumber="15">9</figureCitation>
). Reliable determinations will be based on examination of the aedeagi, which distinctively differ in shapes of the main piece and the distal piece (
<figureCitation id="137C2A295A3BFFEB383E7FF0FB1FFB27" box="[1081,1161,1194,1218]" captionStart="FIGURE 4" captionStartId="8.[151,250,1624,1647]" captionText="FIGURE 4. Ochthebius sitiensis, aedeagus of holotype." pageId="14" pageNumber="15">Figs. 4</figureCitation>
,
<figureCitation id="137C2A295A3BFFEB38927FF0FB22FB27" box="[1173,1204,1194,1218]" captionStart="FIGURE 10" captionStartId="16.[151,250,1605,1628]" captionText="FIGURE 10. Ochthebius bicomicus, aedeagus of holotype." pageId="14" pageNumber="15">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D65275A3BFFEA3CC07F94FB6FFE62" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="8BF836AC5A3BFFEA3CC07F94FB6FFE62" blockId="14.[151,1437,1050,2010]" lastBlockId="15.[151,1437,151,464]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B933EABE5A3BFFEB3CC07F94FEC7FB02" bold="true" box="[199,337,1230,1255]" pageId="14" pageNumber="15">Description</emphasis>
. Size:
<typeStatus id="54FC880E5A3BFFEB3DAA7F94FD99FB03" box="[429,527,1230,1254]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.67/0.72; head 0.30/0.44; pronotum 0.39/0.60, PA 0.53, PB 0.50; elytra 1.04/0.72.
<emphasis id="B933EABE5A3BFFEB3F117FA8FCF5FAEE" bold="true" box="[790,867,1266,1291]" pageId="14" pageNumber="15">Color:</emphasis>
Head black, weakly aeneous; pronotum brown to dark brown except lighter brown narrowly along base, sometimes with faint metallic reflections; elytra brown, serial punctures dark brownish; antennae, maxillary palpi and legs, except coxae, light brown, venter dark brown.
<emphasis id="B933EABE5A3BFFEB3C907E05FF76FA92" bold="true" box="[151,224,1375,1399]" pageId="14" pageNumber="15">Head:</emphasis>
Eyes rather small, in dorsal aspect 78 weakly convex facets in longest series. Frons shining, distinctly but finely sparsely setiferously punctate, punctures 0.3xef or smaller, setae recumbent, setal length and interstices both 23xef but denser near eyes; interocular foveae large, round, deep, width of each about 2/3 distance separating them, each internally with few setiferous punctures. Clypeus sculpture as frons except punctures slightly denser and sides densely pubescent. Labrum shining, moderately densely setiferously punctate; anterior margin very narrowly upturned from side to side, with minute apicomedian tooth.
<emphasis id="B933EABE5A3BFFEB3F7C7D49FC6BF9CE" bold="true" box="[891,1021,1555,1579]" pageId="14" pageNumber="15">Pronotum:</emphasis>
Subcordate, widest at about anterior 1/4; anterior margin very weakly bisinuate, without postocular emarginations or processes, hyaline border well developed; sides sinuately attenuate to posterior angles; rather wide lateral hyaline border beginning just behind anterior angles, edge weakly arcuate nearly straight over median 0.7, continuing around basal angles and along base. Disc with anterior and posterior transverse impressions rather deep, outer edges of impressions gradually sloping, both impressions deepest in midline, anterior impression somewhat T-shaped, with median subsulcus almost attaining posterior impression; impressions densely coarsely punctate; reliefs shining, coarsely moderately densely punctate but size and spacing of punctures quite irregular, size 13xef, smaller in middle of disc, interstices with exceedingly fine and sparse micropunctulate ground sculpture. Lateral depressions subrugosely strongly punctate, slope continuing transverse curvature of disc but appearing slightly inflated. Setae of discal punctures recumbent, fine and indistinct.
<emphasis id="B933EABE5A3BFFEB3DFE7C21FDD9F876" bold="true" box="[505,591,1915,1939]" pageId="14" pageNumber="15">Elytra:</emphasis>
Widest at middle, sides arcuate, explanate margin narrow, about equal to greatest width of tarsus. Apices in both sexes rather narrowly conjointly rounded. Ten-seriate punctate, series not at all striate-impressed except sutural series behind middle, on disc punctures slightly elongate, small but well impressed, about equal size of largest pronotal punctures, ca. 2xef, interstices ca. 0.51xpd, punctures without distinct granules at anterior margin, distally punctures progressively smaller and more longitudinally separated. Intervals flat or very weakly rounded, weakly shining, ca. 2xpd. Serial setae decumbent, fine and short but distinct, each attaining base of next seta in series; nervure setae indistinct.
<emphasis id="B933EABE5A3AFFEA3F3D7A5EFC05FEF9" bold="true" box="[826,915,260,284]" pageId="15" pageNumber="16">Venter:</emphasis>
Mentum shining, moderately sparsely setiferously punctate. Metaventrite pubescent laterally, disc with moderately large, subrhomboidal, strongly shining area, glabrous except few setiferous punctures anteriorly. Abdominal ventrites 15 entirely clothed with dense hydrofuge pubescence; 6 effacedly microreticulate, weakly shining, sparsely pubescent in distal 1/2.
</paragraph>
</subSubSection>
<subSubSection id="C35D65275A3AFFEA3CC07ACEFB20FE49" box="[199,1206,404,428]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BF836AC5A3AFFEA3CC07ACEFB20FE49" blockId="15.[151,1437,151,464]" box="[199,1206,404,428]" pageId="15" pageNumber="16">
<emphasis id="B933EABE5A3AFFEA3CC07ACEFED8FE49" bold="true" box="[199,334,404,428]" pageId="15" pageNumber="16">Etymology.</emphasis>
Named in reference to the two smile-shaped impressions of the pronotal disc.
</paragraph>
</subSubSection>
<subSubSection id="C35D65275A3AFFEA3CC07AEDFC51FE2A" box="[199,967,439,464]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BF836AC5A3AFFEA3CC07AEDFC51FE2A" blockId="15.[151,1437,151,464]" box="[199,967,439,464]" pageId="15" pageNumber="16">
<emphasis id="B933EABE5A3AFFEA3CC07AEDFEF6FE35" bold="true" box="[199,352,439,464]" pageId="15" pageNumber="16">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54FC880E5A3AFFEA3ECB7AE2FD6AFE35" box="[716,764,440,464]" pageId="15" pageNumber="16">type</typeStatus>
locality (
<figureCitation id="137C2A295A3AFFEA3F617AEDFC2DFE2A" box="[870,955,439,463]" captionStart="FIGURES 19 20" captionStartId="29.[151,264,1650,1673]" captionTargetBox="[269,1320,202,1620]" captionTargetId="figure@29.[258,1328,193,1629]" captionTargetPageId="29" captionText="FIGURES 19 20. Geographical distributions of Ochthebius. 19. O. endroedyi. 20. O. anchorus, O. bicomicus, O. granulinus, O. involatus, and O. zulu." httpUri="https://zenodo.org/record/279071/files/figure.png" pageId="15" pageNumber="16">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>